Species that contain: Leiognathus and equula [ 44 ] records

argentea, Equula De Vis [C. W.] 1884:542 [Proceedings of the Linnean Society of New South Wales v. 9 (pt 3); ref. 4899] Cape York, Queensland, Australia. Lectotype: QM I.1699. Paralectotypes: QM (1) missing. Subjectively invalid; secondarily preoccupied by Leiognathus argenteus Lacepède 1802, replaced by Leiognathus devisi Whitley 1929. Lectotype selected by Whitley 1932:105 [ref. 4675]. •Synonym of Leiognathus decorus (De Vis 1884) -- (Jones 1985:578 [ref. 21842] incorrectly with a holotype, Hoese & Bray 2006:1179 [ref. 29000]). •Possible synonym of Nuchequula gerreoides (Bleeker 1851) based on status of decorus. •Synonym of Eubleekeria splendens (Cuvier 1829) -- (Kottelat 2013:338 [ref. 32989], Chakrabarty & Sparks 2022:421 [ref. 39869]). 2°Homonym. Current status: Synonym of Eubleekeria splendens (Cuvier 1829). Leiognathidae: Gazzinae. Habitat: marine.

argenteus, Leiognathus Lacepède [B. G. E.] 1802:448, 449 [Histoire naturelle des poissons (Lacepède) v. 4; ref. 4929] Tranquebar [Tharangambadi], India. Holotype (unique): ZMB 8756 (dry). Type catalog: Bauchot & Desoutter 1989:34 [ref. 15502]. The name equula is Latin for a young horse, the name does not decline (Kottelat 2013:340). Unneeded new name for Scomber edentulus Bloch 1795. •Synonym of Leiognathus equulus (Forsskål 1775) -- (Fricke 1999:260 [ref. 24106]). •Synonym of Leiognathus equula (Forsskål 1775) -- (Kottelat 2013:339 [ref. 32989] with question, Chakrabarty & Sparks 2022:425 [ref. 39869]). Current status: Synonym of Leiognathus equula (Forsskål 1775). Leiognathidae: Leiognathinae. Habitat: freshwater, brackish, marine.

asina, Equula De Vis [C. W.] 1884:544 [Proceedings of the Linnean Society of New South Wales v. 9 (pt 3); ref. 4899] Cape York, Queensland, Australia. Lectotype: QM I.1700. Paralectotypes: ? [non-types] AMS I.355-56 (1, 1), QM I.9809 (?1 of 2). Lectotype selected by Whitley 1932:101-102 [ref. 4675]; later selected by Jones 1985:589 [ref. 21842]; originally 2 syntypes. •Synonym of Leiognathus fasciatus (Lacepède 1803) -- (Jones 1985:587 [ref. 21842], Hoese & Bray 2006:1180 [ref. 29000]). •Synonym of Aurigequula fasciata (Lacepède 1803) -- (Chakrabarty & Sparks 2022:416 [ref. 39869]). Current status: Synonym of Aurigequula fasciata (Lacepède 1803). Leiognathidae: Leiognathinae. Habitat: brackish, marine.

