Institute for Biodiversity Science and Sustainability
Online Version, Updated 6 November 2023

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Catalog of Fishes

Species that contain: Esox and belone [ 20 ] records

aaveri, Esox Curtiss [A.] 1938:40 [A short zoology of Tahiti in the Society islands; ref. 18057] Lagoon near Tautira, Tahiti, Society Islands, French Polynesia, South Pacific. No types known. Original as Esox ááveri. •Synonym of Belone marisrubri (Bloch & Schneider 1801) -- (Mees 1962:45 [ref. 13277]). •Synonym of Tylosurus crocodilus (Péron & Lesueur 1821), subspecies crocodilus -- (Collette 2003:11 [ref. 27306]). •Synonym of Tylosurus crocodilus (Péron & Lesueur 1821). Current status: Synonym of Tylosurus crocodilus (Péron & Lesueur 1821). Belonidae. Habitat: brackish, marine.

bellone, Esox Linnaeus [C.] 1758:314 [Systema Naturae, Ed. X v. 1; ref. 2787] European seas. Syntypes: BMNH 1853.11.12.147 [Gronovius coll.] (1, skin). Type catalog: Wheeler 1958:215 [ref. 13434]. •Treated as valid as Belone bellone (Linnaeus 1758) -- (Mees 1962:17 [ref. 13277]). •Nomen dubium -- (Collette & Parin 1970:17-18 [ref. 7415], Collette 2003:16 [ref. 27306]). •Uncertain in Belonidae -- (Fricke 2008:22 [ref. 30182]). Nomen Dubium. Belonidae. Habitat: marine.

belone, Esox Linnaeus [C.] 1761:126 [Fauna Svecica, sistens Animalia Sveciae Regni, ed. II; ref. 13326] European seas. No types known. Neotype designated by Fricke 1999:35 [ref. 24101], but withdrawn in Fricke 2000:639 [ref. 24537]. Placed on Official List in ICZN Opinion 225 as type species of Belone Cuvier 1816 (See also Esox bellone Linnaeus 1758). See Collette 2003:3 [ref. 27306] for subspecies -- Three subspecies were recognized by Collette and Parin (1970 [ref. 7415]): B. b. belone (Linnaeus 1761) from northeastern Atlantic Ocean; B. b. euxini Günther 1866 from the Black Sea and Sea of Azov; and B. b. acus Risso 1827 from the Mediterranean Sea and adjacent parts of the Atlantic, Madeira, Canary Islands, Azores, and south to the Cape Verde Islands. •Valid as Esox belone Linnaeus 1761 -- (Herrera 1896:31 [ref. 39336]). •Valid as Belone belone (Linnaeus 1761), subspecies belone (Linnaeus 1761) -- (Collette 2003:13 [ref. 27306], Parin et al. 2014:196 [ref. 33547], Collette 2016:2123 [ref. 34601], Mecklenburg et al. 2018:166 [ref. 36370]). •Valid as Belone belone (Linnaeus 1761) -- (Collette & Parin 1970:11 [ref. 7415], Parin 1973:258 [ref. 7191], Fricke 1999:35 [ref. 24101], Bilecenoğlu et al. 2002:60 [ref. 26753], Collette 2003:2 [ref. 27306], Golani 2005:28 [ref. 37112], Fricke 2007:28 [ref. 30577], Ninua & Japoshvili 2008:169 [ref. 31251], Lipej & Dulčić 2010:34 [ref. 36649], Martins & Carneiro 2018:97 [ref. 36550], Artüz & Fricke 2019:554 [ref. 36612], Carneiro et al. 2019:199 [ref. 37250], Elbaraasi et al. 2019:95 [ref. 36864], Kovačić et al. 2020:29 [ref. 37519], Abliazov et al. 2021:9 [ref. 38576], Bañón & Maño 2021:63 [ref. 38883], Dolgov & Zabavnikov 2021:613 [788] [ref. 38754], Golani 2021:31 [ref. 38303], Kovačić et al. 2021:52 [ref. 39014], Niță et al. 2022:30 [ref. 40471]). Current status: Valid as Belone belone (Linnaeus 1761). Belonidae. Distribution: Baltic Sea; North Sea; eastern Atlantic and adjacent Arctic, including Azores and Madeira. Habitat: brackish, marine.

