Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the genus Myoxocephalus: [ 64 ] records

aenaeus, Cottus Mitchill [S. L.] 1814:8 [Report, in part, of Samuel L. Mitchill; ref. 3030] New York, U.S.A. No types known. Appeared as Cottus aeneus in Mitchill 1815:380 [ref. 13292]. •Possibly valid as Megalocottus aenaeus (Mitchill 1814) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus aenaeus (Mitchill 1814) -- (Neelov 1979:123 [ref. 3152], Smith 1986:449 [ref. 22186], Robins & Ray 1986:282 [ref. 23100] as aeneus, Scott & Scott 1988:500 [ref. 25518], McAllister 1990:200 [ref. 14674], Klein-MacPhee in Collette & Klein-MacPhee 2002:349 [ref. 26158], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Afonso et al. 2016:46, 225 [ref. 36199], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Myoxocephalus aenaeus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Atlantic. Habitat: brackish, marine.

anceps, Cottus (Acanthocottus) Sauvage [H.-E.] 1878:145, Pl. 1 (fig. 13) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris (Série 2) v. 1; ref. 3880] U.S.A. Syntypes: MNHN 0000-7241 (1), 0000-7243 (1). Type catalog: Blanc & Hureau 1968:50 [ref. 20738]. •Synonym of Myoxocephalus aenaeus (Mitchill 1814) -- (Smith 1986:450 [ref. 22186]). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

asundensis, Cottus quadricornis Lönnberg [E.] 1932:17, Pl. (figs. 7 a-b) [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Åsunden, Sweden. Syntypes: NRM 20329-32 (1, 5, 3, 3). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

axillaris, Boreocottus Gill [T. N.] 1859:166 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 11; ref. 1760] Awatska Bay, Kamchatka, Russia, Bering Strait. Holotype (unique): ?USNM 24027 (poor condition). •Synonym of Myoxocephalus scorpioides (Fabricius 1780) -- (Neelov 1973:599 [ref. 7219], Neelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:479 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:264 [ref. 33547], Mecklenburg et al. 2016:83 [ref. 34440]). Current status: Synonym of Myoxocephalus scorpioides (Fabricius 1780). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

batrachoides, Myoxocephalus Gilbert [C. H.] & Burke [C. V.] 1912:58, Fig. 11 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Codfish banks west of southern part of Kamchatka, Russia, Albatross station 4798, 51°37'N, 156°21'E, depth 25 fathoms. Holotype (unique): USNM 71809. •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

borkensis, Cottus quadricornis Lönnberg [E.] 1939:4 [Arkiv för Zoologi v. 31 B (no. 2); ref. 17199] Lake Borken [Bork], parishes of Yxnerum and Björsäter, Osthrogothland Province, Sweden. Syntypes: NRM 20327 (2). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

brandtii, Cottus Steindachner [F.] 1867:119 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe v. 4 (no. 14) (for 16 May 1867); ref. 21724] Mouth of Amur River, Russia. Holotype (unique): NMW 11676. Species also appeared on p. 79 of English version [ref. 18764] as brandtii; later illustrated and described in more detail in Steindachner 1867:706, Pl. 3 (figs. 1-2) [p. 6 of separate] [ref. 6393] as brandti in text and brandtii on plate. •Valid as Myoxocephalus brandtii (Steindachner 1867) -- (Watanabe 1960:73 [ref. 11959], Neelov 1979:121 [ref. 3152] as brandti, Yabe in Masuda et al. 1984:326 [ref. 6441] as brandti, Lindberg & Krasyukova 1987:210 [ref. 15964] as brandti, Sokolovsky & Sokolovskaya 1997:60 [ref. 24691] as brandti, Sokolovskaya et al. 1998:10 [ref. 24670] as brandti, Nakabo 2000:645 [ref. 25086] as brandti, Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:404 [ref. 25968], Nakabo 2002:645 [ref. 26001] as brandti, Parin et al. 2002:S80 [ref. 28300], Fedorov et al. 2003:75 [ref. 27579], Parin et al. 2014:260 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138], Prokofiev 2017:648 [ref. 35721], Dyldin et al. 2021:66 [ref. 38804], Dyldin & Orlov 2022:48 [ref. 38972]). Current status: Valid as Myoxocephalus brandtii (Steindachner 1867). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific. Habitat: brackish, marine.

decastrensis, Cottus Kner [R.] 1865:2, Pl. 2 (fig. 1) [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 24; ref. 6174] Decastris Bay at Amur River mouth, northeastern Asia. Holotype (unique): NMW 80072. Appeared as Cottus decastrensis n. sp.? •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662], Neelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:207 [ref. 15964], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138] with question). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

dybowskii, Cottus Hilgendorf [F. M.] 1879:106 [Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1879; ref. 21467] No locality [Japan]. Holotype (unique): ZMB 10628. Type catalog: Paepke & Fricke 1992:282 [ref. 19946]. •Synonym of Myoxocephalus brandtii (Steindachner 1867) -- (Prokofiev 2017:648 [ref. 35721]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Psychrolutidae: Myoxocephalinae.

