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Eschmeyer's Catalog of Fishes
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Species in the genus Pangio: [ 46 ] records


agmus, Acanthophthalmus Burridge [M. E.] 1992:180, Fig. 6 [Copeia 1992 (no. 1); ref. 19452] Sungai Lawa, 1 kilometer upstream of confluence with Baram River, Fourth Division, Sarawak state, Borneo, East Malaysia. Holotype: ROM 58726. Paratypes: BMNH 1990.5.2.1-6 (6); CAS 69586 (6); FMNH 68736 (1); IRSNB 19799 [not 19869] (now 8); MNHN 1990-0261 (6); RMNH 31657 (6); ROM 52847 (87), 53231-35 (12, 12, 1, 23, 296), 53495 (2), 56803 (5); SMF 803 (2); MZUSP [ex IRSNB 799] (2); UMMZ 217310 (6); USNM 308575 (6). •Valid as Pangio agma (Burridge 1992) -- (Kottelat et al. 1993:60 [ref. 23448], Kottelat & Lim 1996:237 [ref. 23210], Kottelat 2012:37 [ref. 32367], Kottelat 2013:182 [ref. 32989], Sulaiman et al. 2018:30 [ref. 35802]). Current status: Valid as Pangio agma (Burridge 1992). Cobitidae. Distribution: Sarawak and Brunei, Borneo. Habitat: freshwater.

alcoides, Pangio Kottelat [M.] & Lim [K. K. P.] 1993:208, Fig. 1 [Raffles Bulletin of Zoology v. 41 (no. 2); ref. 21437] Stream at 154 stone on road to Kuantan from Kuala Terengganu, Rantau Abang, Terengganu, Malaysia. Holotype: ZRC 17209. Paratypes: CMK 8242 (1), 8511 (1); ZRC 17210-14 (5), 24690 (1), 34914 (1). •Valid as Pangio alcoides Kottelat & Lim 1993 -- (Doi 1997:15 [ref. 22832], Kottelat 2012:37 [ref. 32367], Kottelat 2013:182 [ref. 32989], Anoop et al. 2019:149 [ref. 37005]). Current status: Valid as Pangio alcoides Kottelat & Lim 1993. Cobitidae. Distribution: Malaysia. Habitat: freshwater.

alternans, Pangio Kottelat [M.] & Lim [K. K. P.] 1993:210, Fig. 3 [Raffles Bulletin of Zoology v. 41 (no. 2); ref. 21437] Blackwater stream entering the Mahakam River on the left side near Mujub, Kalimantan Timur, Borneo, Indonesia, 0°01'S, 115°43'E. Holotype: MZB 5895. Paratypes: CMK 7737 (7), 7757 (6), 7782 (24); MZB 5896 (10); ZRC 35037-46 (10). •Valid as Pangio alternans Kottelat & Lim 1993 -- (Doi 1997:15 [ref. 22832], Tan & Kottelat 2009:59 [ref. 30261], Kottelat 2012:38 [ref. 32367], Kottelat 2013:182 [ref. 32989]). Current status: Valid as Pangio alternans Kottelat & Lim 1993. Cobitidae. Distribution: Mahakam River drainage, Borneo, Indonesia. Habitat: freshwater.

ammophila, Pangio Britz [R.], Ali [A.] & Raghavan [R.] 2012:46, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 23 (no. 1); ref. 32066] Kumaradhara River, 12°00'N, 75°53'24"E, Subramanya, Karnataka, India, elevation 811 meters. Holotype: CRG-SAC 2012.1.1. Paratypes: CRG-SAC 2012.1.2-3 (2). •Valid as Pangio ammophila Britz, Ali & Raghavan 2012 -- (Kottelat 2012:38 [ref. 32367], Bleher 2018:318 [ref. 36647], Anoop et al. 2019:149 [ref. 37005], Sundar et al. 2022:95 [ref. 39262]). Current status: Valid as Pangio ammophila Britz, Ali & Raghavan 2012. Cobitidae. Distribution: South Asia: Kumaradhara River drainage, southern West Ghats, India. Habitat: freshwater.

anguillaris, Acantophthalmus Vaillant [L. L.] 1902:151, Fig. 46 [Notes from the Leyden Museum v. 24 (note 1); ref. 4490] Kapuas River, Borneo, Indonesia. Holotype (unique): RMNH 7788. •Valid as Acantophthalmus anguillaris Vaillant 1902 -- (Roberts 1989:95 [ref. 6439]). •Valid as Pangio anguillaris (Vaillant 1902) -- (Kottelat et al. 1993:60 [ref. 23448], Kottelat & Lim 1993:214 [ref. 21437], Rainboth 1996:137 [ref. 22772], Doi 1997:15 [ref. 22832], Kottelat 1998:100 [ref. 23436], Kottelat 2001:90 [ref. 25780], Kottelat & Widjanarti 2005:158 [ref. 28284], Parenti & Lim 2005:187 [ref. 28285], Tan & Kottelat 2009:60 [ref. 30261], Hadiaty & Kottelat 2009:66 [ref. 30302], Kottelat 2012:38 [ref. 32367], Valis'eva & Vasil'ev 2012:665 [ref. 32477], Kottelat 2013:182 [ref. 32989], Kottelat 2015:45 [ref. 33971] as aff. anguillaris, Ng et al. 2019:520 [ref. 36606], Anoop et al. 2019:149 [ref. 37005]). Current status: Valid as Pangio anguillaris (Vaillant 1902). Cobitidae. Distribution: Borneo, ?Malaya (Malaysia), ?Mekong River basin, ?Xe Bangfai, Laos. Habitat: freshwater.

