Institute for Biodiversity Science and Sustainability
Online Version, Updated 6 November 2023

SEARCH RESULTS FROM THE

Catalog of Fishes

Species that contain: Himantura and uarnak [ 11 ] records

ellioti, Trygon Blyth [E.] 1860:41 [Journal of the Asiatic Society of Bengal v. 29 (no. 1); ref. 16970] Calcutta fish market, India. No types known. •Questionably a synonym of Himantura uarnak (Gmelin 1789) -- (Kottelat 2013:27 [ref. 32989]). Current status: Synonym of Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Habitat: marine.

forskohlii, Raja Billberg [G. J.] 1833:51 [Linnéska Samfundets Handlingar v. 1 (for 1832); ref. 34146] Red Sea. No types known. Unnecessary replacement name for Raja uarnak Forsskål 1775; see Kullander 2016:112 [ref. 34147]. •Synonym of Himantura uarnak (Gmelin 1789) -- (Kullander 2016:107 [ref. 34147]). Replacement Name. Current status: Synonym of Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Habitat: marine.

oxyrhynchus, Trygon (Himantura) Sauvage [H.-E.] 1878:91 [6] [Bulletin de la Société philomathique de Paris (7th Série) v. 2; ref. 19968] Saigon, southern Vietnam. Holotype (unique): MNHN 0000-9639. Type catalog: Bertin 1939:86 [ref. 20739], Séret & McEachran 1987:19 [ref. 9312]. Specific name is a noun. •Synonym of Himantura uarnak (Forsskål 1775) -- (Kottelat 1984:794 [ref. 11440]). •Valid as Himantura oxyrhyncha (Sauvage 1878) -- (Compagno 1999:496 [ref. 25589], Compagno in Randall & Lim 2000:583 [ref. 25122], Deynat & Fermon 2001:163 [ref. 25563], Vidthayanon & Roberts 2006:124 [ref. 28695], Last el al. 2010:204 [ref. 32461], Naylor et al. 2012:71 [ref. 32267] see remarks, Weigmann 2016:136 [ref. 34211]). •Valid as Himantura oxyrhynchus (Sauvage 1878) -- (Last & Compagno 1999:1503 [ref. 24639], Kottelat 2013:26 [ref. 32989]). •Valid as Fluvitrygon oxyrhynchus (Sauvage 1878) -- (Last et al. 2016:360 [ref. 34581] as oxyrhyncha, Last et al. 2016:543 [ref. 35010], Iqbal et al. 2017:83 [ref. 35489], Taki et al. 2021:38 [ref. 39830]). Current status: Valid as Fluvitrygon oxyrhynchus (Sauvage 1878). Dasyatidae: Urogymninae. Distribution: Eastern Indian Ocean, western Pacific: Cambodia, Thailand, Sumatra and Borneo. Habitat: freshwater, brackish.

punctata, Trygon Günther [A.] 1870:474 [Catalogue of the fishes in the British Museum v. 8; ref. 1995] ? East Indian Archipelago. Holotype (unique): BMNH 1953.8.10.15. •Synonym of Himantura uarnak (Gmelin 1789) [author also seen as Forster 1775] -- (Last & Compagno 1999:1491 [ref. 24639], Bonfil & Abdallah 2004:49 [ref. 27735], Kottelat 2013:27 [ref. 32989] with question). Current status: Synonym of Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Habitat: freshwater, brackish, marine.

scherit, Raia Bonnaterre [J. P.] (ex Fabricius, Forsskål) 1788:6 [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] Red Sea. No types known. Based on Raja omm es scherit of Fabricius 1775. Not used as valid after 1899, therefore reversal of priority in favor of Raja uarnak Gmelin 1789 is followed (Art. 23.9). •Synonym of Himantura uarnak (Gmelin 1789) -- (Fricke 2008:12 [ref. 30182], Kottelat 2013:27 [ref. 32989]). Nomen Oblitum. Current status: Synonym of Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Habitat: brackish, marine.

schoukie, Raja Fabricius [J. C.] in Niebuhr (ex Forsskål) 1775:ix [Descriptiones animalium (Forsskål); ref. 1351] Jeddah, Saudi Arabia, Red Sea; Suaken, Sudan, 19°07'N 37°20'E. No types known. Name considered available, with author as Fabricius, by Fricke 2008:11 [ref. 30182]. Spelled schoukia in Bonnaterre 1788:6 [ref. 4940]. •Unidentifiable in Dasyatidae – (Fricke 2008:11 [ref. 30182]). •Questionably a synonym of Himantura uarnak (Gmelin 1789) -- (Kottelat 2013:26 [ref. 32989] with author as Forsskål). Dasyatidae. Habitat: marine.