edomius, Myoxocephalus Jordan [D. S.] & Starks [E. C.] 1904:274, Fig. 18 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Aino village of Edomo, near Mororan, Japan. Holotype: CAS-SU 7707. Paratypes: USNM 50919 (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Myoxocephalus edomius Jordan & Starks 1904 -- (Watanabe 1960:68 [ref. 11959]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neelov 1979:122 [ref. 3152], Lindberg & Krasyukova 1987:212 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Habitat: marine.

ensiger, Ainocottus Jordan [D. S.] & Starks [E. C.] 1904:283, Fig. 23 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate,Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7709. Paratypes: USNM 50921 (not found in 1980). Additional material: CAS-SU 7596 (4). Type catalog: Böhlke 1953:126 [ref. 12291]. •Valid as Ainocottus ensiger Jordan & Starks 1904 -- (Watanabe 1960:87 [ref. 11959], Kim & Youn 1992:63 [ref. 22287]). •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neelov 1979:121 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Choi et al. 2003:576 [ref. 26218], Nakabo 2002:1526 [ref. 26001], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Mecklenburg et al. 2016:79 [ref. 34440], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

fasciatus, Ainocottus Pavlenko [M. N.] 1910:30 [Trudy Obshchestva Estestvoispytatelej Imperatorskom' Kazanskom' Universiteti, Kazan v. 42 (no. 2); ref. 3393] Peter the Great Bay, Russia, Sea of Japan. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neelov 1979:122 [ref. 3152], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

frykenensis, Cottus quadricornis Lönnberg [E.] 1932:14, Pl. (fig. 5) [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Fryken, Sweden. Syntypes: NRM 30315 (1), Mus. of Karlstad's school (1), Uppsala Univ. Zool. Mus. (1). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

gilberti, Oncocottus hexacornis Fowler [H. W.] 1905:366, Figs. 2-5 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 57; ref. 15353] Point Barrow, northern Alaska, U.S.A. Holotype: ANSP not found [ex WIAP 7203]. Paratypes: WIAP (1). Type catalog: Böhlke 1984:65 [ref. 13621] as types not found, specimens perhaps never deposited at ANSP. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies hexacornis (Richardson 1823) -- (Berg 1949:1129 [ref. 20662]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

glacialis, Cottus Richardson [J.] 1855:349, Pl. 24 [The last of the Arctic voyages in search of Sir J. Franklin; ref. 18631] Northumberland Sound, 76°53'N, western Greenland, depth 9 fathoms. No types known. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

groenlandicus, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:185 [Histoire naturelle des poissons v. 4; ref. 998] Greenland. No types known. Type catalog: Blanc & Hureau 1968:60 [ref. 20738]. Based on Cottus scorpius of Fabricius. •Synonym of Myoxocephalus scorpius (Linnaeus 1758), but a valid subspecies -- (Neelov 1973:599 [ref. 7219], Neelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1255 [ref. 13677], Mecklenburg et al. 2007:176 [ref. 29448]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Scott & Scott 1988:509 [ref. 25518], Yabe in Okamura et al. 1995:165 [ref. 22531], Mecklenburg et al. 2002:478 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Mecklenburg et al. 2011:123 [ref. 31212], Parin et al. 2014:265 [ref. 33547], Mecklenburg et al. 2016:85 [ref. 34440]). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

hexacornis, Cottus Richardson [J.] 1823:726 [22] [Narrative of a journey to the shores of the Polar Sea; ref. 18624] Mouth of Tree River near Coppermine River, Arctic Sea about Herschel Island. On p. 22 of separate. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1129 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Mecklenburg et al. 2002:477 [ref. 25968], Parin et al. 2014:262 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

humilis, Cottus Bean [T. H.] 1881:149 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Chamisso Island, Spafarief Bay at mouth of Eschscholtz Bay, Kotzebue Sound, Alaska, U.S.A., Chukchi Sea, Arctic Ocean. Holotype (unique): USNM 27972. Non-types: MNHN 1890-0465 and 0466 [ex USNM] (2) station 3242. Type catalog: Blanc & Hureau 1968:51 [ref. 20738]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Mecklenburg et al. 2016:75 [ref. 34440]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neelov 1979:122 [ref. 3152], Mecklenburg et al. 2002:481 [ref. 25968], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Habitat: marine.

incitus, Myoxocephalus Watanabe [M.] 1958:294, Figs. 121-122, Pl. 21 [Studies of Japanese cottid fishes; ref. 21357] Same, Aomori Prefecture, Japan, depth 10 fathoms. Holotype (unique): not researched. Also described as new in Watanabe 1960:85, Pl. 27 (fig. 1) [ref. 11959]. •Valid as Myoxocephalus incitus Watanabe 1958 -- (Yabe in Masuda et al. 1984:327 [ref. 6441]). •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Distribution: Western North Pacific. Habitat: brackish, marine.

