Institute for Biodiversity Science and Sustainability
Online Version, Updated 6 November 2023

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Catalog of Fishes

Genera that contain: Garra [ 22 ] records

Ageneiogarra (subgenus of Garra) Garman [S.] 1912:114 [Memoirs of the Museum of Comparative Zoology v. 40 (pt 4); ref. 1544] Fem. Garra imberba Garman 1912. Type by monotypy. •Synonym of Garra Hamilton 1822 -- (Karaman 1971:232 [ref. 2560], Wu et al. 1977:372 [ref. 4807], Lévêque & Daget 1984:301 [ref. 6186], Wu & Wu 1992:292 [ref. 21205], Zhang et al. in Yue et al. 2000:237 [ref. 25272] as Agenneiogarra, Kottelat 2013:103 [ref. 32989], Sun et al. 2018:53 [ref. 35784] as distinct clade). •Valid as Ageneiogarra Garman 1912 -- (Yang et al. 2012:370 [ref. 33458]). Current status: Valid as Ageneiogarra Garman 1912. Cyprinidae: Labeoninae.

Brachygramma Day [F.] 1865:304 [Proceedings of the Zoological Society of London 1865 (pt 1); ref. 5295] Fem. Brachygramma jerdonii Day 1865. Type by monotypy. Also appeared in Day 1865:216 [ref. 1074]. The genus is feminine, not neuter; name based on the special condition of the lateral line. •Synonym of Garra Hamilton 1822 -- (Bănărescu 1997:20 [ref. 23274] in error). •Synonym of Amblypharyngodon Bleeker 1860 -- (Kottelat 2013:73 [ref. 32989] with genus as neuter). Current status: Synonym of Amblypharyngodon Bleeker 1860. Danionidae: Rasborinae.

Crossocheilus Kuhl [H.] & van Hasselt [J. C.] in van Hasselt 1823:132 [Algemeene Konst- en Letter-bode voor het Jaar II Deel (no. 35); ref. 5963] Masc. Crossocheilus oblongus Kuhl & van Hasselt 1823. Type by monotypy. Also spelled Crostocheilus in text and in 1824 [ref. 5104]. Crossocheilus selected by Kottelat (1987:371 [ref. 5962]). Crossocheilos is a misspelling; Crossochilus Günther 1868:71 [ref. 1990] is an unjustified emendation. •Synonym of Garra Hamilton 1822 -- (Behrens-Chapuis et al. 2015:197 [ref. 34083]). •Valid as Crossocheilus Kuhl & van Hasselt 1823 -- (Karaman 1971:236 [ref. 2560], Jayaram 1981:133 [ref. 6497], Roberts 1989:32 [ref. 6439], Chu & Cui in Chu & Chen 1989:243 [ref. 13584], Kottelat 1989:7 [ref. 13605], Ataur Rahman 1989:140 [ref. 24860], Menon & Jacob 1996:62 [ref. 22482], Doi 1997:20 [ref. 22832], Zhang & Chen 1997:321 [ref. 23731], Yang & Winterbottom 1998:49 [ref. 23307], Chen in Chen 1998:171 [ref. 23556], Su et al. 2000:215 [ref. 25281], Zhang et al. in Yue et al. 2000:207 [ref. 25272], Ataur Rahman 2003:159 [ref. 31338], Costa & Vono 2009:36 [ref. 30258], Zheng 2010:3 [ref. 30961], Zhu et al. 2011:46 [ref. 31305], Kottelat & Tan 2011:195 [ref. 31486], Yang et al. 2012:370 [ref. 33458], Kottelat 2013:89 [ref. 32989], Ciccotto & Page 2016:474 [ref. 34501], Zhang et al. 2016:108 [ref. 34477] with author as Hasselt, Ciccotto et al. 2017:270 [ref. 35402], Iqbal et al. 2017:1121 [ref. 35858]). Current status: Valid as Crossocheilus Kuhl & van Hasselt 1823. Cyprinidae: Labeoninae.

