Species that contain: Cottus and quadricornis [ 16 ] records

asundensis, Cottus quadricornis Lönnberg [E.] 1932:17, Pl. (figs. 7 a-b) [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Åsunden, Sweden. Syntypes: NRM 20329-32 (1, 5, 3, 3). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

borkensis, Cottus quadricornis Lönnberg [E.] 1939:4 [Arkiv för Zoologi v. 31 B (no. 2); ref. 17199] Lake Borken [Bork], parishes of Yxnerum and Björsäter, Osthrogothland Province, Sweden. Syntypes: NRM 20327 (2). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

frykenensis, Cottus quadricornis Lönnberg [E.] 1932:14, Pl. (fig. 5) [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Fryken, Sweden. Syntypes: NRM 30315 (1), Mus. of Karlstad's school (1), Uppsala Univ. Zool. Mus. (1). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

hexacornis, Cottus Richardson [J.] 1823:726 [22] [Narrative of a journey to the shores of the Polar Sea; ref. 18624] Mouth of Tree River near Coppermine River, Arctic Sea about Herschel Island. On p. 22 of separate. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1129 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Mecklenburg et al. 2002:477 [ref. 25968], Parin et al. 2014:262 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

kallavesensis, Cottus quadricornis race Lönnberg [E.] 1933:7, Pl. 1 (fig. 1) [Arkiv för Zoologi v. 24 A (pt 3) (no. 12) [1932]; ref. 22149] Lake Kallavesi; Lake Niinivesi in Rautalampi; Lake Virmavesi in Karttula, Lake Päijänne basin, Finland. Syntypes: MZH (9). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies onegensis Berg & Popov 1932 -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

latifrons, Cottus Gratzianov [V. I.] 1904:195, Pl. 11 [Zapiski Imperatorskago russkago geograficheskago obshchestva po obshchei geografii v. 41 (no. 1); ref. 21411] Western coast Kanin Peninsula, northern Russia, Barents Sea. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies labradoricus (Girard 1850) -- (Berg 1949:1130 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952], Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Neelov 1973:597 [ref. 7219], Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

lonnbergi, Cottus quadricornis Lönnberg [E.] 1919:167, 171 [Fauna och Flora (Uppsala) v. 14 (no. 5); ref. 23061] Lake Mälaren, Sweden. Syntypes: ?NRM. Type catalog: Sideleva & Zhidkov 2021:240 [ref. 38705]. Name should not be changed since it is Swedish (see Koerber 2009:63 [ref. 30671] for spelling of Lönnberg and origin of his name). •Validates the infrasubspecific Myoxocephalus quadricornis morpha lonnbergi Berg 1916. •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Sideleva & Zhidkov 2021:240 [ref. 38705]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

oernensis, Cottus quadricornis Lönnberg [E.] 1932:16, Figs. 6 a-b [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Örn, Sweden. Syntypes: (9) ?BMNH 1932.5.17.1 (1), NRM 20323-25 (1, 5, 2). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

paulus, Cottus Williams [J. D.] 2000:302 [Copeia 2000 (no. 1); ref. 24258] Coldwater Spring, 5.7 miles west of Oxford about 500 yards north of U.S. highway 78, tributary of Coosa River, Calhoun County, Alabama, U.S.A. Holotype: UMMZ 187462. Paratypes: ALA 2031 (12), 1138 (39), 1665 (68); CUMV 52064 [ex ALA 1063] (26); TU 34396 (47); UMMZ 187463 (25); USNM 201404 [ex ALA 1063] (26). Replacement name for Cottus pygmaeus Williams 1968, preoccupied by Cottus quadricornis pygmaeus Lönnberg 1932. •Valid as Cottus paulus Williams 2000 -- (Kinziger et al. 2000:1007 [ref. 25019], Nelson et al. 2004:119 [ref. 27807], Boschung & Mayden 2004:424 [ref. 27995], Neely et al. 2007:651 [ref. 29262], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:476 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Replacement Name. Current status: Valid as Cottus paulus Williams 2000. Cottidae. Distribution: Coldwater Spring, Coosa River system, Alabama, U.S.A. Habitat: freshwater.