jaok, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:172 [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka coast, Russia. Holotype (unique): ZMB 31656. Type catalog: Blanc & Hureau 1968:60 [ref. 20738], Paepke & Fricke 1992:284 [ref. 19946]. •Valid as Myoxocephalus jaok (Cuvier 1829) -- (Watanabe 1960:75 [ref. 11959], Neelov 1979:122 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:212 [ref. 15964], Kim & Youn 1992:64 [ref. 22287], Sokolovsky & Sokolovskaya 1997:55 [ref. 24691], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:481 [ref. 25968], Nakabo 2002:645 [ref. 26001], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:75 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Bogutskaya & Naseka 2004:190 [ref. 28183], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:261 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:76 [ref. 34440], Kells et al. 2016:182 [ref. 35888], Dyldin & Orlov 2017:65 [ref. 35138], Mecklenburg et al. 2018:213 [ref. 36370], Dyldin et al. 2021:66 [ref. 38804], Love et al. 2021:123 [ref. 39279], Dyldin & Orlov 2022:48 [ref. 38972]). Current status: Valid as Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific and adjacent Arctic: northern Sea of Japan and Sea of Okhotsk north to Bering Sea and Chukchi Sea, east to eastern Gulf of Alaska (U.S.A.). Habitat: brackish, marine.

kallavesensis, Cottus quadricornis race Lönnberg [E.] 1933:7, Pl. 1 (fig. 1) [Arkiv för Zoologi v. 24 A (pt 3) (no. 12) [1932]; ref. 22149] Lake Kallavesi; Lake Niinivesi in Rautalampi; Lake Virmavesi in Karttula, Lake Päijänne basin, Finland. Syntypes: MZH (9). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies onegensis Berg & Popov 1932 -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

krawchukii, Myoxocephalus quadricornis Mikhalev [Y. V.] 1962:138 [Not found yet; ref. 40286] Reference not traced. •Probably a synonym of Myoxocephalus quadricornis (Linnaeus 1758). Current status: Uncertain as Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish.

labradoricus, Acanthocottus Girard [C. F.] in Storer 1850:247, Pl. 7 (fig. 3) [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Lost. No types known. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1130 [ref. 20662]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Neelov 1973:599 [ref. 7219], Neelov 1979:122 [ref. 3152]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

latifrons, Cottus Gratzianov [V. I.] 1904:195, Pl. 11 [Zapiski Imperatorskago russkago geograficheskago obshchestva po obshchei geografii v. 41 (no. 1); ref. 21411] Western coast Kanin Peninsula, northern Russia, Barents Sea. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies labradoricus (Girard 1850) -- (Berg 1949:1130 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952], Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Neelov 1973:597 [ref. 7219], Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

lonnbergi, Cottus quadricornis Lönnberg [E.] 1919:167, 171 [Fauna och Flora (Uppsala) v. 14 (no. 5); ref. 23061] Lake Mälaren, Sweden. Syntypes: ?NRM. Type catalog: Sideleva & Zhidkov 2021:240 [ref. 38705]. Name should not be changed since it is Swedish (see Koerber 2009:63 [ref. 30671] for spelling of Lönnberg and origin of his name). •Validates the infrasubspecific Myoxocephalus quadricornis morpha lonnbergi Berg 1916. •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Sideleva & Zhidkov 2021:240 [ref. 38705]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

lonnbergi, Myoxocephalus quadricornis morpha Berg [L. S.] 1916:431, Fig. 318 [Les Poissons des eaux douces de la Russie; ref. 277] Lake Ladoga, Russia; lake Mälaren, Sweden. No types known. Original as lönnbergi; should not be changed to loennbergi since the name is not German, but Swedish (see Koerber 2009:63 [ref. 30671] for spelling of Lönnberg and origin of his name.). Not available, as a morpha, infrasubspecific; dates to Lönnberg 1919 or to a later publication by Berg. •In the synonymy of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies as described -- (Berg 1949:1133 [ref. 20662]). •In the synonymy of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Infrasubspecific. Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

marmoratus, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:497 [Histoire naturelle des poissons; ref. 1000] Kamchatka, Russia. Type catalog: Blanc & Hureau 1968:61 [ref. 20738]. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

matsubarai, Myoxocephalus Watanabe [M.] 1958:289, Figs. 117-118, Pl. 20 [Studies of Japanese cottid fishes; ref. 21357] Funka Bay, Hokkaidô, Japan, depth 35 fathoms. Holotype (unique): not researched. Spelled matsubari on plate caption. Also described as new in Watanabe 1960:80, Pl. 26 (fig. 2) [ref. 11959]. •Valid as Myoxocephalus matsubarai Watanabe 1958 -- (Yabe in Masuda et al. 1984:327 [ref. 6441]). •Synonym of Myxocephalus jaok (Cuvier 1829) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Distribution: Western North Pacific. Habitat: marine.