berbis, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:85 [Histoire naturelle des poissons v. 10; ref. 1004] Gulf of Suez, Red Sea, Egypt [Jeddah, Saudi Arabia, Red Sea; Al-Luhayya, Yemen, Red Sea]. Neotype: HUJ 6204. Type catalog: Bauchot & Desoutter 1989:33 [ref. 15502]. Based on Scomber equula var minimus Forsskål 1775. Nomen protectum according to Fricke 2008:34 [ref. 30182]. Neotype designated by Suzuki & Kimura 2023:24 [ref. 40749]. •See Randall 1995:193 [ref. 22896]. •Nomen dubium -- (Chakrabarty et al. 2010:20 [ref. 30812]). •Valid as Leiognathus berbis (Valenciennes 1835) -- (Dor 1984:134 [ref. 29757], Krishnan & Mishra 1993:225 [ref. 24016], Goren & Dor 1994:35 [ref. 25356], Woodland in Randall & Lim 2000:617 [ref. 25122], Woodland et al. 2001:2801 [ref. 26095], Mishra & Krishnan 2003:32 [ref. 27712], Fricke 2008:34 [ref. 30182], Alshawy et al. 2016:1 [ref. 35105], Fricke et al. 2018:191 [ref. 35805], Golani & Fricke 2018:88 [ref. 36273]). •Valid as Equulites berbis (Valenciennes 1835) -- (Kailola 1971:126 [ref. 39475], Suzuki & Kimura 2023:18 [ref. 40749]). Nomen Protectum. Current status: Valid as Equulites berbis (Valenciennes 1835). Leiognathidae: Leiognathinae. Distribution: Red Sea; Indo-West Pacific: Gulf of Oman, Persian Gulf, Saudi Arabia, east Africa and Madagascar, Iran, Pakistan, India, east to Indonesia, Vietnam, Philippines, China, Taiwan, north to Okinawa Island, Japan; Mediterranean Sea and Sea of Marmara (Red Sea immigrant). Habitat: brackish, marine.

bindoides, Equula Bleeker [P.] 1851:372 [Natuurkundig Tijdschrift voor Nederlandsch Indië v. 1 (no. 4); ref. 326] Batavia, Java, Indonesia. Syntypes: RMNH 8218 (9), RMNH 8219 (probably some out of 31), SMNS 10611 (1). Type catalog: Fricke 1991:16 [ref. 13474], Fricke 2005:46 [ref. 29864] Also in Bleeker 1852:83 [ref. 332]. •Valid as Leiognathus bindoides (Bleeker 1851) -- (authors). Current status: Valid as Leiognathus bindoides (Bleeker 1851). Leiognathidae: Leiognathinae. Distribution: Indonesia. Habitat: marine.

bindus, Equula Valenciennes [A.] (ex Russell) in Cuvier & Valenciennes 1835:78 [Histoire naturelle des poissons v. 10; ref. 1004] Locality not stated [Vizagapatam, India]. No types known. Type catalog: Bauchot & Desoutter 1989:33 [ref. 15502]. Based on the 'Bindoo-karah' of Russell 1803:50, Pl. 64 (vol. 1) [ref. 13862]. •Valid as Leiognathus bindus (Valenciennes 1835) -- (Kyushin et al. 1982:131 [ref. 19754], Dor 1984:135 [ref. 29757], Uyeno & Yabumoto in Masuda et al. 1984:158 [ref. 6441], Jones 1985:573 [ref. 21842], Allen & Swainston 1988:76 [ref. 25477], Talwar & Jhingran 1991:829 [ref. 20764], Krishnan & Mishra 1993:225 [ref. 24016], Goren & Dor 1994:35 [ref. 25356], Randall 1995:191 [ref. 22896], Mohsin & Ambak 1996:344 [ref. 27969], Chen et al. 1997:76 [ref. 26476], Cheng & Zhou 1997:296 [ref. 26385], Larson & Williams 1997:359 [ref. 23967], Allen 1997:118 [ref. 23977], Carpenter et al. 1997:169 [ref. 23922], Mishra & Srinivasan 1999:246 [ref. 24754], Mishra et al. 1999:86 [ref. 25315], Nakabo 2000:812 [ref. 25086], Woodland in Randall & Lim 2000:617 [ref. 25122], Bijukumar & Sushama 2000:186 [ref. 25703], Iwatsuki et al. 2000:100 [ref. 26368], Kimura & Peristiwady in Matsuura & Peristiwady 2000:219 [ref. 27511], Hutchins 2001:33 [ref. 25847], Woodland et al. 2001:2802 [ref. 26095], Nakabo 2002:812 [ref. 26001], Manilo & Bogorodsky 2003:S108 [ref. 27377], Ataur Rahman 2003:366 [ref. 31338], Adrim et al. 2004:120 [ref. 27715], Sparks & Dunlap 2004:7 [ref. 27928], Ra et al. 2005:95 [ref. 28304], Hoese & Bray 2006:1178 [ref. 29000], Chakrabarty & Sparks 2007:22 [ref. 29266]). •Valid as Photopectoralis bindus (Valenciennes 1835) -- (Sparks et al. 2005:316 [ref. 28434], Sparks 2006:14 [ref. 28783], Yamanoue et al. 2008:200 [ref. 29613], Chakrabarty et al. 2008:155 [ref. 32414], Chakrabarty et al. 2009:155 [ref. 30295], Kimura 2009:133 [ref. 30426], Chakrabarty et al. 2010:117 [ref. 31090], Abraham et al. 2011:10 [ref. 31311], Allen & Erdmann 2012:442 [ref. 31980], Larson et al. 2013:128 [ref. 32988], Yoshida et al. 2013:128 [ref. 34464], Bogorodsky et al. 2014:423 [ref. 33460], Psomadakis et al. 2015:237 [ref. 34104], Motomura et al. 2017:125 [ref. 35490], Ali et al. 2018:325 [ref. 36183], Golani & Fricke 2018:88 [ref. 36273], Kimura in Kimura et al. 2018:148 [ref. 36460], Suresh et al. 2018:224 [ref. 36495], Eagderi et al. 2019:77 [ref. 37020], Habib & Islam 2020:Supplementary table p. 22 [ref. 38321], Psomadakis et al. 2020:442 [ref. 37272], Sharifuzzaman et al. 2021:[3] [ref. 38872], Chakrabarty & Sparks 2022:428 [ref. 39869], Vilasri et al. 2023:86 [ref. 40764]). Current status: Valid as Photopectoralis bindus (Valenciennes 1835). Leiognathidae: Gazzinae. Distribution: Red Sea; Indo-West Pacific: Gulf of Aden and Persian Gulf east to Philippines and Fiji, north to Taiwan, south to Western Australia and New Caledonia. Habitat: brackish, marine.