Discognathichthys Bleeker [P.] 1860:423 [Natuurkundig Tijdschrift voor Nederlandsch Indië v. 20 (no. 3); ref. 370] Masc. Discognathus variabilis Heckel 1843. Type by subsequent designation. Apparently appeared first in key, without species. Species added by Bleeker 1860:111, 128 [ref. 380] and type designated (p. 128). •Synonym of Garra Hamilton 1822 -- (Karaman 1971:232 [ref. 2560], Roberts 1989:40 [ref. 6439], Krupp & Schneider 1989:367 [ref. 13651], Kottelat 2013:103 [ref. 32989]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Discognathus Heckel [J. J.] 1843:1027 [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Masc. Discognathus variabilis Heckel 1843. Type by subsequent designation. Type designated by Bleeker 1863:24 [ref. 4859], 1863:262 [ref. 403], and 1863:192 [ref. 397]. •Synonym of Garra Hamilton 1822 -- (Karaman 1971:232 [ref. 2560], Wu et al. 1977:372 [ref. 4807], Lévêque & Daget 1984:301 [ref. 6186], Roberts 1989:40 [ref. 6439], Krupp & Schneider 1989:367 [ref. 13651], Wu & Wu 1992:292 [ref. 21205], Zhang et al. in Yue et al. 2000:237 [ref. 25272], Kottelat 2013:103 [ref. 32989]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Discolabeo Fowler [H. W.] 1937:210 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 89; ref. 1425] Masc. Discolabeo fisheri Fowler 1937. Type by original designation (also monotypic). •Valid as Discolabeo Fowler 1937 -- (Chen et al. in Pan et al. 1991:183 [ref. 23876], Zhang et al. 2016:110 [ref. 34477]). •Synonym of Garra Hamilton 1822 -- (Roberts 1989:40 [ref. 6439], Kottelat 2013:103 [ref. 32989]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Garra (subgenus of Cyprinus) Hamilton [F.] 1822:343, 393 [An account of the fishes found in the river Ganges; ref. 2031] Fem. Cyprinus (Garra) lamta Hamilton 1822. Type by subsequent designation. Type designated by Bleeker 1863:192 [ref. 397], 1863:262 [ref. 403] or 1863:24 [ref. 4859]. •Valid as Garra Hamilton 1822 -- (Karaman 1971:232 [ref. 2560], Wu et al. 1977:372 [ref. 4807], Jayaram 1981:134 [ref. 6497], Lévêque & Daget 1984:301 [ref. 6186], Kottelat 1985:262 [ref. 11441], Chu & Cui 1987 [ref. 19903], Roberts 1989:40 [ref. 6439], Chu & Cui in Chu & Chen 1989:269 [ref. 13584], Kottelat 1989:8 [ref. 13605], Ataur Rahman 1989:141 [ref. 24860], Chen et al. in Pan et al. 1991:180 [ref. 23876], Krupp & Schneider 1989:367 [ref. 13651], Lévêque in Lévêque et al. 1990:282 [ref. 21589], Wu & Wu 1992:292 [ref. 21205], Rema Devi 1993:244 [ref. 23727], Poll & Gosse 1995:165 [ref. 24781], Doi 1997:21 [ref. 22832], Chen in Chen 1998:175 [ref. 23556], Zhang et al. in Yue et al. 2000:237 [ref. 25272], Ataur Rahman 2003:160 [ref. 31338], Kullander & Fang 2004:257 [ref. 27857], Zhang 2005:9 [ref. 28349], Zhang 2005:138 [ref. 28426], Zhou et al. 2005:445 [ref. 28432], Zhang 2006:447 [ref. 28912], Proudlove 2006:66 [ref. 28992], Stiassny & Getahun 2007:51 [ref. 29119], De Weirdt et al. 2007:533 [ref. 30024], Li et al. 2008:62 [ref. 29588], Vishwanath & Linthoingambi 2008:101 [ref. 30107], Krupp & Budd 2009:120 [ref. 30148], Nebeshwar et al. 2009:197 [ref. 30177], Chen et al. 2009:438 [ref. 30322], Coad 2010:180 [ref. 36620], Habteselassie et al. 2010:43 [ref. 30774], Zheng 2010:3 [ref. 30961], Zhu et al. 2011:46 [ref. 31305], Kurup & Radhakrishnan 2011:220 [ref. 31583], Nebeshwar et al. 2012:2353 [ref. 31816], Shangningam & Vishwanath 2012:1 [ref. 31861], Zhang & Zhou 2012:22 [ref. 32322], Yang et al. 2012:362 [ref. 33458], Lalronunga et al. 2013:4368 [ref. 32715], Tamang 2013:31 [ref. 32803], Kottelat 2013:103 [ref. 32989], Nebeshwar & Vishwanah 2013:97 [ref. 33006], Arunachalam et al. 2013:62 [ref. 33195], Arunachalam et al. 2013:121 [ref. 33196], Lothongkham et al. 2014:543 [ref. 33283], Arunachalam et al. 2014:58 [ref. 33404], Arunachalam & Nandagopal 2014:43 [ref. 33405], Hamidan et al. 2014:223 [ref. 33626], Vilasri et al. 2014:19 [ref. 33693], Nebeshwar & Vishwanath 2015:305 [ref. 33758], Thoni & Mayden 2015:284 [ref. 33873], Sayyadzadeh et al. 2015:75 [ref. 34042], Shangningam & Vishwanath 2015:263 [ref. 34052], Jouladeh-Roudbar et al. 2015:871 [ref. 34062], Nguyen et al. 2015:893 [ref. 36430], Mousavi-Sabet & Eagderi 2016:45 [ref. 34341], Zheng et al. 2016:299 [ref. 34363], Gurumayum & Kosygin 2016:84 [ref. 34491], Yu et al. 2016:295 [ref. 34494], Ciccotto & Page 2016:475 [ref. 34501], Zhang et al. 2016:110 [ref. 34477], Esmaeili et al. 2016:82 [ref. 34620], Lyon et al. 2016:79 [ref. 34670], Thoni et al. 2016:115 [ref. 34747], Freyhof 2016:496 [ref. 34805], Das et al. 2016:52 [ref. 36383], Kangrang et al. 2016:335 [ref. 34836], Rahman et al. 2016:173 [ref. 34903], Mousavi-Sabet et al. 2016:133 [ref. 34988], Roni et al. 2017:367 [ref. 35205], Esmaeili et al. 2017:46 [ref. 35254], Nebeshwar & Vishwanath 2017:17 [ref. 35484], Roni & Vishwanath 2017:133 [ref. 35625], Rayamajhi & Arunachalam 2017:402 [ref. 35691], Premananda et al. 2017:191 [ref. 35861], Çiçek et al. 2018:5 [ref. 36267], Deng et al. 2018:94 [ref. 36132], Esmaeili et al. 2018:26 [ref. 36089], Pichler et al. 2018:[1] [ref. 35892], Roni & Vishwanath 2018:263 [ref. 35749], Sun et al. 2018:49 [ref. 35784], Yoğurtçuoğlu et al. 2018:281 [ref. 36205], Gong et al. 2018:367 [ref. 36379], Gong et al. 2018:857 [ref. 36387], Moyon & Arunkumar 2018:107 [ref. 36388], Tamang et al. 2019:59 [ref. 36496], Roni et al. 2019:545 [ref. 36796], Arunkumar & Moyon 2019:285 [ref. 36801], Rath et al. 2019:487 [ref. 36896], Mousavi-Sabet et al. 2019:223 [ref. 36979], Page et al. 2019:1 [ref. 37084], Shangningam et al. 2019:148 [ref. 37109], Moritz et al. 2019:311 [ref. 37196], Moyon & Arunkumar 2020:32 [ref. 37218], Kirchner et al. 2020:521 [ref. 37393], Ezung et al. 2020:1333 [ref. 37760], Zamani-Faradonbe et al. 2021:1 [ref. 38079], Bayçelebi et al. 2021:169 [ref. 38238], Ezung et al. 2021:18619 [ref. 38336], Kosygin et al. 2021:325 [ref. 39690], Zamani-Faradonbe et al. 2021:380 [ref. 38714], Shangningam et al. 2021:1 [ref. 38740], Mousavi-Sabet et al. 2022:419 [ref. 39728], Jadhav et al. 2022:1 [ref. 39774], Surachita et al. 2023:1 [ref. 40063], Tangjitjaroen et al. 2023:375 [ref. 40278], Cao et al. 2023:1 [ref. 40340], Ngangbam & Lithoingambi 2023:1 [ref. 40352], Sayyadzadeh et al. 2023:334 [ref. 40780], Thonbamliu et al. 2023:15 [ref. 40353], Thoidingjam et al. 2023:611 [ref. 40356], Zirkunga et al. 2023:65 [ref. 40518], Nebeshwar et al. 2024:189 [ref. 40865]). Current status: Valid as Garra Hamilton 1822. Cyprinidae: Labeoninae.

