Species that contain: Epinephelus and bruneus [ 4 ] records

bruneus, Epinephelus Bloch [M. E.] 1793:15, Pl. 328 (fig. 2) [Naturgeschichte der ausländischen Fische v. 7; ref. 4868] "Norway" [likely China]. Lectotype: ZMB 5221 (left skin). Paralectotypes: (2, lost). Type catalog: Paepke 1999:135 [ref. 24282]. Lectotype selected by Randall & Heemstra 1991:109 [ref. 19601]. On p. 13 of Bloch's Ichthyologie, v. 10 [ref. 21381]. Often misspelled brunneus. •Valid as Epinephelus bruneus Bloch 1793 -- (Lee 1990:39 [ref. 19830], Randall & Heemstra 1991:108 [ref. 19601], Heemstra & Randall 1993:119 [ref. 21042], Heemstra & Randall 1999:2490 [ref. 24815], Nakabo 2000:718 [ref. 25086], Randall in Randall & Lim 2000:609 [ref. 25122], Sadovy & Cornish 2000:82 [ref. 25480], Nakabo 2002:718 [ref. 26001], Choi et al. 2003:309, 590 [ref. 26218], Motomura et al. 2010:96 [ref. 31256], Craig et al. 2011:97 [ref. 31786], Motomura et al. 2015:19 [ref. 34709], Koeda et al. 2016:26 [ref. 35333], Ma & Craig 2018:455 [ref. 36100], Nakae in Kimura et al. 2018:101 [ref. 36460], Kim et al. 2020:115 [ref. 39118], Parenti & Randall 2020:63 [ref. 37354], Murase et al. 2023:39 [ref. 40935]). Current status: Valid as Epinephelus bruneus Bloch 1793. Epinephelidae. Distribution: Northwestern Pacific: Vietnam east to Philippines, north to Korea and Japan. Habitat: marine.

fuscus, Holocentrus Lacepède [B. G. E.] 1802:342, 384 [Histoire naturelle des poissons (Lacepède) v. 4; ref. 4929] "Norway" [likely China]. Lectotype: ZMB 5221 (left half skin). Unneeded new name for Epinephelus bruneus Bloch 1793. •Synonym of Epinephelus bruneus Bloch 1793 -- (Parenti & Randall 2020:63 [ref. 37354]). Current status: Synonym of Epinephelus bruneus Bloch 1793. Epinephelidae. Habitat: marine.

moara, Serranus Temminck [C. J.] & Schlegel [H.] 1843:10, Pl. 4 (fig. 2 [not 1]) [Fauna Japonica Part 1; ref. 4370] Nagasaki, Japan. Lectotype: RMNH D72 (dry). Paralectotypes: RMNH D70 and D71 (2, dry), 181 (1, dry?); ZMB 5215 [ex RMNH] (1, dry). Originally as mo-ara, correction mandatory. Lectotype selected by Boeseman 1947:28 [ref. 12876]. •Synonym of Epinephelus bruneus Bloch 1793 -- (Lee 1990:39 [ref. 19830], Randall & Heemstra 1991:108 [ref. 19601], Heemstra & Randall 1993:119 [ref. 21042], Heemstra & Randall 1999:2490 [ref. 24815], Sadovy & Cornish 2000:82 [ref. 25480] as mo-ara). •Valid as Epinephelus moara (Temminck & Schlegel 1843) -- (Katayama in Masuda et al. 1984:131 [ref. 6441], Machida in Okamura et al. 1985:475, 671 [ref. 8056], Liu et al. 2013:1684 [ref. 32705], Ma & Craig 2018:455 [ref. 36100], Parenti & Randall 2020:76 [ref. 37354], Sonoyama et al. 2020:54 [ref. 37637]). Current status: Valid as Epinephelus moara (Temminck & Schlegel 1843). Epinephelidae. Distribution: Northwestern Pacific: South China Sea to Japan. Habitat: marine.

testudineus, Holocentrus Shaw [G.] 1803:566 [General zoology or systematic natural history v. 4; ref. 4014] Northern seas. Unneeded new name for Epinephelus bruneus Bloch 1793. •Synonym of Epinephelus bruneus Bloch 1793 -- (Parenti & Randall 2020:64 [ref. 37354]). Replacement Name. Current status: Synonym of Epinephelus bruneus Bloch 1793. Epinephelidae. Habitat: marine.