apoda, Pangio Britz [R.] & Maclaine [J. S.] 2007:28, Fig. 8 [Ichthyological Exploration of Freshwaters v. 18 (no. 1); ref. 29149] Tista River at Tista Barrage, Brahmaputra drainage, West Bengal, India. Holotype: UMMZ 247779. Paratypes: BMNH 1932.4.22.1 (1), UMMZ 244848 (1). •Valid as Pangio apoda Britz & Maclaine 2007 -- (Britz et al. 2012:45 [ref. 32066], Kottelat 2012:38 [ref. 32367], Anoop et al. 2019:149 [ref. 37005]). Current status: Valid as Pangio apoda Britz & Maclaine 2007. Cobitidae. Distribution: South Asia: Tista River drainage, Brahmaputra basin, West Bengal, India. Habitat: freshwater.

atactos, Pangio Tan [H. H.] & Kottelat [M.] 2009:57, Fig. 42 [Ichthyological Exploration of Freshwaters v. 20 (no. 1); ref. 30261] Danau Kamining near ampung Trasos, about 5 kilometers to southwards on unpaved road turning-off from Muara Bungo - Muara Tebo road at Kilometer 36, Jambi, Sumatra. Holotype: MZB 10995. Paratypes: CMK 11132 (5); MZB 10996 (2); ZRC 42421 (2), 51763 (5), 51764 (2). Name is an adjective. •Valid as Pangio atactos Tan & Kottelat 2009 -- (Britz & Kottelat 2010:375 [ref. 30691], Kottelat 2012:38 [ref. 32367], Kottelat 2013:182 [ref. 32989]). Current status: Valid as Pangio atactos Tan & Kottelat 2009. Cobitidae. Distribution: Batang Hari River drainage, Sumatra, Indonesia. Habitat: freshwater.

bashai, Pangio Easa [P. S.] & Shaji [C. P.] 1997:181 [Current Science v. 73 (no. 2); ref. 30281] Kerala, India. Name only, not available. Refers to a manuscript description submitted to TFH Magazine, which was not published by 2009 according to online TFH archives. Also seen as Pangio baashai and Pangio bashaii. •Incorrectly treated as valid as Pangio bashai Easa & Shaji -- (Raju Thomas et al. 1999:479 [ref. 25441]). •A nomen nudum -- (Britz et al. 2012:48 [ref. 32066]). Nomen Nudum. Current status: Unknown. Cobitidae.

bhujia, Pangio Anoop [V. K.], Britz [R.], Arjun [C. P.], Dahanukar [N.] & Raghavan [R.] 2019:145, Figs. 1-3 [Zootaxa 4683 (no. 1); ref. 37005] Cherinjal, Kozhikode district, Kerala, India, 11°17'42"N, 75°52'7"E, elevation 40 meters. Holotype: BNHS FWF 980. Paratypes: BNHS FWF 981-990 (10), 991-996 (6). •Valid as Pangio bhujia Anoop, Britz, Arjun, Dahanukar & Raghavan 2019 -- (Sundar et al. 2022:93 [ref. 39262]). Current status: Valid as Pangio bhujia Anoop, Britz, Arjun, Dahanukar & Raghavan 2019. Cobitidae. Distribution: South Asia: aquifer systems in Kozhikode district and Indianoor, northwest Kerala, India. Habitat: freshwater.

bitaimac, Pangio Tan [H. H.] & Kottelat [M.] 2009:59, Fig. 43 [Ichthyological Exploration of Freshwaters v. 20 (no. 1); ref. 30261] Sungai Alai, Jambi, Sumatra, from aquarium fish collectors. Holotype: MZB 10997. Paratypes: CMK 11189 (3), 17813 (6), 20840 (15); MZB 10998 (15); ZRC 29161 (7), 42036 (16), 43230 (1). Plus non-type material. Name is a noun in apposition. •Valid as Pangio bitaimac Tan & Kottelat 2009 -- (Hadiaty & Kottelat 2009:68 [ref. 30302], Kottelat 2012:38 [ref. 32367], Kottelat 2013:182 [ref. 32989], Anoop et al. 2019:149 [ref. 37005]). Current status: Valid as Pangio bitaimac Tan & Kottelat 2009. Cobitidae. Distribution: Batang Hari drainage, Jambi Province, Sumatra, Indonesia. Habitat: freshwater.

borneensis, Acanthophthalmus Boulenger [G. A.] 1894:251 [Annals and Magazine of Natural History (Series 6) v. 13 (no. 75) (art. 26); ref. 535] Baram River, Sarawak state, Borneo, East Malaysia. Syntypes: (several) BMNH 1889.7.31.14 (1), MNHN 1894-0018 (1). Type catalog: Bertin & Estève 1948:91 [ref. 19574]. •Valid as Pangio borneensis (Boulenger 1894) -- (Kottelat et al. 1993:60 [ref. 23448]). •Questionably the same as Pangio shelfordii (Popta 1903) -- (Kottelat & Lim 1993:240 [ref. 21437], Kottelat & Lim 1996:237 [ref. 23210]). •Questionably the same as Pangio agma (Burridge 1992) -- (Kottelat 2012:37 [ref. 32367], Kottelat 2013:182 [ref. 32989]). Current status: Uncertain as Pangio agma (Burridge 1992). Cobitidae. Distribution: Borneo. Habitat: freshwater.

cinnamomea, Cobitis McClelland [J.] 1839:304, 435, Pl. 51 (fig. 5b) [Asiatic Researches v. 19 (pt 2); ref. 2923] Unneeded replacement name for and objective synonym of Cobitis pangia Hamilton 1822 (see Opinion 1695). •Synonym of Acanthophthalmus pangia (Hamilton 1822) -- (Menon 1992:87 [ref. 23970]). •Synonym of Pangio pangia (Hamilton 1822) -- (Menon 1999:166 [ref. 24904] as pangio, Britz & Maclaine 2007:24 [ref. 29149], Kottelat 2012:39 [ref. 32367], Kottelat 2013:183 [ref. 32989]). Replacement Name. Current status: Synonym of Pangio pangia (Hamilton 1822). Cobitidae. Habitat: freshwater.