mertensii, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:496 [Histoire naturelle des poissons; ref. 1000] Petropaulski, Kamchatka. Type catalog: Blanc & Hureau 1968:61 [ref. 20738]. •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neelov 1979:122 [ref. 3152], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Habitat: marine.

mitchilli, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:188 [Histoire naturelle des poissons v. 4; ref. 998] New York, U.S.A. •Synonym of Myoxocephalus aenaeus (Mitchill 1814) -- (Scott & Scott 1988:502 [ref. 25518]). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

mucosus, Acanthocottus Ayres [W. O.] 1854:194 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] New England coast, U.S.A. No types known. Appeared first as above on 13 Oct.; then in Ayres 1854:12 [ref. 18923]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

niger, Cottus Bean [T. H.] 1881:151 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Saint Paul Island, Pribilof Group, Bering Sea, Alaska, U.S.A. Lectotype: USNM 27971. Paralectotypes: MCZ 36021 [ex USNM 27952] (1); USNM 23929 (1), 27952 (orig. 12, now 1). Lectotype established (as figured specimen) in caption to Pl. 298, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. Objectively invalid; preoccupied by Cottus niger Lacepède 1801 but we recommend suppressing that name that has not been used in 100 years and its status is uncertain. •Valid as Myoxocephalus niger (Bean 1881) -- (Watanabe 1960:78 [ref. 11959], Neelov 1979:123 [ref. 3152], Sheiko 1983:18 [ref. 37041], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:213 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:476 [ref. 25968], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:76 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:90 [ref. 37547], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:262 [ref. 33547], Kells et al. 2016:182 [ref. 35888], Love et al. 2021:123 [ref. 39279], Dyldin & Orlov 2022:49 [ref. 38972] with question). Current status: Valid as Myoxocephalus niger (Bean 1881). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific: northern Sea of Japan and Sea of Okhotsk north to southern Bering Sea, east to western Gulf of Alaska (U.S.A.). Habitat: marine.

nigricans, Cottus Lesueur [C. A.] in Vaillant 1896:26 [12], Pl. 19 (unedited) [Bulletin de la Société philomathique de Paris (8th Série) v. 8 (no. 1); ref. 13482] No locality. On p. 12 of separate. •Synonym of Myoxocephalus aenaeus (Mitchill 1814). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

nivosus, Cottus Herzenstein [S. M.] 1890:113 [23] [Mélanges Biologiques, tirés du Bulletin physico-mathématique de l'Académie Impériale des Sciences de St. Pétersbourg v. 13; ref. 2149] St. Olga Bay, Russia, Sea of Japan. Holotype (unique): ZIN 8719. On p. 23 of Bull. Acad. Imp. Sci. St. Petersbourg (n. s.) v. 2. •Synonym of Myoxocephalus brandtii (Steindachner 1867) -- (Neelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:210 [ref. 15964] as brandti, Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:260 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Psychrolutidae: Myoxocephalinae. Habitat: marine.

ocellatus, Acanthocottus Storer [H. R.] 1850:253 [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Entrance to the Gut of Canso, Burial Island in Wilmot Harbor, Nova Scotia, Canada. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

ochotensis, Myoxocephalus niger var. Taranetz [A. Ya.] 1937:115 [Izvestiia Tikhookeanskogo Nauchno-Issledovatel'skogo Instituta Rybnogo Khoziaistva i Okeanografii [Proceedings, Pacific Research Institute of Fisheries and Oceanography, TINRO, Moskva] v. 11; ref. 13384] Holotype: ZIN 25468. Objectively invalid; preoccupied by Myoxocephalus verrucosus subspecies ochotensis Schmidt 1929. •Synonym of Myoxocephalus aeneus (Mitchill 1814) -- (Neelov 1979:123 [ref. 3152]). •Synonym of Myoxocephalus niger (Bean 1881) -- (Lindberg & Krasyukova 1987:213 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547]). 1°Homonym. Current status: Synonym of Myoxocephalus niger (Bean 1881). Psychrolutidae: Myoxocephalinae. Habitat: marine.

ochotensis, Myoxocephalus verrucosus Schmidt [P. Yu.] 1929:417, Fig. 3 [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 30 (no. 3); ref. 16059] Penshinskaya Guba, Shantarsky Islands (Russia), Sea of Okhotsk; Karaginsky Island (Russia), western Bering Sea. •Valid as Myoxocephalus ochotensis Schmidt 1929 -- (Neelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:76 [ref. 27579], Parin et al. 2014:262 [ref. 33547], Mecklenburg et al. 2016:85 [ref. 34440], Dyldin & Orlov 2022:49 [ref. 38972] with question). Current status: Valid as Myoxocephalus ochotensis Schmidt 1929. Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific: Sea of Okhotsk north to western Bering Sea. Habitat: marine.