blochii, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:84 [Histoire naturelle des poissons v. 10; ref. 1004] Kerala State, India. Lectotype: MNHN A-6757. Paralectotypes: MNHN A-6757 (1, poor condition) Malabar, A-6759 (2) Mahe; ZMB 8757 (1, right skin) Tranquebar. Type catalog: Bauchot & Desoutter 1989:19 [ref. 15502], Paepke 1999:93 [ref. 24282]. Lectotype selected by Kimura et al. 2008:25 [ref. 29525]. •Valid as Leiognathus blochii (Valenciennes 1835) -- (Jones 1985:576 [ref. 21842], Talwar & Jhingran 1991:830 [ref. 20764], Krishnan & Mishra 1993:226 [ref. 24016], Kottelat et al. 1993:111 [ref. 23448], Woodland in Randall & Lim 2000:617 [ref. 25122], Bijukumar & Sushama 2000:186 [ref. 25703], Hutchins 2001:33 [ref. 25847], Woodland et al. 2001:2803 [ref. 26095], Manilo & Bogorodsky 2003:S108 [ref. 27377], Khan 2003:9 [ref. 27710], Hoese & Bray 2006:1178 [ref. 29000]). •Valid as Nuchequula blochii (Valenciennes 1835) -- (Chakrabarty & Sparks 2007:13 [ref. 29266], Kimura et al. 2008:25 [ref. 29525], Kimura 2009:133 [ref. 30426], Kottelat 2013:340 [ref. 32989], Yoshida et al. 2013:126 [ref. 34464], Psomadakis et al. 2015:236 [ref. 34104], Suresh et al. 2018:223 [ref. 36495], Psomadakis et al. 2020:440 [ref. 37272], Sharifuzzaman et al. 2021:[3] [ref. 38872], Chakrabarty & Sparks 2022:426 [ref. 39869]). Current status: Valid as Nuchequula blochii (Valenciennes 1835). Leiognathidae: Gazzinae. Distribution: Indo-West Pacific: Pakistan east to Gulf of Thailand. Habitat: brackish, marine.

