Species that contain: Lefua and costata [ 7 ] records

andrewsi, Lefua Fowler [H. W.] 1922:1 [American Museum Novitates No. 38; ref. 15339] Shing Lung Shan, eastern Tombs [now Hebei, Zunhua, ,40°11'N, 117°58'E], Hopeh Province, China. Holotype: AMNH 7974. Paratypes: AMNH 7990 (1). •Synonym of Lefua costata (Kessler 1876) -- (Berg 1949:887 [ref. 20661], Zhu 1989:21 [ref. 17744], Vasil'eva 1998:96 [ref. 23591], Choi et al. 2003:178 [ref. 26218], Kottelat 2006:55 [ref. 28831], Kottelat 2012:87 [ref. 32367]). Current status: Synonym of Lefua costata (Kessler 1876). Nemacheilidae. Habitat: freshwater.

coreanus, Elxis Jordan [D. S.] & Starks [E. C.] 1905:201, Fig. 7 [Proceedings of the United States National Museum v. 28 (no. 1391); ref. 2530] Gensan, Korea. Holotype: USNM 45243 (mixed with paratypes). Paratypes: MCZ 35960 [ex USNM 45243] (2), CAS-SU 8661 [not 8061] (77), USNM 45241-42 (249, 50), 45243 (now 11, including holotype), 45245 (1), 45248-49 (8, 14). Type catalog: Böhlke 1953:39 [ref. 12291]. •Synonym of Lefua costata (Kessler 1876) -- (Berg 1949:887 [ref. 20661], Kim 1997:560 [ref. 24032], Vasil'eva 1998:96 [ref. 23591], Kottelat 2006:55 [ref. 28831], Kottelat 2012:87 [ref. 32367]). Current status: Synonym of Lefua costata (Kessler 1876). Nemacheilidae. Habitat: freshwater.

costata, Diplophysa Kessler [K. F.] 1876:29, Pl. 3 (fig. 4) [v. 2 (pt 4); ref. 2595] Hu-lun [Dalai-Nor, Dalai-nor] Lake at 43°N [43°18'N, 116°39'E], Liaoning, Inner Mongolia, China, elevation 4200 feet. Holotype (unique): ZIN 2477. For remarks on sysematics see Kottelat 2006:55 [ref. 28831]. •Valid as Lefua costata (Kessler 1876) -- (Berg 1949:887 [ref. 20661], Wang 1984:81 [ref. 22951], Zhu 1989:20 [ref. 17744], Zhu 1995:107 [ref. 25213], Bogutskaya & Naseka 1996:48 [ref. 22798], Cheng & Zhou 1997:195 [ref. 26385], Kim 1997:285 [ref. 24032], Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:96 [ref. 23591], Kim et al. 2000:224 [ref. 25434], Pietsch et al. 2001:140 [ref. 25688], Wang et al. 2001:167 [ref. 26566], Kim & Park 2002:197 [ref. 25913], Bogutskaya et al. 2001:45 [ref. 26178], Choi et al. 2003:178, 518 [ref. 26218], Jang et al. 2003:119 [ref. 27681], Naseka & Bogutskaya 2004:284 [ref. 27838], Bogutskaya & Naseka 2004:115 [ref. 28183], Mihara et al. 2005:157 [ref. 28201], Kottelat 2006:55 [ref. 28831], Ocock et al. 2006:29 [ref. 28933], Bogutskaya et al. 2008:340 [ref. 30085], Kottelat 2012:87 [ref. 32367], Zhang et al. 2016:129 [ref. 34477], Hosoya et al. 2018:199 [ref. 36197], Ohgita & Matsunuma 2020:10 [ref. 37953], Du et al. 2021:4 [ref. 38600], Guo 2021:487 [ref. 38618], Dyldin et al. 2023:[9] [ref. 40328]). Current status: Valid as Lefua costata (Kessler 1876). Nemacheilidae. Distribution: East Asia: northern China, Russia, Korea and Mongolia; introduced in central Japan. Habitat: freshwater.

dixoni, Nemachilus Fowler [H. W.] 1899:181 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 51; ref. 17572] Tan Lan Ho River, tributary of Shan Lan Ho River, about 30 miles northeast of Lama-Miau or Dolon-nor, Pechili Province, China. Holotype (unique): ANSP 16393. Type catalog: Böhlke 1984:63 [ref. 13621]. Original genus is correctly Nemacheilus. •Synonym of Lefua costata (Kessler 1876) -- (Berg 1949:887 [ref. 20661], Zhu 1989:21 [ref. 17744], Vasil'eva 1998:96 [ref. 23591], Kottelat 2006:55 [ref. 28831], Kottelat 2012:87 [ref. 32367]). Current status: Synonym of Lefua costata (Kessler 1876). Nemacheilidae. Habitat: freshwater.