cuneovirgatus, Acanthophthalmus Raut [H.] 1957:30, Figs. 4, 5 (lower), 6 [Die Aquarien- und Terrarienzeitschrift (DATZ) v. 10 (no. 2); ref. 12164] Johore, southern Malaysia. Holotype: ZMB 21332. Paratypes: ZMB 21333 (1). •Valid as Acanthophthalmus cuneovirgatus Raut 1957 -- (Burridge 1992:177 [ref. 19452]). •Valid as Pangio cuneovirgata (Raut 1957) -- (Kottelat et al. 1993:60 [ref. 23448], Kottelat & Lim 1993:215 [ref. 21437], Doi 1997:15 [ref. 22832], Ng & Tan 1999:356 [ref. 24159], Kottelat 2012:38 [ref. 32367], Valis'eva & Vasil'ev 2012:664 [ref. 32477], Kottelat 2013:182 [ref. 32989], Ng et al. 2019:520 [ref. 36606]). Current status: Valid as Pangio cuneovirgata (Raut 1957). Cobitidae. Distribution: Southeast Asia. Habitat: freshwater.

doriae, Eucirrhichthys Perugia [A.] 1892:1009 [1] [Annali del Museo Civico di Storia Naturale di Genova (Serie 2) v. 12; ref. 3432] Sarawak state, Borneo, East Malaysia. Syntypes: MSNG 9231 (4 or 2, apparently lost), ZMA 114898 (1). Type catalog: Tortonese 1961:188 [ref. 10343], Nijssen et al. 1982:28 [ref. 19236]. On p. 1 of separate. •Valid as Pangio doriae (Perugia 1892) -- (Kottelat et al. 1993:60 [ref. 23448], Kottelat & Lim 1993:217 [ref. 21437], Kottelat & Lim 1996:237 [ref. 23210], Doi 1997:15 [ref. 22832], Ng & Tan 1999:356 [ref. 24159], Kottelat & Widjanarti 2005:158 [ref. 28284], Tan & Kottelat 2009:60 [ref. 30261], Kottelat 2012:38 [ref. 32367], Kottelat 2013:182 [ref. 32989], Anoop et al. 2019:149 [ref. 37005]). Current status: Valid as Pangio doriae (Perugia 1892). Cobitidae. Distribution: Borneo, Malaysia. Habitat: freshwater.

elongata, Pangio Britz [R.] & Maclaine [J. S.] 2007:18, Fig. 1 [Ichthyological Exploration of Freshwaters v. 18 (no. 1); ref. 29149] Mouth of the Mingaw Kloh, Tenasserim, Myanmar. Holotype: BMNH 1992.11.16.13. Paratypes: BMNH 1893.2.16.69 (1), USNM 44774 (1). •Valid as Pangio elongata Britz & Maclaine 2007 -- (Kottelat 2012:38 [ref. 32367], Kottelat 2013:182 [ref. 32989]). Current status: Valid as Pangio elongata Britz & Maclaine 2007. Cobitidae. Distribution: Tenasserim, Myanmar. Habitat: freshwater.

fasciatus, Acanthophthalmus Bleeker [P.] 1860:74 [Acta Societatis Regiae Scientiarum Indo-Neêrlandicae v. 7 (art. 2); ref. 380] Krawang, Java, Indonesia. Neotype: RMNH 2688. Neotype designated by Kottelat & Lim 1993:224 [ref. 21437]. •Absolute synonym of Pangio kuhlii (Valenciennes 1846) -- (Kottelat & Lim 1993:224 [ref. 21437], Kottelat 2012:38 [ref. 32367], Kottelat 2013:183 [ref. 32989]). Current status: Synonym of Pangio kuhlii (Valenciennes 1846). Cobitidae. Habitat: freshwater.

fasciatus, Acanthophthalmus van Hasselt [J. C.] 1823:133 [Algemeene Konst- en Letter-bode voor het Jaar II Deel (no. 35); ref. 5963] Not available, no distinguishing features, from Java -- (see Alfred 1961:86 [ref. 20553], Kottelat 1987:371 [ref. 5962]). Also in van Hasselt 1824:377 [ref. 5964]. •In the synonymy of Acanthophthalmus kuhlii (Valenciennes 1846) -- (Burridge 1992:181 [ref. 19452]). •In the synonymy of Pangio kuhlii (Valenciennes 1846) -- (Roberts 1993:26 [ref. 21799], Kottelat & Lim 1993:224 [ref. 21437], Kottelat 2012:38 [ref. 32367], Kottelat 2013:183 [ref. 32989]). Nomen Nudum. Current status: Synonym of Pangio kuhlii (Valenciennes 1846). Cobitidae.

fasciatus, Acanthophthalmus Kuhl [H.] in Cuvier & Valenciennes 1846:77 [Histoire naturelle des poissons v. 18; ref. 1011] Not available, name mentioned in passing under Cobitis kuhlii Valenciennes 1846; from Jakarta [Batavia], Java, Indonesia. Nomen Nudum. Cobitidae.

filinaris, Pangio Kottelat [M.] & Lim [K. K. P.] 1993:219, Fig. 9 [Raffles Bulletin of Zoology v. 41 (no. 2); ref. 21437] Sungai Tersat, a tributary of Sungai Terengganu, Terengganu, Malaysia, 4°57'51"N, 102°57'45"E. Holotype: ZRC 34915. Paratypes: CMK 9756 (8); ZRC 657 (5), 9505 (1), 24550 (1), 34916-36 (21), 34937-45 (9). •Valid as Pangio filinaris Kottelat & Lim 1993 -- (Doi 1997:15 [ref. 22832], Ng & Tan 1999:356 [ref. 24159], Britz & Kottelat 2010:375 [ref. 30691], Kottelat 2012:38 [ref. 32367], Kottelat 2013:182 [ref. 32989]). Current status: Valid as Pangio filinaris Kottelat & Lim 1993. Cobitidae. Distribution: Malaysia. Habitat: freshwater.