cancila, Esox Hamilton [F.] 1822:213, 380, Pl. 27 (fig. 70) [An account of the fishes found in the river Ganges; ref. 2031] Ponds and smaller rivers of Gangetic provinces [India, Bangladesh]. No types known. Hamilton's original illustration reproduced by Britz 2019:Pl. 121 [ref. 37532]. •Valid as Belone cancila (Hamilton 1822) -- (Mees 1962:20 [ref. 13277]). •Placed in Xenentodon -- (Collette & Berry 1965 [ref. 13367]). •Valid as Xenentodon cancila (Hamilton 1822) -- (Shrestha 1978:40 [ref. 15970], Pethiyagoda 1991:175 [ref. 20075], Talwar & Jhingran 1991:743 [ref. 20764], Kottelat et al. 1993:89 [ref. 23448], Indra 1994:421 [ref. 24015], Sen 1995:581 [ref. 23778], Rainboth 1996:172 [ref. 22772], Rema Devi et al. 1996:138 [ref. 24021], Banik & Choudhury 1996:579 [ref. 24736], Menon 1999:265 [ref. 24904], Fuller et al. 1999:287 [ref. 25838], Freyhof et al. 2000:96 [ref. 25184], Rafique 2000:326 [ref. 25220], Raju Thomas et al. 2000:444 [ref. 25319], Karmakar 2000:34 [ref. 25662], Lovejoy & Collette 2001:329 [ref. 25232], Raju Thomas et al. 2002:50 [ref. 26359], Mirza 2003:21 [ref. 27265], Collette 2003:12 [ref. 27306], Ataur Rahman 2003:71 [ref. 31338], Mundy 2005:283 [ref. 28379], Shrestha 2008:204 [ref. 29923], Conway et al. 2011:1756 [ref. 31594], Ahmed et al. 2013:287 [ref. 32872], Gurung et al. 2013:4884 [ref. 32969], Kottelat 2013:298 [ref. 32989], Narasimhaiah et al. 2016:69 [ref. 34975], Arunkumar & Manimekalan 2018:1739 [ref. 36172], Suresh et al. 2018:161 [ref. 36495], Psomadakis et al. 2020:337 [ref. 37272] as cf. cancila, Shangningam & Kosygin 2022:60 [ref. 39587], Tudu et al. 2022:44 [ref. 40484], Chan et al. 2023:152 [ref. 40365]). Current status: Valid as Xenentodon cancila (Hamilton 1822). Belonidae. Distribution: South Asia, Southeast Asia: Pakistan, India, Sri Lanka, Bangladesh, Bhutan, Myanmar, Malaysia and Thailand. Introduced elsewhere, including Hawaiian Islands (U.S.A.). Habitat: freshwater.

graii, Belone Sykes [W. H.] 1839:163 [Proceedings of the Zoological Society of London 1838 (pt 6) (for 27 Nov. 1838); ref. 18720] Deccan, India. No types known. Appeared first as above, then in Sykes 1839:60 [ref. 4306]; more complete treatment in Sykes 1841:350, Pl. 60 (fig. 1) [ref. 18868]. •Synonym of Esox cancila Hamilton 1822 -- (Mees 1962 [ref. 13277]). •Synonym of Xenentodon cancila (Hamilton 1822) -- (Menon 1999:265 [ref. 24904], Collette 2003:12 [ref. 27306], Kottelat 2013:298 [ref. 32989]). Current status: Synonym of Xenentodon cancila (Hamilton 1822). Belonidae. Habitat: freshwater.

hindostanicus, Esox (Belone) Falconer [H.] 1868:589 [Miscellaneous Notes Indian Zoology v. 1; ref. 13399] Nullahs [tempory stream] and stagnant waters at Suharunpore [Sukarumpoor], India. No types known. Original description is reproduced in Myers 1960:245-246 [ref. 9582]. •Synonym of Belone cancila (Hamilton 1822) -- (Mees 1962:20 [ref. 13277]). •Synonym of Xenentodon cancila (Hamilton 1822) -- (Myers 1960:246 [ref. 9582], Talwar & Jhingran 1991:743 [ref. 20764], Collette 2003:12 [ref. 27306]). Current status: Synonym of Xenentodon cancila (Hamilton 1822). Belonidae. Habitat: freshwater.