Hemigarra Karaman [M. S.] 1971:241 [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 67; ref. 2560] Fem. Tylognathus elegans Günther 1868. Type by original designation (also monotypic). •Probably a synonym of Hemigrammocapoeta Pellegrin 1927 -- (Krupp & Schneider 1989:372 [ref. 13651], Coad 1995:18 [ref. 23608]). •Synonym of Garra Hamilton 1822 -- (Behrens-Chapuis et al. 2015:197 [ref. 34083], Freyhof 2016:496 [ref. 34805]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Hemigrammocapoeta Pellegrin [J.] 1927:34 [Bulletin de la Société Zoologique de France v. 52 (no. 1); ref. 5042] Fem. Hemigrammocapoeta culiciphaga Pellegrin 1927. Type by monotypy. •Valid as Hemigrammocapoeta Pellegrin 1927 -- (Krupp & Schneider 1989:371 [ref. 13651], Coad 2010:186 [ref. 36620], Jouladeh-Roudbar et al. 2015:873 [ref. 34062], Küçük et al. 2015:359 [ref. 34064]). •Synonym of Garra Hamilton 1822 -- (Behrens-Chapuis et al. 2015:197 [ref. 34083]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Horalabiosa Silas [E. G.] 1954:28 [Records of the Indian Museum (Calcutta) v. 51 (pt 1); ref. 4025] Fem. Horalabiosa joshuai Silas 1954. Type by original designation (also monotypic). •Status uncertain -- (Bănărescu 1997:22 [ref. 23274]). •Valid as Horalabiosa Silas 1954 -- (Johnson & Soranam 2001:392 [ref. 26173]). •Synonym of Garra Hamilton 1822 -- (Kottelat 2013:103 [ref. 32989]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Iranocypris Bruun [A. F.] & Kaiser [E. W.] 1944:4 [Danish Scientific Investigations in Iran (Copenhagen) Part 4; ref. 674] Fem. Iranocypris typhlops Bruun & Kaiser 1944. Type by original designation (also monotypic). Journal dates to 1948, preprint appeared in 1944? •Valid as Iranocypris Bruun & Kaiser 1944 -- (Banister & Bunni 1980:151 [ref. 175], Proudlove 2006:63 [ref. 28992], Jouladeh-Roudbar et al. 2015:873 [ref. 34062]). •Synonym of Garra Hamilton 1822 -- (Sayyadzadeh et al. 2015:77 [ref. 34042], Mousavi-Sabet & Eagderi 2016:45 [ref. 34341]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Lissorhynchus Bleeker [P.] 1860:422 [Natuurkundig Tijdschrift voor Nederlandsch Indië v. 20 (no. 3); ref. 370] Masc. Gobio lissorhynchus McClelland 1839. Type by subsequent monotypy. Apparently appeared first in key, without species, as Lissorhynchus. One species included in Bleeker 1860:85-86 [ref. 380]. Type by subsequent monotypy. Misspelled Lissorhynchos by Bleeker 1860:86, Lissorhynchus on p. 85. •Synonym of Garra Hamilton 1822 -- (Karaman 1971:232 [ref. 2560], Roberts 1989:40 [ref. 6439], Kottelat 2013:102 [ref. 32989]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Mayoa Day [F.] 1870:553 [Proceedings of the Zoological Society of London 1869 (pt 3) (art. 3) (for 25 Nov. 1869); ref. 1077] Fem. Mayoa modesta Day 1870. Type by monotypy. •Synonym of Garra Hamilton 1822 -- (Karaman 1971:232 [ref. 2560], Lévêque & Daget 1984:301 [ref. 6186], Bănărescu 1997:23 [ref. 23274], Kottelat 2013:103 [ref. 32989]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Neotylognathus (subgenus of Tylognathus) Kosswig [C.] 1950:409 [Zoologischer Anzeiger v. 145; ref. 5022] Masc. Tylognathus (Neotylognathus) klatti Kosswig 1950. Type by monotypy. •Synonym of Hemigrammocapoeta Pellegrin 1927, but a valid subgenus Neotylognathus as described -- (Karaman 1971:236 [ref. 2560]). •Synonym of Garra Hamilton 1822. Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Parapsilorhynchus Hora [S. L.] 1921:13 [Records of the Indian Museum (Calcutta) v. 22 (pt 1); ref. 2201] Masc. Psilorhynchus tentaculatus Annandale 1919. Type by original designation. Type designated on p. 13, not P. discophorus of authors. See Das 2003:368 [ref. 27706] as Parasilorhynchus (also as Parapsilorhynchus). •Synonym of Garra Hamilton 1822 -- (Roberts 1989:40 [ref. 6439]). •Valid as Parapsilorhynchus Hora 1921 -- (Jayaram 1981:132 [ref. 6497], Yazdani & Singh 1991:43 [ref. 24004], Singh 1994:282 [ref. 22587], Rema Devi & Menon 1995 [ref. 22106], Bănărescu 1997:25 [ref. 23274], Baliarsingh et al. 2017:22 [ref. 35577], Baliarsingh & Kosygin 2017:44 [ref. 35747], Jadhav et al. 2020:563 [ref. 37394]). Current status: Valid as Parapsilorhynchus Hora 1921. Cyprinidae: Labeoninae.