echigonia, Lefua Jordan [D. S.] & Richardson [R. E.] 1907:263, Fig. 1 [Proceedings of the United States National Museum v. 33 (no. 1570); ref. 10643] Nagaoka, Echigo, Japan. Holotype: CAS-SU 20164. Paratypes: USNM (1). Type catalog: Böhlke 1953:39 [ref. 12291]. •Synonym of Lefua costata (Kessler 1876), but a valid subspecies -- (Kawanabe & Mizuno 1989:400 [ref. 25598]). •Valid as Lefua echigonia Jordan & Richardson 1907 -- (Kuroda 1953:276 [ref. 39545], Okada 1961:546 [ref. 6363], Sawada in Masuda et al. 1984:59 [ref. 6441], Watanabe 1998:263 [ref. 23491], Nakabo 2000:277 [ref. 25086], Matsuura et al. 2000:168 [ref. 26000], Nakabo 2002:277 [ref. 26001], Mihara et al. 2005:157 [ref. 28201], Kottelat 2012:87 [ref. 32367], Hosoya et al. 2018:199 [ref. 36197], Ito et al. 2019:485 [ref. 36659]). Current status: Valid as Lefua echigonia Jordan & Richardson 1907. Nemacheilidae. Distribution: East Asia: Honshu Island (Japan). Habitat: freshwater.

nikkonis, Elxis Jordan [D. S.] & Fowler [H. W.] 1903:768, Fig. 1 [Proceedings of the United States National Museum v. 26 (no. 1332); ref. 2461] Chitose in Iburi, Hokkaido, Japan. Holotype: CAS-SU 7848. Paratypes or additional material: CAS-SU 7147 (1), 7485 (8). Type catalog: Böhlke 1953:39 [ref. 12291]. •Synonym of Lefua costata (Kessler 1876), but a valid subspecies -- (Kawanabe & Mizuno 1989:401 [ref. 25598], Pietsch et al. 2001:140 [ref. 25688]). •Valid as Lefua nikkonis (Jordan & Fowler 1903) -- (Okada 1961:549 [ref. 6363], Sawada in Masuda et al. 1984:59 [ref. 6441], Watanabe 1998:263 [ref. 23491], Nakabo 2000:277 [ref. 25086], Matsuura et al. 2000:169 [ref. 26000], Nakabo 2002:277 [ref. 26001], Mihara et al. 2005:157 [ref. 28201], Kottelat 2012:87 [ref. 32367], Dyldin & Orlov 2016:670 [ref. 34853], Hosoya et al. 2018:200 [ref. 36197], Ito et al. 2019:486 [ref. 36659], Ohgita & Matsunuma 2020:11 [ref. 37953], Dyldin et al. 2021:19 [ref. 38804], Dyldin & Orlov 2021:63 [ref. 38158], Dyldin et al. 2023:[9] [ref. 40328]). Current status: Valid as Lefua nikkonis (Jordan & Fowler 1903). Nemacheilidae. Distribution: East Asia: Hokkaido (Japan), Sakhalin Island (Russia). Habitat: freshwater.

pleskei, Octonema Herzenstein [S. M.] in Warpachowski & Herzenstein 1888:48, Pl. (fig. 5) [Trudy St.-Peterburgskogo Obscestva Estestvoispytatelej = Travaux de la Société des Naturalistes de St. Pétersbourg. v. 19 (no. 8) (for 1887); ref. 2152] Ilistaya [Lefu] River near Nikolajewka, Khanka Lake basin, Amur River drainage, Russia. Syntypes: BMNH 1891.10.7.73 [ex ZIN] (1), ZIN 7209 (orig. 5, now 4). •Synonym of Lefua costata (Kessler 1876) -- (Berg 1949:887 [ref. 20661], Zhu 1989:20 [ref. 17744], Bogutskaya & Naseka 1996:48 [ref. 22798], Vasil'eva 1998:96 [ref. 23591]). •Valid as Lefua pleskei (Herzenstein 1888) -- (Naseka & Bogutskaya 2004:284 [ref. 27838] dated 1887, Bogutskaya & Naseka 2004:115 [ref. 28183] dated 1887, Bogutskaya et al. 2008:340 [ref. 30085] dated 1887, Parin et al. 2014:103 [ref. 33547] dated 1887, Dyldin & Orlov 2016:671 [ref. 34853], Hosoya et al. 2018:199 [ref. 36197], Ohgita & Matsunuma 2020:11 [ref. 37953], Dyldin et al. 2021:20 [ref. 38804], Dyldin & Orlov 2021:63 [ref. 38158], Dyldin et al. 2023:[9] [ref. 40328]). Current status: Valid as Lefua pleskei (Herzenstein 1888). Nemacheilidae. Distribution: East Asia: Russia and Japan. Habitat: freshwater, brackish.