houttuyni, Esox Walbaum [J. J.] 1792:88 [Petri Artedi sueci genera piscium Part 3; ref. 4572] No locality. No types known. Placed on Official Index (Opinion 900); suppressed for purpose of priority but not for homonymy. •Valid as Belone houttuyni (Walbaum 1792) -- (Mees 1962:33 [ref. 13277]). •In the synonymy of Strongylura marina (Walbaum 1792) -- (Collette & Berry 1965:391 [ref. 13367], Castro-Aguirre et al. 1999:201 [ref. 24550], Collette 2003:7 [ref. 27306]). Suppressed. Current status: Synonym of Strongylura marina (Walbaum 1792). Belonidae. Habitat: freshwater, brackish, marine.

houttuyni, Esox belone var. Bloch [M. E.] & Schneider [J. G.] 1801:391 [M. E. Blochii, Systema Ichthyologiae; ref. 471] No locality. No types known. Apparently independent of Esox houttuyni Walbaum 1792 but based on the same source. Objectively invalid; preoccupied by Esox houttuyni Walbaum 1792. •Synonym of Strongylura marina (Walbaum 1792) -- (Collette 2003:7 [ref. 27306]). 1°Homonym. Current status: Synonym of Strongylura marina (Walbaum 1792). Belonidae. Habitat: freshwater, brackish, marine.

imperialis, Esox Rafinesque [C. S.] 1810:59, Pl. 9 (fig. 2) [Caratteri di alcuni nuovi generi e nuove specie di animali e piante della Sicilia; ref. 3594] Palermo, Sicily, Italy, Mediterranean Sea. No types known. Also appeared in Rafinesque 1810:34 [ref. 3595]. Sphyraena acus was placed on the Official Index and Esox imperialis on the Official List. See also Sphyraena acus Lacepède 1803 [note: because of changes, synonyms of T. imperialis may not be synonyms of Tylosurus imperialis imperialis] and a new petition will be submitted to preserve use of acus as in current literature. This subspecies occurs in the Mediterranean Sea. •Valid as Belone imperialis (Rafinesque 1810) -- (Mees 1962:38 [ref. 13277]). •Synonym of Tylosurus acus (Lacepède 1803) -- (Collette & Berry 1965:391 [ref. 13367]). •Synonym of Tylosurus acus (Lacepède 1803), but a valid subspecies imperialis -- (Parin 1973:260 [ref. 7191], Masuda et al. 1984 [ref. 6441], Collette & Parin in Whitehead et al. 1986:607 [ref. 13676], Collette & Parin 1990:596 [ref. 16473], Schmitter-Soto et al. 2000:154 [ref. 27754], Collette & Banford 2001:53 [ref. 25982], Bilecenoğlu et al. 2002:60 [ref. 26753], Collette 2003:9, 10 [ref. 27306], Paxton et al. 2006:719 [ref. 28995], Lipej & Dulčić 2010:34 [ref. 36649], Collette 2016:2121 [ref. 34601], Collette 2016:2126 [ref. 34601]). •Valid as Tylosurus imperialis (Rafinesque 1810) -- (Fricke et al. 2007:70 [ref. 29533], Wirtz et al. 2013:121 [ref. 32972], Parenti 2019:101 [ref. 37117], Bariche & Fricke 2020:42 [ref. 37515], Ketsilis-Rinis & Koutsidi in Bo et al. 2020:617 [ref. 37876], Michailidis in Gerovasileiou et al. 2020:355 [ref. 37890], Kovačić et al. 2021:52 [ref. 39014]). Current status: Valid as Tylosurus imperialis (Rafinesque 1810). Belonidae. Distribution: Mediterranean Sea; eastern Atlantic: Morocco south to Cape Verde Islands. Habitat: marine.

linnei, Belone Malm [A. W.] 1877:553 [ ref. 2881] European seas. No types known. Original as Linnéi. An unneeded new name for Esox belone Linnaeus 1758 (to avoid "Strickland tautonomy"). •Synonym of Belone belone (Linnaeus 1761), subspecies belone (Linnaeus 1761) -- (Parin 1973:258 [ref. 7191], Collette 2003:3, 15 [ref. 27306]). Current status: Synonym of Belone belone (Linnaeus 1761). Belonidae. Habitat: brackish, marine.