fusca, Apua Blyth [E.] 1860:169 [Journal of the Asiatic Society of Bengal v. 29 (no. 2); ref. 477] Tenasserim, Myanmar. Syntypes: (3) ?AMS B.7500 (1), ZSI F2647/1 (2). Type catalog: Menon & Yazdani 1968:120 [ref. 20743]. •Synonym of Pangio pangia (Hamilton 1822) -- (Talwar & Jhingran 1991:531 [ref. 20764], Menon 1999:166 [ref. 24904], Ataur Rahman 2003:171 [ref. 31338]). •Synonym of Acanthophthalmus pangia (Hamilton 1822) -- (Menon 1992:88 [ref. 23970]). •Valid as Pangio fusca (Blyth 1860) -- (Kottelat 1998:100 [ref. 23436], Kottelat 2001:90 [ref. 25780], Britz & Maclaine 2007:20 [ref. 29149], Kottelat 2012:38 [ref. 32367], Valis'eva & Vasil'ev 2012:665 [ref. 32477], Kottelat 2013:182 [ref. 32989], Kottelat 2015:45 [ref. 33971] as aff. fusca). Current status: Valid as Pangio fusca (Blyth 1860). Cobitidae. Distribution: Myanmar, doubtfully Laos. Habitat: freshwater.

goaensis, Acanthophthalmus Tilak [R.] 1972:62, Figs. 1-4 [Journal of the Inland Fishery Society of India v. 4; ref. 7655] Colem River, Goa, India. Holotype (unique): ZSI F6150/2. See Das 2003:352 [ref. 27706] with date as 1973. •Valid as Acanthophthalmus goaensis Tilak 1972 -- (Tilak 1990:295 [ref. 24003], Menon 1992:91 [ref. 23970]). •Valid as Pangio goaensis (Tilak 1972) -- (Talwar & Jhingran 1991:530 [ref. 20764], Menon 1999:165 [ref. 24904], Raju Thomas et al. 1999:479 [ref. 25441], Raju Thomas et al. 2002:50 [ref. 26359], Britz et al. 2012:45 [ref. 32066], Kottelat 2012:38 [ref. 32367]). Current status: Valid as Pangio goaensis (Tilak 1972). Cobitidae. Distribution: India. Habitat: freshwater.

incognito, Pangio Kottelat [M.] & Lim [K. K. P.] 1993:221, Fig. 11 [Raffles Bulletin of Zoology v. 41 (no. 2); ref. 21437] kilometer 42 on road from Lindu to Kuching, west of Sungei Stinggang, Sarawak state, Borneo, East Malaysia. Holotype: ZRC 34913. Paratypes: CMK 9735 (2); Sarawak Mus. uncat. (2); ZRC 34910-11 (2). •Valid as Pangio incognito Kottelat & Lim 1993 -- (Kottelat & Lim 1996:237 [ref. 23210], Doi 1997:15 [ref. 22832], Britz & Kottelat 2010:375 [ref. 30691], Kottelat 2012:38 [ref. 32367], Kottelat 2013:182 [ref. 32989]). Current status: Valid as Pangio incognito Kottelat & Lim 1993. Cobitidae. Distribution: Malaysia. Habitat: freshwater.

javanicus, Acanthophthalmus Bleeker [P.] 1860:75 [Acta Societatis Regiae Scientiarum Indo-Neêrlandicae v. 7 (art. 2); ref. 380] Bogor, Java, Indonesia. Possible former syntypes in RMNH 7061 (some of 18). Unnecessary replacement name for Cobitis oblonga Valenciennes 1846. Previously mentioned as Acanthophthalmus javanicus by Hasselt 1823:133 [ref. 5963] and Hasselt 1824:377 [ref. 5964] without distinguishing characters, therefore nomina nuda. •Synonym of Pangio oblonga (Valenciennes 1846) -- (Kottelat & Lim 1993:234 [ref. 21437], Kottelat 2001:90 [ref. 25780], Kottelat 2012:39 [ref. 32367], Kottelat 2013:183 [ref. 32989]). Replacement Name. Current status: Synonym of Pangio oblonga (Valenciennes 1846). Cobitidae. Habitat: freshwater.

javanicus, Acantophthalmus van Hasselt [J. C.] 1823:133 [Algemeene Konst- en Letter-bode voor het Jaar II Deel (no. 35); ref. 5963] Not available, no distinguishing features; from Java (see Alfred 1961:86 [ref. 20553]). Also in van Hasselt 1824:377 [ref. 5964]. •In the synonymy of Pangio oblonga (Valenciennes 1846) -- (Kottelat & Lim 1993:234 [ref. 21437], Roberts 1993:26 [ref. 21799], Kottelat 2013:183 [ref. 32989]). Nomen Nudum. Current status: Synonym of Pangio oblonga (Valenciennes 1846). Cobitidae.

kuhlii, Cobitis Valenciennes [A.] in Cuvier & Valenciennes 1846:77 [Histoire naturelle des poissons v. 18; ref. 1011] Jakarta, Java, Indonesia. Neotype: RMNH 2688. Type information: Roberts 1993:26 [ref. 21799]. Neotype (RMNH 2688) designated by Burridge 1992:181 [ref. 19452], but a holotype was listed by Bertin & Estève 1948:91 [ref. 19574] (not valid). Valid as Pangio kuhlii after Acanthophthalmus was suppressed in Opinion 1695 in 1992. •Valid as Acanthophthalmus kuhlii (Valenciennes 1846) -- (Burridge 1992:181 [ref. 19452]). •Valid as Pangio kuhlii (Valenciennes 1846) -- (Kottelat et al. 1993:60 [ref. 23448], Roberts 1993:26 [ref. 21799], Kottelat & Lim 1993:224 [ref. 21437], Kottelat & Lim 1996:237 [ref. 23210], Doi 1997:15 [ref. 22832], Ng & Tan 1999:356 [ref. 24159], Fuller et al. 1999:177 [ref. 25838], Kottelat & Widjanarti 2005:158 [ref. 28284], Tan & Kottelat 2009:61 [ref. 30261], Kottelat 2012:38 [ref. 32367], Valis'eva & Vasil'ev 2012:664 [ref. 32477], Kottelat 2013:183 [ref. 32989]). Current status: Valid as Pangio kuhlii (Valenciennes 1846). Cobitidae. Distribution: Java, Sumatra, Kalimantan, Malaya and Vietnam. Habitat: freshwater.