polaris, Cottus Sabine [E.] 1824:213 [ccxiii] [A supplement to the appendix of Captain Parry's voyage; ref. 18649] North Georgia, east side Boothia Peninsula, Northwest Territories, Canada. No types known. •Synonym of Triglopsis quadricornis (Linnaeus 1758), but a valid subspecies -- (Chereshnev 1996:606 [ref. 24659]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Lee et al. 1980:826 [ref. 22416], Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

pygmaeus, Cottus Williams [J. D.] 1968:334, Fig. 1 [Copeia 1968 (no. 2); ref. 7934] Spring, 5.7 miles west of Oxford about 500 yards north of U.S. highway 78, tributary of Coosa River, Calhoun County, Alabama, U.S.A. Holotype: UMMZ 187462. Paratypes: ALA 2031 (12), 1138 (39), 1665 (68); CUMB 52064 [ex ALA 1063] (26); TU 34396 (47); UMMZ 187463 (25); USNM 201404 [ex ALA 1063] (26). Type catalog: Suttkus 1970:128 [ref. 20861], Bart & Taylor 1993:67 [ref. 20906]. Objectively invalid; preoccupied by Cottus quadricornis pygmaeus Lönnberg 1932; replaced by Cottus paulus Williams 2000. •Treated as valid as Cottus pygmaeus Williams 1968 -- (Lee et al. 1980:821 [ref. 22416], Page & Burr 1991:252 [ref. 18983], Boschung 1992:102 [ref. 23239], Mettee et al. 1996:495 [ref. 25836]). 1°Homonym. Current status: Synonym of Cottus paulus Williams 2000. Cottidae. Habitat: freshwater.

pygmaeus, Cottus quadricornis Lönnberg [E.] 1933:9, Pl. 1 (fig. 2) [Arkiv för Zoologi v. 24 A (pt 3) (no. 12) [1932]; ref. 22149] Lake Puruvesi, Finland. Syntypes: MZH. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies onegensis Berg & Popov 1932 -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

quadricornis, Cottus Linnaeus [C.] 1758:264 [Systema Naturae, Ed. X v. 1; ref. 2787] Baltic Sea. Syntypes: NRM 2491-92 (1, 1). Type catalog: Fernholm & Wheeler 1983:235-236 [ref. 20707] who suggest that the largest syntype would make the best lectotype. •Valid as Triglopsis quadricornis (Linnaeus 1758) -- (Neelov 1979:133 [ref. 3152], Fedorov in Whitehead et al. 1986:1260 [ref. 13677], Kottelat 1997:170 [ref. 22952], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:154 [ref. 23595], Novikov et al. 2000:161 [ref. 40929], Bogutskaya et al. 2001:46 [ref. 26178], Parin et al. 2002:S89 [ref. 28300], Bogutskaya & Naseka 2004:191 [ref. 28183], Fricke 2007:29 [ref. 30577], Kottelat & Freyhof 2007:519 [ref. 29996], Sideleva et al. 2022:3 [ref. 40100]). •Possibly valid as Megalocottus quadricornis (Linnaeus 1758) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus quadricornis (Linnaeus 1758) -- (Berg 1949:1128 [ref. 20662], Neelov 1973:597 [ref. 7219], Lee et al. 1980:826 [ref. 22416], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:504 [ref. 25518], McAllister 1990:200 [ref. 14674], Page & Burr 1991:245 [ref. 18983], Scott & Crossman 1998:842 [ref. 24570], Mecklenburg et al. 2002:477 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Scharpf 2008:11 [ref. 30399], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212], Page & Burr 2011:468 [ref. 31215], Page et al. 2013:124 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:263 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:46, 227 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:80 [ref. 34440], Mecklenburg et al. 2018:217 [ref. 36370], Love et al. 2021:123 [ref. 39279]). Current status: Valid as Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Distribution: Circumpolar in Arctic Ocean, in Pacific and north Atlantic. Habitat: freshwater, brackish, marine.

relicta, Cottus quadricornis var. Lilljeborg [N.] 1891:149 [Sveriges och Norges fauna. Fiskarne. v. 1; ref. 21766] Lake Vättern, Sweden. No types known. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies relictus (Lilljeborg 1891) -- (Berg 1949:1133 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

vaenernensis, Cottus quadricornis Lönnberg [E.] 1932:8, Fig. 3 [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Vänern, Sweden. Syntypes: NRM 20336-38 (1, 4, 1), 40338 (1). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

vermelnensis, Cottus quadricornis Nybelin [O.] 1940:77, Fig. 1 [Fauna och Flora (Uppsala) v. 35 (no. 2); ref. 23062] Lake Värmeln, Sweden. Syntypes: whereabouts unknown. •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.