brevirostris, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:83 [Histoire naturelle des poissons v. 10; ref. 1004] Malabar, India. Syntypes: MNHN A-6763 (2). Type catalog: Bauchot & Desoutter 1989:19 [ref. 15502]. •Valid as Leiognathus brevirostris (Valenciennes 1835) -- (Zheng 1989:322 [ref. 21202], Ye in Pan et al. 1991:391 [ref. 23876], Krishnan & Mishra 1993:226 [ref. 24016], Mohsin & Ambak 1996:345 [ref. 27969], Mishra & Srinivasan 1999:246 [ref. 24754], Sadovy & Cornish 2000:110 [ref. 25480], Bijukumar & Sushama 2000:186 [ref. 25703], Mishra & Krishnan 2003:33 [ref. 27712], Habib & Islam 2020:Supplementary table p. 22 [ref. 38321]). •See Randall 1995:192 [ref. 22896]. •Synonym of Leiognathus bindus (Valenciennes 1835) -- (Jones 1985:573 [ref. 21842], Talwar & Jhingran 1991:829 [ref. 20764], Chakrabarty & Sparks 2007:22 [ref. 29266]). •Synonym of Photopectoralis bindus (Valenciennes 1835). Current status: Synonym of Photopectoralis bindus (Valenciennes 1835). Leiognathidae: Leiognathinae. Habitat: brackish, marine.

caballa, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:73 [Histoire naturelle des poissons v. 10; ref. 1004] Indian Ocean; Red Sea. Syntypes: MNHN A-6618 (1) Guam, MNHN A-6722 (1) Red Sea; MNHN 0000-3091 (1) Malabar; one syntype from Red Sea and one from Mumbai lost. Type catalog: Bauchot & Desoutter 1989:19 [ref. 15502]. •Synonym of Leiognathus equulus (Forsskål 1775) -- (Dor 1984:135 [ref. 29757]). •Synonym of Leiognathus equula (Forsskål 1775) -- (Kottelat 2013:340 [ref. 32989], Chakrabarty & Sparks 2022:425 [ref. 39869]). Current status: Synonym of Leiognathus equula (Forsskål 1775). Leiognathidae: Leiognathinae. Habitat: freshwater, brackish, marine.

cara, Equula Cuvier [G.] 1829:212 [Le Règne Animal (Edition 2) v. 2; ref. 995] Indian seas [Vizagapatam, India]. No types known. Type catalog: Bauchot & Desoutter 1989:33 [ref. 15502]. Available from footnote as, "Eq. cara, N., Russel, 66." Also appeared in Valenciennes in Cuvier & Valenciennes 1835:95 [ref. 1004]. •Synonym of Leiognathus fasciatus (Lacepède 1803). •Synonym of Aurigequula fasciata (Lacepède 1803) -- (Chakrabarty & Sparks 2022:416 [ref. 39869]). Current status: Synonym of Aurigequula fasciata (Lacepède 1803). Leiognathidae: Gazzinae. Habitat: brackish, marine.

coma, Equula Cuvier [G.] 1829:212 [Le Règne Animal (Edition 2) v. 2; ref. 995] Indian seas [Visakhapatnam]. No types known. Type catalog: Bauchot & Desoutter 1989:33 [ref. 15502]. Available from footnote as, "Eq. coma, Russ. et Seb., III, xxvii, 4, 63." Also in Cuvier & Valenciennes 1835:76 [ref. 1004]. •Synonym of Leiognathus equula (Forsskål 1775) -- (Kottelat 2013:339 [ref. 32989]). Current status: Synonym of Leiognathus equula (Forsskål 1775). Leiognathidae: Leiognathinae. Habitat: freshwater, brackish, marine.