octodecemspinosus, Cottus Mitchill [S. L.] 1814:7 [Report, in part, of Samuel L. Mitchill; ref. 3030] New York, U.S.A. No types known. Also in Mitchill 1815:380 [ref. 13292]. Originally as 18-spinosus; correction mandatory; as octodecem-spinosus in Mitchill 1815:380 [ref. 13292]. •Valid as Cottus octodecemspinosus Mitchill 1814 -- (Herrera 1896:63 [ref. 39336] as octodecimspinosus). •Possibly valid as Megalocottus octodecemspinosus (Mitchill 1814) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus octodecemspinosus (Mitchill 1814) -- (Neelov 1979:123 [ref. 3152], Smith 1986:450 [ref. 22186], Robins & Ray 1986:282 [ref. 23100] as octodecimspinosus, Scott & Scott 1988:502 [ref. 25518], McAllister 1990:200 [ref. 14674], Murdy et al. 1997:147 [ref. 23144], Klein-MacPhee in Collette & Klein-MacPhee 2002:351 [ref. 26158], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Afonso et al. 2016:46, 226 [ref. 36199], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Myoxocephalus octodecemspinosus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Atlantic. Habitat: brackish, marine.

oernensis, Cottus quadricornis Lönnberg [E.] 1932:16, Figs. 6 a-b [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Örn, Sweden. Syntypes: (9) ?BMNH 1932.5.17.1 (1), NRM 20323-25 (1, 5, 2). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

okadai, Porocottus Watanabe [M.] 1958:301, Figs. 126, 127, Pl. 22 [Studies of Japanese cottid fishes; ref. 21357] Off Date, Hokkaido, Japan, depth 30 fathoms. Holotype (unique): not researched. Also described as new in Watanabe 1960:90, Pl. 27 (fig. 2) [ref. 11959]. •Valid as Porocottus okadai Watanabe 1958 -- (Watanabe 1960:90 [ref. 11959]). •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

onegensis, Myoxocephalus quadricornis Berg [L. S.] & Popov [A. A.] 1932:158 [Doklady Akademii Nauk SSSR 1932; ref. 16359] Onega Lake and Petrozavodsk, Onega Lake, Russia. Syntypes: ZIN 3842 (1), 9026a (1). Type catalog: Sideleva & Zhidkov 2021:241 [ref. 38705]. Based on Berg 1916:432, fig. 319 and on specimens. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Sideleva & Zhidkov 2021:241 [ref. 38705]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

ontariensis, Triglopsis Jordan [D. S.] & Thompson [W. F.] 1910:75, Fig. 1 [Proceedings of the United States National Museum v. 38 (no. 1728); ref. 10651] Lake Ontario (deep water), off Toronto, Canada. Holotype (unique): USNM 64599. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Scott & Crossman 1998:846 [ref. 24570]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

pachypus, Cottus Günther [A.] 1860:161 [Catalogue of the fishes in the British Museum v. 2; ref. 1963] Port Leopold [Qikiqtaaluk, Somerset Island, Nunavut, Canada]. Syntypes: (2) BMNH 1855.9.19.48 (1), 1855.9.19.761 (1). •Synonym of Myoxocephalus scorpioides (Fabricius 1780) -- (Jordan Evermann 1898:1973 [ref. 2444]). Current status: Synonym of Myoxocephalus scorpioides (Fabricius 1780). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

polaris, Cottus Sabine [E.] 1824:213 [ccxiii] [A supplement to the appendix of Captain Parry's voyage; ref. 18649] North Georgia, east side Boothia Peninsula, Northwest Territories, Canada. No types known. •Synonym of Triglopsis quadricornis (Linnaeus 1758), but a valid subspecies -- (Chereshnev 1996:606 [ref. 24659]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Lee et al. 1980:826 [ref. 22416], Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

polyacanthocephalus, Cottus Pallas [P. S.] 1814:133 [Zoographia Rosso-Asiatica v. 3; ref. 3351] No definite locality [= Kodiak, Aleutian Islands]. Holotype (unique): ZMB 4757 (dry). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. Type information: Svetovidov 1978:27 [ref. 6025]. •Valid as Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Watanabe 1960:66 [ref. 11959], Neelov 1979:121 [ref. 3152], Eschmeyer & Herald 1983:176 [ref. 9277], Yabe in Masuda et al. 1984:326 [ref. 6441], Lindberg & Krasyukova 1987:210 [ref. 15964], McAllister 1990:200 [ref. 14674], Sokolovsky & Sokolovskaya 1997:57 [ref. 24691], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:482 [ref. 25968], Nakabo 2002:645 [ref. 26001], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:77 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Mecklenburg et al. 2007:176 [ref. 29448], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:261 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Pietsch & Orr 2015:43 [ref. 34694], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:78 [ref. 34440], Kells et al. 2016:184 [ref. 35888], Dyldin & Orlov 2017:66 [ref. 35138], Mecklenburg et al. 2018:215 [ref. 36370], Kim et al. 2020:103 [ref. 39118], Dyldin et al. 2021:66 [ref. 38804], Love et al. 2021:123 [ref. 39279], Dyldin & Orlov 2022:49 [ref. 38972]). Current status: Valid as Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific and adjacent Arctic: eastern Sea of Japan and sea of Okhotsk north to southern Bering Sea, then south to Puget Sound, Washington (U.S.A.). Habitat: marine.