Phreatichthys Vinciguerra [D.] 1924:240 [Annali del Museo Civico di Storia Naturale `Giacomo Doria' v. 51 (for 1923); ref. 4523] Masc. Phreatichthys andruzzii Vinciguerra 1924. Type by monotypy. •Valid as Phreatichthys Vinciguerra 1924 -- (Lévêque & Daget 1984:330 [ref. 6186], Poll & Gosse 1995:161 [ref. 24781], Proudlove 2006:59 [ref. 28992]). •Synonym of Garra Hamilton 1822. Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Placocheilus Wu [H.-W.] in Wu, Lin, Chen, Chen & He 1977:382 [The cyprinid fishes of China v. 2; ref. 4807] Masc. Discognathus caudofasciatus Pellegrin & Chevey 1936. Type by original designation (also monotypic). Misspelled Placoheilus in index (p. 593). Zhang et al. 2002:209 [ref. 27464] indicate that Wu based this genus on a misidentified type species which Zhang et al. 2002 describe as a new species robustus. •Synonym of Garra Hamilton 1822 -- (Li et al. 2008:62 [ref. 29588], Lothongkham et al. 2014:543 [ref. 33183]). •Valid as Placocheilus Wu 1977 -- (Cui & Li 1984 [ref. 8236], Chu & Cui in Chu & Chen 1989:278 [ref. 13584], Chen in Chen 1998:180 [ref. 23556], Zhang et al. in Yue et al. 2000:265 [ref. 25272], Zheng et al. 2002:209 [ref. 27464], Zhang 2005:139 [ref. 28426], Zhou et al. 2005:445 [ref. 28432], Zheng 2010:3 [ref. 30961], Zhu et al. 2011:46 [ref. 31305], Chen et al. 2012:307 [ref. 31862], Zhang & Zhou 2012:22 [ref. 32322], Kottelat 2013:141 [ref. 32989], Zhang et al. 2016:113 [ref. 34477]). Current status: Valid as Placocheilus Wu 1977. Cyprinidae: Labeoninae.