lidi, Pangio Hadiaty [R. K.] & Kottelat [M.] 2009:65, Figs. 1-2 [Zootaxa No. 2171; ref. 30302] Mahakam River basin, Ulu Belayan, Sungai Petunk Kanan, 0°32'06"N, 116°10'55"E, eastern Borneo. Holotype: MZB 16528. Paratypes: CMK 20805 (1). •Valid as Pangio lidi Hadiaty & Kottelat 2009 -- (Kottelat 2012:38 [ref. 32367], Kottelat 2013:183 [ref. 32989], Anoop et al. 2019:149 [ref. 37005]). Current status: Valid as Pangio lidi Hadiaty & Kottelat 2009. Cobitidae. Distribution: Mahakam River basin, eastern Borneo, Indonesia. Habitat: freshwater.

longimanus, Pangio Britz [R.] & Kottelat [M.] 2010:373, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 20 (no. 4) [Dec. 2009]; ref. 30691] Confluence of Nam Leuk and Nam Gnong streams, 18°22'04"N, 103°05'27"E, Vientiane Province, Laos. Holotype: ZRC 51933. Paratypes: CMK 199692 (2). •Valid as Pangio longimanus Britz & Kottelat 2010 -- (Kottelat 2012:39 [ref. 32367], Valis'eva & Vasil'ev 2012:665 [ref. 32477], Kottelat 2013:183 [ref. 32989], Anoop et al. 2019:149 [ref. 37005]). Current status: Valid as Pangio longimanus Britz & Kottelat 2010. Cobitidae. Distribution: Mekong River drainage, Central Laos. Habitat: freshwater.

lumbriciformis, Pangio Britz [R.] & Maclaine [J. S.] 2007:22, Fig. 4 [Ichthyological Exploration of Freshwaters v. 18 (no. 1); ref. 29149] Nan Kwe stream, near Myitkina, Ayeyarwaddy basin, Myanmar. Holotype: BMNH 2006.9.29.6. Paratypes: BMNH 2006.9.29.7-10 (4). •Valid as Pangio lumbriciformis Britz & Maclaine 2007 -- (Tan & Kottelat 2009:61 [ref. 30261], Hadiaty & Kottelat 2009:68 [ref. 30302], Kottelat 2012:39 [ref. 32367], Kottelat 2013:183 [ref. 32989]). Current status: Valid as Pangio lumbriciformis Britz & Maclaine 2007. Cobitidae. Distribution: Irrawaddy basin, Myanmar. Habitat: freshwater.

malayanus, Acanthophthalmus kuhlii Tweedie [M. W. F.] 1956:58, Pl. 6a [Bulletin of the Raffles Museum No. 27; ref. 12237] Tahan River, Pahang, Malaysia. Holotype: BMNH 1957.1.23.1. Paratypes: BMNH 1957.1.23.2-3 (2), ZRC 1444 (2). Type catalog: Alfred 1970:70 [ref. 23931]. The holotype is apparently at the BMNH and not ZRC 1444 as given by Burridge 1992:185 [ref. 19452]. •Synonym of Acanthophthalmus kuhlii (Valenciennes 1846) -- (Burridge 1992:181 [ref. 19452]). •Valid as Pangio malayana (Tweedie 1956) -- (Kottelat & Lim 1993:231 [ref. 21437], Doi 1997:15 [ref. 22832], Ng & Tan 1999:357 [ref. 24159], Kottelat & Widjanarti 2005:158 [ref. 28284], Kottelat 2012:39 [ref. 32367], Valis'eva & Vasil'ev 2012:666 [ref. 32477], Kottelat 2013:183 [ref. 32989]). Current status: Valid as Pangio malayana (Tweedie 1956). Cobitidae. Distribution: Southeast Asia. Habitat: freshwater.

mariae, Acanthophthalmus Inger [R. F.] & Chin [P. K.] 1962:118, Fig. 56 [Fieldiana Zoology v. 45; ref. 12303] Sabah, tributary of Kinabatangan at Deramakot, Kinabatangan District, Malaysia. Holotype: FMNH 68161. Paratypes: BMNH 1960.12.30.6-7 [ex FMNH 68162] (2); Chin coll. [ex FMNH 68163] (5); FMNH 68162 (41, now 37), 68163 (57, now 50), 68164-65 (14, 2); RMNH 23739 [ex FMNH] (2); ZRC 941 [? ex Chin coll.] (5). Type catalog: Alfred 1970:70 [ref. 23931], Ibarra & Stewart 1987:4 [ref. 12367]. Originally as mariae but named for two women named Mary, so corrected to mariarum, now in use. •Questionably a synonym of Acantophthalmus oblongus (Valenciennes 1846) -- (Roberts 1989:98 [ref. 6439]). •Valid as Pangio mariarum (Inger & Chin 1962) -- (Kottelat et al. 1993:60 [ref. 23448], Kottelat & Lim 1996:238 [ref. 23210], Doi 1997:15 [ref. 22832], Martin-Smith & Tan 1998:589 [ref. 23647], Kottelat 2012:39 [ref. 32367], Kottelat 2013:183 [ref. 32989], Sulaiman et al. 2018:30 [ref. 35802], Wilkinson & Tan 2018:601 [ref. 36226]). Current status: Valid as Pangio mariarum (Inger & Chin 1962). Cobitidae. Distribution: Southeast Asia: Northern Borneo. Habitat: freshwater.