porosus, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:498 [Histoire naturelle des poissons; ref. 1000] Baffin Bay, between western Greenland and Baffin Island. Holotype (unique): MNHN 0000-6761. Type catalog: Blanc & Hureau 1968:51 [ref. 20738]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

pygmaeus, Cottus quadricornis Lönnberg [E.] 1933:9, Pl. 1 (fig. 2) [Arkiv för Zoologi v. 24 A (pt 3) (no. 12) [1932]; ref. 22149] Lake Puruvesi, Finland. Syntypes: MZH. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies onegensis Berg & Popov 1932 -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

quadricornis, Cottus Linnaeus [C.] 1758:264 [Systema Naturae, Ed. X v. 1; ref. 2787] Baltic Sea. Syntypes: NRM 2491-92 (1, 1). Type catalog: Fernholm & Wheeler 1983:235-236 [ref. 20707] who suggest that the largest syntype would make the best lectotype. •Valid as Triglopsis quadricornis (Linnaeus 1758) -- (Neelov 1979:133 [ref. 3152], Fedorov in Whitehead et al. 1986:1260 [ref. 13677], Kottelat 1997:170 [ref. 22952], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:154 [ref. 23595], Novikov et al. 2000:161 [ref. 40929], Bogutskaya et al. 2001:46 [ref. 26178], Parin et al. 2002:S89 [ref. 28300], Bogutskaya & Naseka 2004:191 [ref. 28183], Fricke 2007:29 [ref. 30577], Kottelat & Freyhof 2007:519 [ref. 29996], Sideleva et al. 2022:3 [ref. 40100]). •Possibly valid as Megalocottus quadricornis (Linnaeus 1758) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus quadricornis (Linnaeus 1758) -- (Berg 1949:1128 [ref. 20662], Neelov 1973:597 [ref. 7219], Lee et al. 1980:826 [ref. 22416], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:504 [ref. 25518], McAllister 1990:200 [ref. 14674], Page & Burr 1991:245 [ref. 18983], Scott & Crossman 1998:842 [ref. 24570], Mecklenburg et al. 2002:477 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Scharpf 2008:11 [ref. 30399], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212], Page & Burr 2011:468 [ref. 31215], Page et al. 2013:124 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:263 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:46, 227 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:80 [ref. 34440], Mecklenburg et al. 2018:217 [ref. 36370], Love et al. 2021:123 [ref. 39279]). Current status: Valid as Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Distribution: Circumpolar in Arctic Ocean, in Pacific and north Atlantic. Habitat: freshwater, brackish, marine.

raninus, Myoxocephalus Jordan [D. S.] & Starks [E. C.] 1904:277, Fig. 20 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Aomori, Japan. Holotype: CAS-SU 7708. Paratypes: ?CAS 56330 [ex IU 10595] (1); USNM 50920 (13); BMNH 1905.2.4.373-374 (2). Additional material: CAS-SU 7591 (15, missing), 7592 (5), 7595 (5), 7938 (now 83); USNM 51285-86 (1, 1), 23815 [ex Iowa Mus.] (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Watanabe 1960:82 [ref. 11959], Neelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:207 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:480 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

relicta, Cottus quadricornis var. Lilljeborg [N.] 1891:149 [Sveriges och Norges fauna. Fiskarne. v. 1; ref. 21766] Lake Vättern, Sweden. No types known. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies relictus (Lilljeborg 1891) -- (Berg 1949:1133 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

scorpioides, Cottus Fabricius [O.] 1780:157 [Fauna groenlandica, systematice sistens animalia Groenlandiae occidentalis, etc.; ref. 17464] Western Greenland. No types known. •Possibly valid as Megalocottus scorpioides (Fabricius 1780) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus scorpioides (Fabricius 1780) -- (Neelov 1973:599 [ref. 7219], Neelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1254 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:505 [ref. 25518], McAllister 1990:200 [ref. 14674], Yabe in Okamura et al. 1995:165 [ref. 22531], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:479 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:264 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:49, 228 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:82 [ref. 34440], Mecklenburg et al. 2018:219 [ref. 36370], Love et al. 2021:123 [ref. 39279]). Current status: Valid as Myoxocephalus scorpioides (Fabricius 1780). Psychrolutidae: Myoxocephalinae. Distribution: Arctic Ocean, northwestern Atlantic and North Pacific: eastern Siberian and Chukchi seas to northern Bering Sea, Beaufort Sea to Greenland and Gulf of St. Lawrence. Habitat: brackish, marine.