Platycara McClelland [J.] 1838:944, 947 [Journal of the Asiatic Society of Bengal v. 7 (for Nov. 1838); ref. 2924] Neut. Balitora brucei Gray 1830. Type by being a replacement name. Apparently appeared first as above in late 1838. Also in McClelland 1839:246, 299, 427, 944, 947 [ref. 2923]; as published in ref. 2924 it is less clear that it is a replacement name, and Jordan and Fowler both regard P. nasuta McClelland as type as first designated by Jordan 1919:195 [ref. 2410]. •Synonym of Garra Hamilton 1822 -- (Karaman 1971:232 [ref. 2560], Lévêque & Daget 1984:301 [ref. 6186], Roberts 1989:40 [ref. 6439], Bănărescu 1997:25 [ref. 23274], Kottelat 2013:103 [ref. 32989]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Tylognathoides (subgenus of Tylognathus) Tortonese [E.] 1938:15 [of separate] [Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino. (Ser. 3) v. 46 (no. 85); ref. 10283] Masc. Tylognathus festai Tortonese 1939. Type by monotypy. •Valid as Tylognathoides Tortonese 1938 -- (Karaman 1971:239 [ref. 2560]). •Probably a synonym of Hemigrammocapoeta Pellegrin 1927 -- (Bănărescu 1997:28 [ref. 23274]). •Probably a synonym of Garra Hamilton 1822. Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Tylognathus Heckel [J. J.] 1843:1027 [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Masc. Varicorhinus diplostomus Heckel 1838. Type by subsequent designation. Type designated by Bleeker 1863:194 [ref. 397] or 1863:25 [ref. 4859]. •Synonym of Crossocheilus Kuhl & van Hasselt 1823 -- (Karaman 1971:236 [ref. 2560]). •Synonym of Labeo Cuvier 1816 -- (Bănărescu 1997:28 [ref. 23274]). •Synonym of Lobocheilus van Hasselt 1823 [in error] -- (Zhang et al. in Yue et al. 2000:197 [ref. 25272]). •Synonym of Bangana Hamilton 1822 -- (Zhang & Chen 2006:43 [ref. 28775], Kottelat 2013:74 [ref. 32989], Kottelat 2017:494 [ref. 35262]). •Synonym of Garra Hamilton 1822 -- (Behrens-Chapuis et al. 2015:197 [ref. 34083]). Current status: Synonym of Bangana Hamilton 1822. Cyprinidae: Labeoninae.