toshi, Himantura Whitley [G. P.] 1939:258 [Australian Zoologist v. 9 (pt 3); ref. 4695] Clarence River Estuary, New South Wales, Australia. Holotype (unique): AMS IA.39. •Synonym of Himantura uarnak (Forsskål 1775) -- (Paxton et al. 1989:42 [ref. 12442]). •Valid as Himantura toshi Whitley 1939 -- (Last & Stevens 1994:405 [ref. 23873], Allen 1997:48 [ref. 23977], Last & Compagno 1999:1503 [ref. 24639], Compagno 1999:496 [ref. 25589], Hutchins 2001:15 [ref. 25847], Paxton et al. 2006:194 [ref. 28995], Last et al. 2008:303 [ref. 29696], Schwartz 2008:38 [ref. 36773], Allen & Erdmann 2012:63 [ref. 31980], Larson et al. 2013:19 [ref. 32988], Weigmann 2016:137 [ref. 34211], Allen & Erdmann 2024:66 [ref. 40796]). •Valid as Maculabatis toshi (Whitley 1939) -- (Last et al. 2016:362 [ref. 34581], Last et al. 2016:66 [ref. 34669], Last et al. 2016:581 [ref. 35010], White et al. 2017:234 [ref. 35860], White & Ko'ou 2018:56 [ref. 35922]). Current status: Valid as Maculabatis toshi (Whitley 1939). Dasyatidae: Urogymninae. Distribution: Indo-West Pacific: southern New Guinea and Western Australia, Northern Territory and Queensland (northern Australia). Habitat: brackish, marine.

tutul, Himantura Borsa [P.], Durand [J.-D.], Shen [K.-N.], Alyza [I. A.], Solihin [D. D.] & Berrebi [P.] 2013:89, Fig. 5 [Comptes Rendus Biologies v. 336; ref. 32736] Mkoani fish landing, Pemba Island, Tanzania. Holotype: MNHN ICTI-5184 (2 sections of tail). Paratypes: CSIRO H 5284.05 (2). Plus non-type specimens. See also Borsa 2016:[1] 1170 [ref. 35022] and Weigmann 2016:4 [ref. 35023]. This species has been confused with Himantura leoparda Manjaji-Matsumoto & Last 2008 (Borsa 2016:[2]1171 [ref. 35022]). •Valid as Himantura tutul Borsa, Durand, Shen, Alyza, Solihin & Berrebi 2013 -- (Borsa 2016:[4] 1173 [ref. 35022], Miesen et al. 2016:80 [ref. 34492], Fernando et al. 2019:208 [ref. 36642], Kumar et al. 2020:[2] [ref. 37567]). •Synonym of Himantura uarnak (Gmelin 1789) -- (Last et al. 2016:563 [ref. 35010], Weigmann 2016:[4] 1179 [ref. 34211] see remarks, Weigmann 2016:1177 [ref. 35023]). Current status: Synonym of Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Distribution: Indian Ocean: East Africa east to Indonesia (if valid). Habitat: brackish, marine.