scorpius, Cottus Linnaeus [C.] 1758:265 [Systema Naturae, Ed. X v. 1; ref. 2787] European Ocean. Syntypes: BMNH 1853.11.12.17 [Gronovius coll.] (1, skin), LS 17 (left half-skin plus parts), NRM 2357 (2). Type catalog: Goode & Bean 1885:196 [ref. 1841], Wheeler 1958:240 [ref. 13434], Fernholm & Wheeler 1983:234-235 [ref. 20707], Wheeler 1985:46 [ref. 20705]. •Valid as Myoxocephalus scorpius (Linnaeus 1758) -- (Neelov 1973:598 [ref. 7219], Neelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1254 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:507 [ref. 25518], Matarese et al. 1989:402 [ref. 26885], McAllister 1990:200 [ref. 14674], Yabe in Okamura et al. 1995:165 [ref. 22531], Klein-MacPhee in Collette & Klein-MacPhee 2002:353 [ref. 26158], Mecklenburg et al. 2002:478 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Fricke 2007:29 [ref. 30577], Mecklenburg et al. 2007:176 [ref. 29448], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212] with comments. Page et al. 2013:124 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:264 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:49, 229 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:84 [ref. 34440], Mecklenburg et al. 2018:221 [ref. 36370], Carneiro et al. 2019:237 [ref. 37250], Love et al. 2021:123 [ref. 39279], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Distribution: Circumpolar, northern Hemisphere. Habitat: brackish, marine.

sinensis, Cottus (Aspicottus) Sauvage [H.-E.] 1873:55 [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 9 (pt 2, Bulletin); ref. 18661] China. Holotype (unique): MNHN 0000-5265. Type catalog: Blanc & Hureau 1968:52 [ref. 20738]. •Valid as Myoxocephalus sinensis (Sauvage 1873) -- (Zhu 1995:189 [ref. 25213]). Current status: Valid as Myoxocephalus sinensis (Sauvage 1873). Psychrolutidae: Myoxocephalinae. Distribution: China. Habitat: marine.

spinosus, Porocottus Watanabe [M.] 1958:311, Figs. 132-133, Pl. 24 [Studies of Japanese cottid fishes; ref. 21357] Kurshiro, Hokaidô, Japan, depth 35 fathoms. Syntypes: (3). Also described as new in Watanabe 1960:98, Pl. 3 (fig. 2) [ref. 11959]. •Valid as Porocottus spinosus Watanabe 1958 -- (Watanabe 1960:98 [ref. 11959]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Habitat: marine.

stelleri, Myoxocephalus Tilesius [W. G. von] 1811:273, Pl. 13 (fig. 1) [Mémoires de l'Académie Impériale des Sciences de Sankt Pétersbourg (5e Série) v. 3; ref. 4408] Kamchatka, Russia. No types known. •Valid as Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662], Neelov 1979:121 [ref. 3152], Yabe in Masuda et al. 1984:326 [ref. 6441], Lindberg & Krasyukova 1987:207 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Kim 1997:407 [ref. 24032], Sokolovsky & Sokolovskaya 1997:58 [ref. 24691], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:153 [ref. 23595], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Pietsch et al. 2001:145 [ref. 25688], Zemnukhov et al. 2001:420 [ref. 25412], Kim & Park 2002:327 [ref. 25913], Mecklenburg et al. 2002:480 [ref. 25968], Nakabo 2002:645 [ref. 26001], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S82 [ref. 28300], Hagiwara 2003:37 [ref. 27188], Fedorov et al. 2003:77 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Bogutskaya & Naseka 2004:191 [ref. 28183], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:265 [ref. 33547], Kells et al. 2016:186 [ref. 35888], Dyldin & Orlov 2017:66 [ref. 35138], Kim et al. 2020:103 [ref. 39118], Dyldin et al. 2021:67 [ref. 38804], Love et al. 2021:124 [ref. 39279], Dyldin & Orlov 2022:49 [ref. 38972]). Current status: Valid as Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Distribution: North Pacific: Sea of Japan (Korea, Japan, Russia) and Sea of Okhotsk north to Commander Islands (Russia) and Aleutian Islands (U.S.A.), then south to southeastern Alaska (U.S.A.). Habitat: brackish, marine.

stimpsonii, Triglopsis Hoy [P. R.] 1872:99 [Transactions of the Wisconsin Academy of Sciences Arts and Letters 1870-72; ref. 18119] Lake Michigan, off Racine, Milwaukee, U.S.A. (stomach content). Holotype (unique): whereabouts unknown. Apparently not available; appeared above and also as Gill (MS) in Jordan 1878:50 [ref. 10444] as stimpsoni, both without distingusihing features. Subsequent availability not researched. •In the synonymy of Myoxocephalus quadricornis (Linnaeus 1758) -- (Scott & Crossman 1998:846 [ref. 24570]). Nomen Nudum. Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae.