longirostris, Belone Schinz [H. R.] 1822:312 [Das Thierreich v. 2; ref. 3926] European Seas. No types known. Unneeded new name for and objective synonym of Esox belone Linnaeus 1761 (see Daget 1968:1058 [ref. 26392]). •Valid as Tylosurus longirostris Schinz 1822 -- (Herrera 1896:34 [ref. 39336] with author as Gill). •Objective synonym of Belone belone (Linnaeus 1761) -- (Daget 1968:1058 [ref. 26392], Collette 2003:2 [ref. 27306]). •Synonym of Belone belone (Linnaeus 1761), subspecies belone (Linnaeus 1761) -- (Collette 2003:2, 15 [ref. 27306]). Current status: Synonym of Belone belone (Linnaeus 1761). Belonidae. Habitat: brackish, marine.

longirostris, Esox Mitchill [S. L.] 1818:322 [American Monthly Magazine and Critical Review v. 2 (no. 5, art. 1); ref. 30214] Hudson River near Albany, New York, U.S.A. No types known. •Synonym of Belone houttuyni (Walbaum 1792) -- (Mees 1962:33 [ref. 13277]). •Synonym of Strongylura marina (Walbaum 1792) -- (Smith 1986:408 [ref. 22186], Collette 2003:7 [ref. 27306]). Current status: Synonym of Strongylura marina (Walbaum 1792). Belonidae. Habitat: freshwater, brackish, marine.

marinus, Acus Müller [P. L. S.] 1774:341 [Des Ritters Carl von Linné vollständiges Natursystem nach der 12ten Ausgabe, und nach Anleitung Houttynischen Werkes, mit einer ausführlichen Erklärung ausgefertigt. 6 vols. 4; ref. 6461] Not available, published in a rejected work on Official Index (Opinion 701). Acus used only as a Latin vernacular name in reference to Esox belone Linnaeus 1761; Acus marinus was attributed to Müller but is a Latin translation of the German vernacular name 'Meernadel', used by a subsequent author. Belonidae.

marinus, Esox Walbaum [J. J.] (ex Schöpf) 1792:88 [Petri Artedi sueci genera piscium Part 3; ref. 4572] No locality stated [New York, U.S.A.]. No types known. Based on 'Esox, Sea Pike, Sea Snipe, zu Neuyork' in Schöpf 1788:177 [ref. 18700]. All of Valenciennes' specimens of Belone timucu are Strongylura marina (Collette et al. 1997:27 [ref. 22813]). Placed on Official List in Opinion 900 [ref. 13280]. •Synonym of Belone houttuyni (Walbaum 1792) -- (Mees 1962:33 [ref. 13277]). •Valid as Strongylura marina (Walbaum 1792) -- (Lee et al. 1980:489 [ref. 22416], Smith 1986:408 [ref. 22186], Robins & Ray 1986:106 [ref. 23100], Cervigón 1991:229 [ref. 24472], Boschung 1992:84 [ref. 23239], Cervigón 1992:269 [ref. 23827], Mettee et al. 1996:441 [ref. 25836], Murdy et al. 1997:118 [ref. 23144], González Bencomo et al. 1997:158 [ref. 24960], McEachran & Fechhelm 1998:936 [ref. 23897], Aguilera 1998:47 [ref. 24221], Keith et al. 2000:28 [ref. 24547], Castro-Aguirre et al. 1999:201 [ref. 24550], Fuller et al. 1999:287 [ref. 25838], Marín 2000:70 [ref. 26252], Schmitter-Soto et al. 2000:154 [ref. 27754], Lovejoy & Collette 2001:329 [ref. 25232], Ross et al. 2001:347 [ref. 25978], Camargo & Isaac 2001:139 [ref. 27639], Collette in Collette & Klein-MacPhee 2002:288 [ref. 26158], Collette 2003:1109 [ref. 26981], Collette 2003:7 [ref. 27306], Menezes & Figueiredo in Menezes et al. 2003:67 [ref. 27192], Smith et al. 2003:14 [ref. 27621], Banford et al. 2004:834 [ref. 27645], Nelson et al. 2004:99 [ref. 27807], Vaske et al. 2005:77 [ref. 36155], Miller 2006:203 [ref. 28615], Scharpf 2007:10 [ref. 30398], García-Hernández et al. 2009:92 [ref. 37265], Page & Burr 2011:461 [ref. 31215], Matamoros et al. 2009:15 [ref. 31241], Angulo et al. 2013:997 [ref. 33194], Di Dario et al. 2013:263 [ref. 37503], Page et al. 2013:102 [ref. 32708], Catelani et al. 2014:932 [ref. 37496], Raz-Guzmán et al. 2018:345 [ref. 36407], Robins et al. 2018:189 [ref. 35886], Ribeiro et al. 2019:5 [ref. 36717], Collette & Bemis 2019:21 [ref. 37342], Caires et al. in Marceniuk et al. 2021:317 [ref. 38945], Angulo et al. 2023:65 [ref. 40676], Carvalho-Filho 2023:91 [ref. 40480], Martínez-Servín et al. 2023:111 [ref. 40726]). Current status: Valid as Strongylura marina (Walbaum 1792). Belonidae. Distribution: Western Atlantic: Massachusetts (U.S.A.) south to southern Brazil, including Gulf of Mexico and western Caribbean Sea, but absent from the Bahamas and the West Indies. Habitat: freshwater, brackish, marine.