muraeniformis, Acanthophthalmus (Cobitophis) de Beaufort [L. F.] 1933:32 [Bulletin of the Raffles Museum No. 8; ref. 12381] Thomson Road, 3rd mile, Singapore. Syntypes: (6, orig. Raffles Mus. F.739-744) CAS-SU 32602 (1) Malaya, ZMA 103185 (3), ZRC 1052 (2). Type catalog: Böhlke 1953:39 [ref. 12291], Alfred 1970:70 [ref. 23931]. •Synonym of Pangio shelfordii (Popta 1903) -- (Kottelat & Lim 1993:240 [ref. 21437]). •Valid as Pangio muraeniformis (de Beaufort 1933) -- (Lim et al. 1990:43 [ref. 16753], Doi 1997:15 [ref. 22832], Kottelat 2012:39 [ref. 32367], Kottelat 2013:183 [ref. 32989], Ng et al. 2019:520 [ref. 36606]). Current status: Valid as Pangio muraeniformis (de Beaufort 1933). Cobitidae. Distribution: Malaya (Malaysia), Singapore. Habitat: freshwater.

myersi, Acanthophthalmus Harry [R. R.] 1949:69 [Proceedings of the Biological Society of Washington v. 62; ref. 12417] Nong Khor, southeastern Thailand. Holotype: USNM 103300. Paratypes: MCZ 37251 [ex USNM 103300] (1), CAS-SU 14888 (1), USNM 144222 (1). Type catalog: Böhlke 1953:39 [ref. 12291]. •Synonym of Acanthophthalmus kuhlii (Valenciennes 1846) -- (Burridge 1992:181 [ref. 19452]). •Valid as Pangio myersi (Harry 1949) -- (Doi 1997:15 [ref. 22832], Kottelat 2001:90 [ref. 25780], Kottelat 2012:39 [ref. 32367], Valis'eva & Vasil'ev 2012:665 [ref. 32477], Kottelat 2013:183 [ref. 32989], Taki et al. 2021:187 [ref. 39830]). Current status: Valid as Pangio myersi (Harry 1949). Cobitidae. Distribution: Southeast Asia: Mekong River basin, southeastern Thailand and western Cambodia. Habitat: freshwater.

oblonga, Cobitis Valenciennes [A.] (ex Kuhl & van Hasselt) in Cuvier & Valenciennes 1846:76 [Histoire naturelle des poissons v. 18; ref. 1011] Bogor, Java, Indonesia. Neotype: ZRC 35047. Based on a drawing sent by van Hasselt, specimen lost. Neotype designated by Kottelat & Lim 1993:234 [ref. 21437]. •Valid as Acantophthalmus oblongus (Valenciennes 1846) -- (Roberts 1989:97 [ref. 6439]). •Valid as Pangio oblonga (Valenciennes 1846) -- (Kottelat et al. 1993:60 [ref. 23448], Kottelat & Lim 1993:234 [ref. 21437], Roberts 1993:26 [ref. 21799], Rainboth 1996:138 [ref. 22772], Doi 1997:15 [ref. 22832], Kottelat 1998:100 [ref. 23436], Kottelat 2001:90 [ref. 25780], Kottelat & Widjanarti 2005:158 [ref. 28284], Britz et al. 2012:48 [ref. 32066], Kottelat 2012:39 [ref. 32367], Valis'eva & Vasil'ev 2012:665 [ref. 32477], Kottelat 2013:183 [ref. 32989], Taki et al. 2021:187 [ref. 39830]). Current status: Valid as Pangio oblonga (Valenciennes 1846). Cobitidae. Distribution: Southeast Asia: Sundaland. Habitat: freshwater.

pangia, Cobitis Hamilton [F.] 1822:355, 394 [An account of the fishes found in the river Ganges; ref. 2031] Brahmaputra River, at Gualpara [Goalpara], Assam, India. No types known. See Opinion 1695. Information on type locality see Britz 2019:46 [ref. 37532]. The original Hamilton illustration is reproduced from McClelland 1839 in Britz & Maclaine 2007:25, fig. 6 [ref. 29149].; also reproduced by Britz 2019:Pl. 221 [ref. 37532]. •Valid as Acanthophthalmus pangia (Hamilton 1822) -- (Ataur Rahman 1989:152 [ref. 24860], Menon 1992:87 [ref. 23970], Tilak 1990:295 [ref. 24003]). •Valid as Pangio pangia (Hamilton 1822) -- (Tilak 1990:55 [ref. 24001], Talwar & Jhingran 1991:531 [ref. 20764], Sen 1995:555 [ref. 23778], Doi 1997:15 [ref. 22832], Kottelat 1998:100 [ref. 23436], Menon 1999:166 [ref. 24904] as pangio, Raju Thomas et al. 1999:479 [ref. 25441], Kunda 2000:96 [ref. 25663], Karmakar 2000:31 [ref. 25662], Britz & Maclaine 2007:24 [ref. 29149], Shrestha 2008:139 [ref. 29923], Britz et al. 2012:45 [ref. 32066], Kottelat 2012:39 [ref. 32367], Ahmed et al. 2013:286 [ref. 32872], Kottelat 2013:183 [ref. 32989], Bleher 2018:320 [ref. 36647], Anoop et al. 2019:149 [ref. 37005], Shangningam & Kosygin 2022:60 [ref. 39587]). Current status: Valid as Pangio pangia (Hamilton 1822). Cobitidae. Distribution: South Asia: Ganges River basin to Chindwin River basin (Nepal, India, Bhutan, Bangladesh and Myanmar). Habitat: freshwater.

