Species that contain: Engraulis and encrasicolus [ 17 ] records

amara, Engraulis Risso [A.] 1827:456 [Histoire naturelle des principales productions de l'Europe méridionale v. 3; ref. 3757] Nice, France, northwestern Mediterranean Sea. No types known. Type catalog: Whitehead & Bauchot 1986:47 [ref. 9739]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

aquitanicus, Engraulis encrasicolus Aleksandrov [A. I.] 1927:77 [Trudy Kerčenskoj Naučnoj Rybochozjajstvennoj Stancii v. 1 (2-3); ref. 21154] Bay of Biscay and probably near shores of Portugal. No types known. After "race Atlantique, groupe méridional" Fage 1920. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

argyrophanus, Engraulis Valenciennes [A.] (ex Kuhl & van Hasselt) in Cuvier & Valenciennes 1848:49 [Histoire naturelle des poissons v. 21; ref. 1013] Equatorial Atlantic off western Africa. Lectotype: MNHN A-7614. Paralectotypes: MNHN B-3103 [ex MNHN A-7614] (4), RMNH 3331 (2). Type catalog: Whitehead & Bauchot 1986:31-32 [ref. 9739]. Lectotype selected by Whitehead 1967:125 [ref. 6464]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

atlantica, Engraulis encrassicholus forma de Buen [F.] 1926:49 [Resultado de las Campañas Realizadas por Acuerdos Internacionales, Madrid No. 2; ref. 5054] Gulf of Cádiz and coasts of Larache, Spain. Based on literature sources. Species should have been encrasicolus. Apparently independent of Pusanov & Zeeb 1926; earliest not reserached. •Synonym of Engraulis encrasicolus (Linnaeus 1758) Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

atlanticus, Engraulis encrasicholus Pusanov [I. I.] & Zeeb [Y.] 1926:93 [Trudy Krimskogo Naucno-Issledovatel'skogo Instituta Biologii i Biologiceskogo Fakul'teta v. 1; ref. 18576] Atlantic. After "race Atlantique" of Fage 1920. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

capensis, Engraulis Gilchrist [J. D. F.] 1913:62, Fig. [Marine biological report: for the year ended ... and for the half year ending ...; to be presented to the Provincial Council; Union of South Africa, Province of the Cape of Good Hope. No. 1 (art. 2); ref. 14154] South Africa. No types known. •Synonym of Engraulis japonicus Temminck & Schlegel 1846 -- (Whitehead & Wongratana 1986:205 [ref. 6284]). •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead 1990:228 [ref. 14039]). •Valid as Engraulis capensis Gilchrist 1913 -- (Whitehead et al. 1988:315 [ref. 5725], Bianchi & Carpenter in Bianchi et al. 1993:138 [ref. 25472], Fricke et al. 2018:52 [ref. 35805], Anderson 2022:169 [ref. 39690]). Current status: Valid as Engraulis capensis Gilchrist 1913. Engraulidae: Engraulinae. Distribution: Southeastern Atlantic, Western Indian Ocean: Namibia south to Cape of Good Hope (South Africa), then north to Somalia, east to Madagascar and Mascarenes (La Réunion, Mauritius, Rodrigues). Habitat: marine.