marinus, Esox belone var. Bloch [M. E.] & Schneider [J. G.] 1801:391 [M. E. Blochii, Systema Ichthyologiae; ref. 471] North America. No types known. Apparently independent of Esox marinus Walbaum 1792. Objectively invalid; preoccupied by Esox marinus Walbaum 1792. •Synonym of Strongylura marina (Walbaum 1792) -- (Collette 2003:7 [ref. 27306]). 1°Homonym. Current status: Synonym of Strongylura marina (Walbaum 1792). Belonidae. Habitat: freshwater, brackish, marine.

marisrubri, Esox belone var. Bloch [M. E.] & Schneider [J. G.] 1801:391 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Red Sea. No types known. Type catalog: Paepke & Seegers 1986:146 [ref. 19981]. Originally maris rubri. Placed on Official Index (Opinion 900); suppressed for purpose of priority but not for homonymy. •Considered valid by Mees 1962:44 [ref. 13277]. •Synonym of Belone choram Rüppell 1837 -- (Parin 1967 [ref. 10272]). •In the synonymy of Tylosurus crocodilus (Péron & Lesueur 1821) -- (Dor 1984:65 [ref. 29757], Collette & Berry 1965:391 [ref. 13367]). •In the synonymy of Tylosurus choram (Rüppell 1837) -- (Collette 2003:10 [ref. 27306]). Suppressed. Current status: Synonym of Tylosurus choram (Rüppell 1837). Belonidae. Habitat: marine.

teatae, Esox Curtiss [A.] 1938:41 [A short zoology of Tahiti in the Society islands; ref. 18057] Lagoon near Tautira, Tahiti, Society Islands [French Polynesia, South Pacific]. No types known. •Synonym of Belone platyura Bennett 1832 -- (Mees 1962:59 [ref. 13277]). •Synonym of Platybelone argalus (Lesueur 1821), subspecies platyura (Bennett 1832) -- (Parin 1967:13 [ref. 10272], Collette 2003:4, 16 [ref. 27306]). •Synonym of Platybelone platyura (Bennett 1832) -- (authors). Current status: Synonym of Platybelone argalus (Lesueur 1821). Belonidae. Habitat: brackish, marine.