uarnak, Raja Gmelin [J. F.] (ex Fabricius, Forsskål) 1789:1509 [Caroli a Linné ... Systema Naturae per regna tria naturae v. 1 (pt 3); ref. 18139] [No locality stated] Red Sea. No types known. Types given by Klausewitz 1960:294 [ref. 21330] are specimens used by Rüppell 1837 and are not types of uarnak Gmelin. Based on Raja sephen var. b of Fabricius [ex Forsskål, in Niebuhr] 1775:18, viii [ref. 1351]. Sometimes misspelled uarnack. Nomen protectum according to Fricke 2008:12 [ref. 30182]. Lectotype (SMF ) designated by Klausewitz 1960:294 [ref. 21330], based on Rüppell specimen; lectotype invalid as specimen was collected in 1828, after Gmelin's description. Neotype (CAS 247241) designated by Borsa et al. 2020:[5] [ref. 37984], but specimen too small to sufficiently characterize the species, therefore neotype invalid; ref. 37984 is also lacking fixed content and layout, and neotype designation therefore unavailable according to ICZN 8.1.3.2. •Valid as Dasyatis uarnak (Gmelin 1789) [author usually given as Forsskål 1775] -- (Dor 1984:18 [ref. 29757], Compagno & Roberts 1984:4 [ref. 6167], Mohsin & Ambak 1996:77 [ref. 2969]). •Valid as Himantura uarnak (Gmelin 1789) -- (Krefft & Stehmann 1973:72 [ref. 7167], Nakaya in Masuda et al. 1984:16 [ref. 6441], McEachran & Capapé in Whitehead et al. 1984:201 [ref. 13675], Compagno 1986:139 [ref. 5648], Allen & Swainston 1988:28 [ref. 25477], Paxton et al. 1989:42 [ref. 12442], Talwar & Jhingran 1991:46 [ref. 20764], Last & Stevens 1994:406 [ref. 23873], Goren & Dor 1994:5 [ref. 25356], Randall 1995:46 [ref. 22896], Chen et al. 1997:9 [ref. 26476], Larson & Williams 1997:346 [ref. 23967], Basusta et al. 1998:65 [ref. 23545], Golani 1998:97 [ref. 27567], Yoshigou & Yoshino 1999:39 [ref. 23925], Myers 1999:39 [ref. 23965], Fricke 1999:32 [ref. 24106], Morón et al. 1999:151 [ref. 24253], Last & Compagno 1999:1491 [ref. 24639], Vossoughi & Vosoughi 1999:302 [ref. 25467], Johnson 1999:718 [ref. 25471], Compagno 1999:496 [ref. 25589], Nakabo 2000:178 [ref. 25086], Compagno in Randall & Lim 2000:583 [ref. 25122], Iwatsuki et al. 2000:96 [ref. 26368], Deynat & Fermon 2001:161 [ref. 25563], Hutchins 2001:15 [ref. 25847], Bilecenoğlu et al. 2002:26 [ref. 26753], Nakabo 2002:178 [ref. 26001], Sujatha 2002:157 [ref. 26899], Manilo & Bogorodsky 2003:S94 [ref. 27377], Bonfil & Abdallah 2004:49 [ref. 27735], Heemstra & Heemstra 2004:83 [ref. 28072], Compagno et al. 2005:73 [ref. 29145], Golani 2005:14 [ref. 37112], Paxton et al. 2006:194 [ref. 28995], Fricke et al. 2007:20 [ref. 29533], Manjaji-Matsumoto & Last 2008:293 [ref. 29695], Last et al. 2008:303 [ref. 29696], Fricke 2008:12 [ref. 30182], Schwartz 2008:38 [ref. 36773], Last et al. 2010:214 [ref. 32461] with author as Forsskål, Allen & Erdmann 2012:64 [ref. 31980], Naylor et al. 2012:70 [ref. 32267] see remarks, Borsa et al. 2013:87 [ref. 32736], Larson et al. 2013:19 [ref. 32988] with author as Forsskål, Kottelat 2013:27 [ref. 32989], Ebert et al. 2013:362 [ref. 33045], Psomadakis et al. 2015:102 [ref. 34104], Weigmann 2016:138 [ref. 34211], Miesen et al. 2016:80 [ref. 34492], Last et al. 2016:361 [ref. 34581], Last et al. 2016:387 [ref. 34638], Last et al. 2016:563 [ref. 35010], Motomura et al. 2017:25 [ref. 35490], Ali et al. 2018:304 [ref. 36183], Bogorodsky & Randall 2018:[20] 258 [ref. 36492], Fricke et al. 2018:26 [ref. 35805], Golani & Fricke 2018:15 [ref. 36273] with author as Forsskål, Kumar et al. 2018:297 [ref. 36178], Suresh et al. 2018:49 [ref. 36495], Almeida & Biscoito 2019:173 [ref. 37302], Eagderi et al. 2019:9 [ref. 37020], Zajonz et al. 2019:60 [ref. 36871], Bariche & Fricke 2020:116 [ref. 37515], Golzarianpour et al. 2020:330 [ref. 37674], Habib & Islam 2020:Supplementary table p. 3 [ref. 38321], Psomadakis et al. 2020:206 [ref. 37272], Serena et al. 2020:502 [ref. 37963], Bogorodsky et al. 2021:520 [ref. 38242], Ebert et al. 2021:92 [ref. 38234], Golani 2021:11 [ref. 38303] with author as Forsskål 1775, Haroon & Kibria 2021:109 [ref. 39241] with author as Forsskål, Kovačić et al. 2021:17 [ref. 39014], Barone et al. 2022:58 [ref. 39527], Hsu et al. 2022:3 [ref. 39567], Manjaji-Matsumoto et al. 2022:604 [ref. 39674], Zajonz et al. 2022:25 [ref. 40522] with author as Forsskål 1775, Allen & Erdmann 2024:66 [ref. 40796]). Nomen Protectum. Current status: Valid as Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Distribution: Red Sea; Indo-West Pacific: Eastern Cape and KwaZulu-Natal (South Africa), East Africa, Persian Gulf, Socotra (Yemen), Seychelles, Madagascar and Mauritius (Mascarenes) east to Philippines; Mediterranean Sea (Red Sea immigrant). Habitat: freshwater, brackish, marine.

uarnak, Raja sephen var. Forsskål [P. S.] 1775:18, viii [Descriptiones animalium (Forsskål); ref. 1351] Red Sea. No types known. As a variety on p. viii, local name only on p. 18. Types given by Klausewitz 1960:294 [ref. 21330] are specimens used by Rüppell 1837 and are not types of uarnak Forsskål. Not available, published as vernacular Arabic name only. Name made available by Gmelin 1789:1509 [ref. 18139]; see Fricke 2008:12 [ref. 30182]. •Valid as Dasyatis uarnak (Forsskål 1775) -- (Dor 1984:18 [ref. 29757], Compagno & Roberts 1984:4 [ref. 6167], Mohsin & Ambak 1996:77 [ref. 27969]). •Synonym of Himantura uarnak (Gmelin 1789). Current status: Synonym of Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Distribution: Red Sea, Indo-West Pacific, including Red Sea; Mediterranean Sea (Red Sea immigrant); Japan south to northern Australia (with author as Forsskål 1775). Habitat: brackish, marine.

variegatus, Trygon McClelland [J.] 1841:60, Pl. 2 (fig. 2) [Calcutta Journal of Natural History v. 1; ref. 17619] Salt lake near Calcutta, India. No types known. •Synonym of Himantura uarnak (Gmelin 1789) -- (Kottelat 2013:27 [ref. 32989]). Current status: Synonym of Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Habitat: marine.


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