encrasicolus, Clupea Linnaeus [C.] 1758:318 [Systema Naturae, Ed. X v. 1; ref. 2787] 18 nautical miles off Sète, Gulf of Lyon, France, northwestern Mediterranean Sea. MNHN 2002-1775. Neotype designated by Fricke 1999:20 [ref. 24101], but withdrawn in Fricke 2000:639 [ref. 24537]. Neotype designated by Borsa et al. 2004:1113 [ref. 28607]. Spelled encrasiolus in Brünnich 1768:83 [ref. 17290]. Sometimes spelled encrasicholus or encrassicolus.. •Valid as Engraulis encrasicolus (Linnaeus 1758) -- (Herrera 1896:30 [ref. 39336] as encrasicholus, Svetovidov 1973:111 [ref. 7169], Dor 1984:43 [ref. 29757], Nelson 1984:424 [ref. 5190], Whitehead in Whitehead et al. 1984:282 [ref. 13675], Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039], Goren & Dor 1994:11 [ref. 25356], Coad 1995:9 [ref. 23608], Coad 1996:84 [ref. 25304], Santos et al. 1997:121 [ref. 23531], Arruda 1997:30 [ref. 24952] as encrasicholus, Fricke 1999:20 [ref. 24101], Bilecenoğlu et al. 2002:39 [ref. 26753], Falcón et al. 2003:146 [ref. 39343], Manilo & Bogorodsky 2003:S97 [ref. 27377], Bogutskaya & Naseka 2004:30 [ref. 28183], Borsa et al. 2005:1121 [ref. 28607], Golani 2005:20 [ref. 37112], Fricke 2007:26 [ref. 30577], Fricke et al. 2007:25 [ref. 29533], Vasil'eva 2007:44 [ref. 30517], Fricke 2008:15 [ref. 30182], Lipej & Dulčić 2010:20 [ref. 36649], Wirtz et al. 2013:118 [ref. 32972], Parin et al. 2014:51 [ref. 33547], Nizinski & Munroe 2016:1717 [ref. 34559], Bogorodsky & Randall 2018:[22] 260 [ref. 36492], Golani & Fricke 2018:32 [ref. 36273], Martins & Carneiro 2018:29 [ref. 36550], Artüz & Fricke 2019:553 [ref. 36612], Carneiro et al. 2019:74 [ref. 37250], Elbaraasi et al. 2019:92 [ref. 36864], Reiner 2019:79 [ref. 38139], Bariche & Fricke 2020:45 [ref. 37515], Kovačić et al. 2020:12 [ref. 37519], Abliazov et al. 2021:8 [ref. 38576], Bañón & Maño 2021:52 [ref. 38883], Golani 2021:20 [ref. 38303], Kovačić et al. 2021:22 [ref. 39014], Bonhomme et al. 2022:597 [ref. 39183] as cf. encrasicolus, Dyldin et al. 2022:[18] [ref. 39193], Fermon et al. 2022:129 [ref. 39448], Niță et al. 2022:43 [ref. 40471]). Current status: Valid as Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Distribution: Western Baltic Sea; North Sea; Mediterranean Sea; Sea of Marmara; Black Sea; Sea of Azov; eastern Atlantic: Norway south to South Africa, including Azores, Madeira, Canary Islands, and São Tomé and Principe; Red Sea (Mediterranean Sea immigrant); southwestern Indian Ocean. Habitat: brackish, marine.

guineensis, Anchoviella Rossignol [M.] & Blache [J.] 1961:285, Fig. 1 [Bulletin du Muséum National d'Histoire Naturelle (Série 2) v. 33 (no. 3); ref. 9375] Off Cape Lopez, Gabon, depth 20 meters. Syntypes: MNHN 1961-0310 (10). Additional material: ?MNHN [ex ORSTOM Dakar] uncat. (70). Type catalog: Whitehead & Bauchot 1986:30 [ref. 9739]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Nelson 1984:424 [ref. 5190], Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:229 [ref. 14039], Nizinski & Munroe 2016:1717 [ref. 34559]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