timucu, Belone Valenciennes [A.] in Cuvier & Valenciennes 1846:426 [Histoire naturelle des poissons v. 18; ref. 1011] Rio de Janeiro, Brazil; Cayenne, French Guiana. Syntypes: MNHN 0000-0721 (1) Cayenne, 0000-0722 (1) Brazil, 0000-0944 (3) Brazil, 0000-4508 (1) Rio, 0000-4510 (1) Rio, 0000-4511 (1) Rio. Secondarily preoccupied in Strongylura by Esox belone timucu Walbaum 1792. Mees 1962:33 [ref. 13277] states, "... type locality here restricted to Rio de Janeiro, lectotype the type of B. almeida Quoy & Gaimard, which makes timucu Valenciennes (nec Walbaum) an objective synonym of almeida; but almeida has syntypes and Mees gives no catalog numbers, so Mees 1962:33 [ref. 13277] did not establish a lectotype. •Synonym of Strongylura marina (Walbaum 1792) -- (Collette et al. 1997:27 [ref. 22813], Collette 2003:7 [ref. 27306]). 2°Homonym. Current status: Synonym of Strongylura marina (Walbaum 1792). Belonidae. Habitat: freshwater, brackish, marine.

timucu, Esox belone var. Walbaum [J. J.] 1792:88 [Petri Artedi sueci genera piscium Part 3; ref. 4572] Lake Papary, about 24 miles south of Natal, Brazil. Neotype: CAS-SU 22073. Neotype designated by Collette 1968:191 [ref. 12725]. •Status uncertain -- (Mees 1962:73 [ref. 13277]). •Synonym of Strongylura marina (Walbaum 1792) -- (Mees 1962 [ref. 13277]). •Valid as Strongylura timucu (Walbaum 1792) -- (Collette 1968:191 [ref. 12725], Robins & Ray 1986:106 [ref. 23100], Cervigón 1991:230 [ref. 24472], Boschung 1992:84 [ref. 23239], Cervigón 1992:270 [ref. 23827], Greenfield & Thomerson 1997:95 [ref. 22947], McEachran & Fechhelm 1998:938 [ref. 23897], Castro-Aguirre et al. 1999:203 [ref. 24550], Keith et al. 2000:28 [ref. 24547], Marín 2000:70 [ref. 26252], Schmitter-Soto et al. 2000:154 [ref. 27754], Lovejoy & Collette 2001:329 [ref. 25232], Camargo & Isaac 2001:139 [ref. 27639], Feitoza et al. 2003:68 [ref. 26943], Collette 2003:1111 [ref. 26981], Collette 2003:9 [ref. 27306], Menezes & Figueiredo in Menezes et al. 2003:67 [ref. 27192], Smith et al. 2003:15 [ref. 27621], Banford et al. 2004:834 [ref. 27645], Nelson et al. 2004:99 [ref. 27807], Vaske et al. 2005:77 [ref. 36155], Miller 2006:204 [ref. 28615], Matamoros et al. 2009:15 [ref. 31241], García-Hernández et al. 2009:92 [ref. 37265], Page et al. 2013:103 [ref. 32708], Angulo et al. 2013:997 [ref. 33194], Di Dario et al. 2013:263 [ref. 37503], Smith-Vaniz & Jelks 2014:32 [ref. 33341], Marceniuk et al. 2017:39 [ref. 35324], Raz-Guzmán et al. 2018:345 [ref. 36407], Robins et al. 2018:191 [ref. 35886], Spier et al. 2018:8 [ref. 36103], Ribeiro et al. 2019:5 [ref. 36717], Collette & Bemis 2019:37 [ref. 37342], Caires et al. in Marceniuk et al. 2021:318 [ref. 38945], Angulo et al. 2023:65 [ref. 40676], Carvalho-Filho 2023:92 [ref. 40480], Martínez-Servín et al. 2023:111 [ref. 40726]). Current status: Valid as Strongylura timucu (Walbaum 1792). Belonidae. Distribution: Western Atlantic, southern Florida (U.S.A.) and Bahamas south to southern Brazil, including Gulf of Mexico and Caribbean Sea. Habitat: freshwater, brackish, marine.

vulgaris, Belone Fleming [J.] 1828:184 [A history of British animals; ref. 1339] Coasts of Great Britain. No types known. Based in part on Esox belone Linnaeus 1761. •Synonym of Belone belone (Linnaeus 1761), subspecies belone (Linnaeus 1761) -- (Collette & Parin 1970:11 [ref. 7415], Parin 1973:258 [ref. 7191], Collette 2003:2, 16 [ref. 27306]). Current status: Synonym of Belone belone (Linnaeus 1761). Belonidae. Habitat: brackish, marine.


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