daura, Equula Cuvier [G.] 1829:212 [Le Règne Animal (Edition 2) v. 2; ref. 995] Indian seas [Vizagapatam, India]. No types known. Type catalog: Bauchot & Desoutter 1989:34 [ref. 15502]. Available from footnote as, "Eq. Daura, N., Russel, 65." •Valid as Leiognathus daura (Cuvier 1829) -- (Randall 1995:192 [ref. 22896], Mohsin & Ambak 1996:346 [ref. 27969], Mishra et al. 1999:86 [ref. 25315], Woodland in Randall & Lim 2000:617 [ref. 25122], Kimura & Peristiwady in Matsuura & Peristiwady 2000:220 [ref. 27511], Woodland et al. 2001:2804 [ref. 26095], Allen & Adrim 2003:38 [ref. 26830], Manilo & Bogorodsky 2003:S108 [ref. 27377], Kimura 2009:131 [ref. 30426]). •Valid as Karalla daura (Cuvier 1829) -- (Chakrabarty & Sparks 2008:7 [ref. 29788], Chakrabarty et al. 2009:153 [ref. 30295], Chakrabarty et al. 2010:120 [ref. 31090], Abraham et al. 2011:9 [ref. 31311], Allen & Erdmann 2012:442 [ref. 31980], Chakrabarty et al. 2008:153 [ref. 32414], Psomadakis et al. 2015:235 [ref. 34104], Eagderi et al. 2019:76 [ref. 37020], Psomadakis et al. 2020:439 [ref. 37272], Sharifuzzaman et al. 2021:[3] [ref. 38872], Chakrabarty & Sparks 2022:424 [ref. 39869]). Current status: Valid as Karalla daura (Cuvier 1829). Leiognathidae: Gazzinae. Distribution: Indo-West Pacific: East Africa (Tanzania), Gulf of Aden, Gulf of Oman and Persian Gulf east to Philippines and eastern Indonesia. Habitat: marine.

decora, Equula De Vis [C. W.] 1884:543 [Proceedings of the Linnean Society of New South Wales v. 9 (pt 3); ref. 4899] Cape York, Queensland, Australia [Coral Sea, western Pacific]. Lectotype: QM I.1698. Paralectotypes: QM I.4877 (2). Lectotype selected Whitley 1932:105 [ref. 4675]. •Valid as Leiognathus decorus (De Vis 1884) -- (Jones 1985:578 [ref. 21842], Talwar & Jhingran 1991:830 [ref. 20764], Kottelat et al. 1993:111 [ref. 23448], Randall 1995:192 [ref. 22896], Larson & Williams 1997:359 [ref. 23967], Carpenter et al. 1997:169 [ref. 23922], Johnson 1999:735 [ref. 25471], Woodland in Randall & Lim 2000:617 [ref. 25122], Hutchins 2001:33 [ref. 25847], Woodland et al. 2001:2805 [ref. 26095], Manilo & Bogorodsky 2003:S108 [ref. 27377], Ikejima et al. 2004:908 [ref. 27755], Hoese & Bray 2006:1179 [ref. 29000]). •Valid as Nuchequula decora (De Vis 1884) -- (Chakrabarty & Sparks 2007:16 [ref. 29266]). •Synonym of Nuchequula gerreoides (Bleeker 1851) -- (Kimura et al. 2008:31 [ref. 29525], Kottelat 2013:340 [ref. 32989], Fricke et al. 2019:150 [ref. 36673], Chakrabarty & Sparks 2022:427 [ref. 39869]). Current status: Synonym of Nuchequula gerreoides (Bleeker 1851). Leiognathidae: Gazzinae. Habitat: marine.

devisi, Leiognathus Whitley [G. P.] 1929:113, Fig. 2 [Records of the Australian Museum v. 17 (no. 3); ref. 4665] Cape York, Queensland, Australia. Lectotype: QM I.1699. Paralectotypes: QM (1) missing. Replacement name for Equula argentea De Vis 1884, secondarily preoccupied in Leiognathus by Leiognathus argenteus Lacepède 1802 -- (Jones 1985:578 [ref. 21842]). Description by De Vis was based on two syntypes, only one remained and was wrongly designated as a holotype by Whitley 1929:115 [ref. 4665]. Whitley 1932:105 [ref. 4675] stated "Lectotype of Equula argentea De Vis = holotype of Leiognathus devisi Whitley, but devisi is a replacement name taking the same two types as Equula argentea De Vis. •Synonym of Leiognathus decorus (De Vis 1884) -- (Hoese & Bray 2006:1179 [ref. 29000]). •Synonym of Nuchequula gerreoides (Bleeker 1851). •Synonym of Eubleekeria splendens (Cuvier 1829) -- (Kottelat 2013:338 [ref. 32989], Chakrabarty & Sparks 2022:421 [ref. 39869]). Replacement Name. Current status: Synonym of Eubleekeria splendens (Cuvier 1829). Leiognathidae: Gazzinae. Habitat: marine.