thompsonii, Triglopsis Girard [C. F.] 1851:19 [Proceedings of the Boston Society of Natural History v. 4 (1851-1854); ref. 1816] Lake Ontario off Oswego, New York, U.S.A. Syntypes: MCZ 35979 [ex USNM 3175] (1), USNM 3175 (8, poor condition). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Scott & Crossman 1998:843 [ref. 24570], Goto et al. 2014: [6] [ref. 33456]). •Valid as Myoxocephalus thompsonii (Girard 1851) -- (Lee et al. 1980:827 [ref. 22416], Smith 1986:359 [ref. 22186], McAllister 1990:200 [ref. 14674] as thompsoni, Page & Burr 1991:244 [ref. 18983] as thompsoni, Bailey et al. 2004:173 [ref. 27882], Nelson et al. 2004:121 [ref. 27807], Murray et al. 2004:642 [ref. 27932], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:467 [ref. 31215], Page et al. 2013:124 [ref. 32708]). Current status: Valid as Myoxocephalus thompsonii (Girard 1851). Psychrolutidae: Myoxocephalinae. Distribution: Canada and New York (U.S.A.). Habitat: freshwater.

tuberculatus, Myoxocephalus Soldatov [V. K.] & Pavlenko [M. N.] 1922:342, Fig. 1 [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23; ref. 15431] Many localities, near Shantarskiye [Shantar] Islands, western Okhotsk Sea, depth 10-87 meters. Lectotype: ZIN 20615. Paralectotypes: (26) ZIN 19430 (6), 20614 (2, lost), 20615-24 (2, 4, 1, 2, 4, 2, 2, 2, 1, 1). Lectotype selected by Neelov 1979:122 [ref. 3152]. •Valid as Myoxocephalus tuberculatus Soldatov & Pavlenko 1922 -- (Parin et al. 2014:265 [ref. 33547], Dyldin et al. 2020:114 [ref. 37569], Dyldin et al. 2021:67 [ref. 38804], Dyldin & Orlov 2022:50 [ref. 38972]). Current status: Valid as Myoxocephalus tuberculatus Soldatov & Pavlenko 1922. Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific: northern Okhotsk Sea, including Shantar Islands, Taui Bay and western Kamchatka (Russia) Habitat: brackish, marine.

vaenernensis, Cottus quadricornis Lönnberg [E.] 1932:8, Fig. 3 [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Vänern, Sweden. Syntypes: NRM 20336-38 (1, 4, 1), 40338 (1). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

variabilis, Cottus Ayres [W. O.] 1842:68 [Proceedings of the Boston Society of Natural History v. 1 (1841-1844); ref. 15926] Brookhaven, Long Island, New York, U.S.A. Appeared first as name only in Ayres 1842:58 [ref. 15927]. Also in Ayres 1842:259 [ref. 15930]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

vermelnensis, Cottus quadricornis Nybelin [O.] 1940:77, Fig. 1 [Fauna och Flora (Uppsala) v. 35 (no. 2); ref. 23062] Lake Värmeln, Sweden. Syntypes: whereabouts unknown. •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

verrucosus, Cottus Bean [T. H.] 1881:152 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Plover Bay, anchorage in Providence Bay, Chukchi Peninsula, northeastern Siberia, Russia, Bering Sea. Holotype (unique): USNM 27547. Non-types: MNHN 1896-0467 (1) station 3232. Type catalog: Blanc & Hureau 1968:52 [ref. 20738]. See nomenclatural comments in Mecklenburg et al. 2007:175-176 [ref. 29448]. •Valid as Myoxocephalus verrucosus (Bean 1881) -- (Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S83 [ref. 28300], Fedorov et al. 2003:78 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Parin et al. 2014:266 [ref. 33547], Chernova et al. 2022:9 [ref. 39177]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Mecklenburg et al. 2002:405 [ref. 25968], Mecklenburg et al. 2007:175 [ref. 29448], Mecklenburg et al. 2011:123 [ref. 31212]). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific and adjacent Arctic Ocean (if valid). Habitat: marine.

virginianus, Cottus Cuvier [G.] (ex Willoughby) 1829:163 [Le Règne Animal (Edition 2) v. 2; ref. 995] Available from footnote as, "(1) C. virginianus, Will. x, 15, ou octodecim spinosus, Mitchill. Trans. New-York, iv, p. 380." •Synonym of Myoxocephalus octodecemspinosus (Mitchill 1814). Current status: Synonym of Myoxocephalus octodecemspinosus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

yesoensis, Myoxocephalus Snyder [J. O.] 1911:538 [Proceedings of the United States National Museum v. 40 (no. 1836); ref. 4152] Hakodate market, Hokkaido, Japan. Holotype (unique): USNM 68238. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662]). •Valid as Myoxocephalus yesoensis Snyder 1911 -- (Watanabe 1960:70 [ref. 11959]). •Synonym of Myxocephalus brandtii (Steindachner 1867) -- (Lindberg & Krasyukova 1987:210 [ref. 15964] as yessoensis and brandti, Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Psychrolutidae: Myoxocephalinae. Habitat: marine.