pathala, Pangio Sundar [R. L.], Arjun [C. P.], Sidharthan [A.], Dahanukar [N.] & Raghavan [R.] 2022:90, Fig. 1 [Zootaxa 5138 (no. 1); ref. 39262] Well at Thiruvanvandoor, near the town of Chengannur, Kerala, India, 9°20'23.09"N, 76°34'48.54"E, elevation 7 meters. Holotype: KUFOS FT.2020.1. Paratypes: KUFOS FT.2020.2 (1), KUFOS FT.2020.3 (1). •Valid as Pangio pathala Sundar, Arjun, Sidharthan, Dahanukar & Raghavan 2022. Current status: Valid as Pangio pathala Sundar, Arjun, Sidharthan, Dahanukar & Raghavan 2022. Cobitidae. Distribution: South Asia: aquifer systems at Thiruvanvandoor, Western Ghats, Kerala State, India. Habitat: freshwater.

perakensis, Cobitophis Herre [A. W. C. T.] 1940:8, Pl. 2 [Bulletin of the Raffles Museum No. 16; ref. 2130] Lake above Chenderoh Dam, Perak, Malaysia. Holotype: CAS-SU 33004. Paratypes: (4) BMNH 1938.12.1.126 (1), CAS-SU 33005 (3), ZRC 1442 (3). Type catalog: Böhlke 1953:39 [ref. 12291]. •Synonym of Pangio anguillaris (Vaillant 1902) -- (Kottelat & Lim 1993:214 [ref. 21437], Tan & Kottelat 2009:61 [ref. 30261], Kottelat 2012:38 [ref. 32367], Kottelat 2013:182 [ref. 32989]). Current status: Synonym of Pangio anguillaris (Vaillant 1902). Cobitidae. Habitat: freshwater.

piperata, Pangio Kottelat [M.] & Lim [K. K. P.] 1993:236, Fig. 20 [Raffles Bulletin of Zoology v. 41 (no. 2); ref. 21437] Stream at about kilometer 6 on Kuala Brang to Kuala Terengganu road, Terengganu, Malaysia, 6°04'25"N, 103°03'20"E. Holotype: ZRC 35003. Paratypes: CMK 8497 (7), 8537 (1); ZRC 493 (26), 494 (70), 495 (18), 23049-054 (6), 23074-087 (14), 28189-216 (28), 28294-317 (24). •Valid as Pangio piperata Kottelat & Lim 1993 -- (Doi 1997:15 [ref. 22832], Ng & Tan 1999:357 [ref. 24159], Kottelat 2001:93 [ref. 25780] as cf. piperata, Parenti & Lim 2005:187 [ref. 28285], Britz & Kottelat 2010:375 [ref. 30691], Kottelat 2012:39 [ref. 32367], Kottelat 2013:183 [ref. 32989]). Current status: Valid as Pangio piperata Kottelat & Lim 1993. Cobitidae. Distribution: Southeast Asia: Malaysia, Sumatra and Borneo. Habitat: freshwater.

pulla, Pangio Kottelat [M.] & Lim [K. K. P.] 1993:238, Fig. 21 [Raffles Bulletin of Zoology v. 41 (no. 2); ref. 21437] Blackwater stream, 28-29 kilometers on road to Tangkiling, tributary of Sungei Rungan, Kalimantan Tengah, Borneo, Indonesia. Holotype: ZRC 35022. Paratypes: CAS 80441 (1). •Valid as Pangio pulla Kottelat & Lim 1993 -- (Doi 1997:15 [ref. 22832], Tan & Kottelat 2009:60 [ref. 30261], Kottelat 2012:39 [ref. 32367], Kottelat 2013:184 [ref. 32989], Anoop et al. 2019:149 [ref. 37005]). Current status: Valid as Pangio pulla Kottelat & Lim 1993. Cobitidae. Distribution: Borneo, Indonesia. Habitat: freshwater.

robiginosus, Acanthophthalmus Raut [H.] 1957:31, Figs. 5 (upper), 7, 8 [Die Aquarien- und Terrarienzeitschrift (DATZ) v. 10 (no. 2); ref. 12164] Tjipaja-en near Rangkas-Betong village, Bantean area, western Java, Indonesia. Holotype: ZMB 21334. Paratypes: ZMB 21335 (1) •Synonym of Acanthophthalmus cuneovirgatus Raut 1957 -- (Burridge 1992:177 [ref. 19452]). •Questionably a synonym of Pangio cuneovirgata (Raut 1957) -- (Kottelat & Lim 1993:215 [ref. 21437]). •Valid as Pangio robiginosa (Raut 1957) -- (Doi 1997:15 [ref. 22832], Kottelat 2012:39 [ref. 32367], Kottelat 2013:184 [ref. 32989]). Current status: Valid as Pangio robiginosa (Raut 1957). Cobitidae. Distribution: Java, Indonesia. Habitat: freshwater.

rubiginosus, Canthophrys Swainson [W.] 1839:310 [The natural history and classification v. 2; ref. 4303] Based on Ham., Cob. No. 6 -- which is Hamilton 1822:355, "6th Species.-- Cobitis pangia." Unneeded new name for Cobitis pangia Hamilton 1822. •Synonym of Acanthophthalmus pangia (Hamilton 1822) -- (Menon 1992:87 [ref. 23970]). •Synonym of Pangio pangia (Hamilton 1822) -- (Menon 1999:166 [ref. 24904] as pangio, Kottelat 2012:39 [ref. 32367], Kottelat 2013:183 [ref. 32989]). Current status: Synonym of Pangio pangia (Hamilton 1822). Cobitidae. Habitat: freshwater.