Typhlogarra Trewavas [E.] 1955:552 [Annals and Magazine of Natural History (Series 12) v. 8 (no. 91) (art. 67); ref. 4454] Fem. Typhlogarra widdowsoni Trewavas 1955. Type by original designation (also monotypic). •Valid as Typhlogarra Trewavas 1955 -- (Bănărescu 1997:28 [ref. 23274], Proudlove 2006:64 [ref. 28992], Coad 2010:197 [ref. 36620]). •Synonym of Garra Hamilton 1822 -- (Sayyadzadeh et al. 2015:88 [ref. 34042], Behrens-Chapuis et al. 2015:197 [ref. 34083]). Current status: Synonym of Garra Hamilton 1822. Cyprinidae: Labeoninae.

Vinagarra Nguyen [V. H.] & Bui [T. A.] 2009:52 [Science and Technology journal of Agriculture & Rural Development (Nông Nghiêp & Phát Triên Nông Thôn) 2009; ref. 33206] Fem. Garra laichowensis Nguyen & Doan 1969. Type by original designation (also monotypic). Name spelled as Vingarra and Vinagrra as well as Vinagarra -- this name correctly formed (Endruweit 2014:148 [ref. 33205]). •Valid as Vinagarra Nguyen & Bui 2009 -- (Endruweit 2014:147 [ref. 33205]). Current status: Valid as Vinagarra Nguyen & Bui 2009. Cyprinidae: Labeoninae.


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