maeoticus, Engraulis encrasicholus Pusanov [I. I.] & Zeeb [Y.] 1926:94 [Trudy Krimskogo Naucno-Issledovatel'skogo Instituta Biologii i Biologiceskogo Fakul'teta v. 1; ref. 18576] Sea of Azov, Russia/Ukraine. Syntypes: whereabouts unknown. Bonhomme et al. 2022:[6] 598 [ref. 39183] treat MNHN 2002-1716 (holotype of Engraulis albidus Borsa, Collet & Durand 2005) from the Mediterranean coast of southern France as the holotype of Engraulis encrasicolus maeoticus Pusanov & Zeeb 1926, without designating a neotype; this action is not valid, as the syntypes originated from the Sea of Azov and were collected in the 1920s. •Synonym of Engraulis encrasicolus (Linnaeus 1758), but a valid subspecies as described -- (Ninua & Japoshvili 2008:164 [ref. 31251]). •Possibly valid as Engraulis maeoticus Pusanov & Zeeb 1926 -- (Bonhomme et al. 2022:[6] 598 [ref. 39183]). •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725], Bogutskaya & Naseka 2004:30 [ref. 28183], Parin et al. 2014:51 [ref. 33547]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

mediterraneus, Engraulis encrasicolus Pusanov [I. I.] & Zeeb [Y.] 1926:93 [Trudy Krimskogo Naucno-Issledovatel'skogo Instituta Biologii i Biologiceskogo Fakul'teta v. 1; ref. 18576] Mediterranean Sea. After "race Méditerranéenne" Fage 1920. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725], Bogutskaya & Naseka 2004:30 [ref. 28183]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

meletta, Engraulis Cuvier [G.] 1829:323 [Le Règne Animal (Edition 2) v. 2; ref. 995] Mediterranean Sea. No types known. Type catalog: Whitehead & Bauchot 1986:49 [ref. 9739]. Based on brief description and citation to "Duham. sect. VI, pl. iii, f. 5" [Duhamel 1777:157-158, Pl. 3 (fig. 5)]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

oliveri, Engraulis russoi var. Scuderi [I.] 1957:242 [Bollettino di Pesca, Piscicoltura e Idrobiologia, Roma (n. s.) v. 12 (no. 2); ref. 18715] Lago Verdi, brackish marine pond along the Tyrrhenian coast of Sicily near Oliveri, Italy. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

pontica, Engraulis encrasicolus mediterraneus natio Pusanov [I. I.] & Zeeb [Y.] 1926:93 [Trudy Krimskogo Naucno-Issledovatel'skogo Instituta Biologii i Biologiceskogo Fakul'teta v. 1; ref. 18576] Sevastopol, Crimea, Ukraine, Black Sea. No types known. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Bogutskaya & Naseka 2004:30 [ref. 28183]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

ponticus, Engraulis encrasicholus Aleksandrov [A. I.] 1927:78 [Trudy Kerčenskoj Naučnoj Rybochozjajstvennoj Stancii v. 1 (2-3); ref. 21154] Western part of Black Sea. No types known. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Bogutskaya & Naseka 2004:31 [ref. 28183], Parin et al. 2014:51 [ref. 33547]). •Synonym of Engraulis encrasicolus (Linnaeus 1758), but a valid subspecies as described -- (Ninua & Japoshvili 2008:164 [ref. 31251]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Distribution: The subspecies occurs in the Black Sea basin. Habitat: brackish, marine.

russoi, Engraulis Dulzetto [F.] 1947:183, Figs. 1-6 [Memorie di matematica e di scienze fisiche e naturali della Società Italiana delle Scienze. v. 26; ref. 17424] Messina, Sicily, Italy, Mediterranean Sea. Neotype: MSNG 39522. Neotype designated by Tortonese 1967 [ref. 10366]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:229 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

symaetensis, Engraulis encrasicholus Dulzetto [F.] 1938:397, Pls. 1 (fig. 2), 2 (fig. 4) [Atti della Reale Accademia d'Italia, Memorie della Classe di Scienze Fisiche, Matematiche e Naturali v. 11 (no. 8); ref. 17423] Catania, eastern Sicily, Italy, Mediterranean Sea. Neotype: MSNG 39522. Neotype designated by Tortonese 1967. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.

vulgaris, Engraulis Nilsson [S.] 1832:25 [Prodromus ichthyologiae Scandinavicae; ref. 3204] Scandinavia. No types known. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae: Engraulinae. Habitat: brackish, marine.