semicinctus, Acanthophthalmus Fraser-Brunner [A.] 1940:172, Fig. 3 [Annals and Magazine of Natural History (Series 11) v. 6 (no. 32); ref. 17234] Mawai District, Johore, Malaysia. Holotype: BMNH 1938.12.1.113. Paratypes: (5) BMNH 1938.12.1.114-115 (2), 1940.2.8.2-3 (2). Tan & Kottelat 2009:61 [ref. 30261] give precedence to Pangio semicincta over its simultaneous synonym P. sumatrana. •Valid as Acanthophthalmus semicinctus Fraser-Brunner 1940 -- (Roberts 1989:98 [ref. 6439]). •Synonym of Acanthophthalmus kuhlii (Valenciennes 1846) -- (Burridge 1992:181 [ref. 19452]). •Questionably a synonym of Pangio kuhlii (Valenciennes 1846) -- (Kottelat & Lim 1993:224 [ref. 21437]). •Synonym of Pangio kuhlii (Valenciennes 1846) -- (Fuller et al. 1999:177 [ref. 25838]). •Valid as Pangio semicincta (Fraser-Brunner 1940) -- (Doi 1997:15 [ref. 22832], Kottelat & Widjanarti 2005:158 [ref. 28284], Parenti & Lim 2005:187 [ref. 28285], Tan & Kottelat 2009:61 [ref. 30261], Kottelat 2012:39 [ref. 32367], Kottelat 2013:184 [ref. 32989], Ng et al. 2019:520 [ref. 36606]). Current status: Valid as Pangio semicincta (Fraser-Brunner 1940). Cobitidae. Distribution: Sumatra, Borneo, Malaya (Malaysia). Habitat: freshwater.

shelfordii, Acanthophthalmus Popta [C. M. L.] 1903:231, Fig. [Notes from the Leyden Museum v. 23 (no. 4) (art. 33) (for Oct. 1901); ref. 15039] Sarawak River near Kuching, Sarawak state, Borneo, East Malaysia. Holotype (unique): RMNH 7661. •Valid as Acantophthalmus shelfordii Popta 1903 -- (Roberts 1989:98 [ref. 6439] as shelfordi). •Valid as Pangio shelfordii (Popta 1903) -- (Kottelat et al. 1993:61 [ref. 23448], Kottelat & Lim 1993:240 [ref. 21437], Kottelat & Lim 1996:238 [ref. 23210], Doi 1997:15 [ref. 22832], Ng & Tan 1999:357 [ref. 24159], Kottelat & Widjanarti 2005:159 [ref. 28284], Tan & Kottelat 2009:58 [ref. 30261], Kottelat 2012:40 [ref. 32367], Kottelat 2013:184 [ref. 32989]). Current status: Valid as Pangio shelfordii (Popta 1903). Cobitidae. Distribution: Asia. Habitat: freshwater.

signicauda, Pangio Britz [R.] & Maclaine [J. S.] 2007:26, Fig. 7 [Ichthyological Exploration of Freshwaters v. 18 (no. 1); ref. 29149] Nanmate stream, 25°23'03"N, 97°00'38"E, between Myitkina and Hopin, Ayeyarwaddy basin, Myanmar. Holotype: USNM 378386. Paratypes: BMNH 2006.9.29.1-5 (5), USNM 388185 (13). •Valid as Pangio signicauda Britz & Maclaine 2007 -- (Kottelat 2012:40 [ref. 32367], Kottelat 2013:184 [ref. 32989]). Current status: Valid as Pangio signicauda Britz & Maclaine 2007. Cobitidae. Distribution: Southern Asia: Ayeyarwaddy River basin, Myanmar. Habitat: freshwater.

sumatranus, Acanthophthalmus kuhlii Fraser-Brunner [A.] 1940:175 [Annals and Magazine of Natural History (Series 11) v. 6 (no. 32); ref. 17234] Lahat, Palembang Province, Sumatra, Indonesia. Holotype (unique): BMNH 1866.5.2.41. Tan & Kottelat 2009:61 [ref. 30261] give precedence to Pangio semicincta over its simultaneous synonym P. sumatrana. •Synonym of Acanthophthalmus kuhlii (Valenciennes 1846) -- (Burridge 1992:181 [ref. 19452]). • Questionably a synonym of Pangio kuhlii (Valenciennes 1846) -- (Kottelat & Lim 1993:224 [ref. 21437]). •Synonym of Pangio semicincta (Fraser-Brunner 1940) -- (Tan & Kottelat 2009:61 [ref. 30261], Kottelat 2012:39 [ref. 32367], Kottelat 2013:184 [ref. 32989]). Current status: Synonym of Pangio semicincta (Fraser-Brunner 1940). Cobitidae. Habitat: freshwater.

superbus, Acantophthalmus Roberts [T. R.] 1989:98, Fig. 74 [Memoirs of the California Academy of Sciences No. 14; ref. 6439] Kapuas basin, small forest stream about 1 kilometer up Sungai Tajan from Tajan, 87 kilometer east of Pontianak, 0°02"S, 110°07'E, Indonesia. Holotype: MZB 3499. Paratypes: CAS 49339-40 (3, 3); MZB 3500-01 (1, 1); USNM 230259 (2). •Valid as Pangio superba (Roberts 1989) -- (Kottelat et al. 1993:61 [ref. 23448], Kottelat & Widjanarti 2005:159 [ref. 28284], Tan & Kottelat 2009:58 [ref. 30261], Kottelat 2012:40 [ref. 32367], Kottelat 2013:184 [ref. 32989]). Current status: Valid as Pangio superba (Roberts 1989). Cobitidae. Distribution: Borneo, Indonesia. Habitat: freshwater.

vermicularis, Acanthophthalmus Weber [M.] & de Beaufort [L. F.] 1916:34 [Fishes Indo-Australian Archipelago v. 3; ref. 4604] Kampar Kiri River, Sumatra, Indonesia. Holotype (unique): ZMA 100260. Type catalog: Nijssen et al. 1982:26 [ref. 19236]. •Synonym of Acanthophthalmus anguillaris Vaillant 1902 -- (Roberts 1989:94 [ref. 6439]). •Synonym of Pangio anguillaris (Vaillant 1902) -- (Kottelat & Lim 1993:214 [ref. 21437], Tan & Kottelat 2009:61 [ref. 30261], Kottelat 2012:38 [ref. 32367]). Current status: Synonym of Pangio anguillaris (Vaillant 1902). Cobitidae. Habitat: freshwater.


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