dussumieri, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:77, Pl. 283 [Histoire naturelle des poissons v. 10; ref. 1004] Coromandel coast, southeastern India. Syntypes: MNHN A-6720 (1), A-6721 (1). Type catalog: Bauchot & Desoutter 1989:20 [ref. 15502]. •Valid as Leiognathus dussumieri (Valenciennes 1835) -- (Kottelat et al. 1993:111 [ref. 23448], Fricke 1999:260 [ref. 24106], Mishra & Srinivasan 1999:246 [ref. 24754], Woodland in Randall & Lim 2000:617 [ref. 25122], Woodland et al. 2001:2806 [ref. 26095], Manilo & Bogorodsky 2003:S108 [ref. 27377], Mishra & Krishnan 2003:33 [ref. 27712], Fricke et al. 2009:59 [ref. 30213]). •Valid as Karalla dussumieri (Valenciennes 1835) -- (Chakrabarty & Sparks 2008:7 [ref. 29788], Chakrabarty et al. 2008:153 [ref. 32414], Chakrabarty et al. 2009:153 [ref. 30295], Abraham et al. 2011:9 [ref. 31311], Kottelat 2013:338 [ref. 32989], Fricke et al. 2018:190 [ref. 35805], Psomadakis et al. 2020:440 [ref. 37272], Chakrabarty & Sparks 2022:425 [ref. 39869]). Current status: Valid as Karalla dussumieri (Valenciennes 1835). Leiognathidae: Gazzinae. Distribution: Indo-West Pacific: Madagascar, western Mascarenes (La Réunion), India and Sri Lanka east to Indonesia and Philippines. Habitat: brackish, marine.

edentulus, Scomber Bloch [M. E.] 1795:109, Pl. 428 [Naturgeschichte der ausländischen Fische v. 9; ref. 464] No locality given [East India, evidently Tranquebar [Tharangambadi], India]. Holotype (unique): ZMB 8756 (left skin). Type catalog: Paepke 1999:93 [ref. 24282]. On p. 109 of Bloch's Ichthyologie, v. 12 [ref. 21381]. Holotype figured in Sparks & Dunlap 2004:15 [ref. 27928] •Synonym of Leiognathus equulus (Forsskål 1775) -- (Dor 1984:135 [ref. 29757], Fricke 1999:260 [ref. 24106], Ataur Rahman 2003:365 [ref. 31338]). •Synonym of Leiognathus equula (Forsskål 1775) -- (Kottelat 2013:339 [ref. 32989] with question). Current status: Synonym of Leiognathus equula (Forsskål 1775). Leiognathidae: Leiognathinae. Habitat: freshwater, brackish, marine.

elongata, Equula Günther [A.] 1874:369 [2] [Annals and Magazine of Natural History (Series 4) v. 14 (no. 83) (art. 47); ref. 2006] North Sulawesi. Holotype (unique): BMNH 1872.4.6.105. •Valid as Leiognathus elongatus (Günther 1874) -- (Kyushin et al. 1982:127 [ref. 19754], Dunlap & McFall-Ngai 1984:884 [ref. 21845], Uyeno & Yabumoto in Masuda et al. 1984:158 [ref. 6441], Jones 1985:581 [ref. 21842], Smith 1986:620 [ref. 5712], Allen & Swainston 1988:76 [ref. 25477], Mohsin & Ambak 1996:346 [ref. 27969], Allen 1997:118 [ref. 23977], Kuiter 1997:164 [ref. 25488], Fricke 1999:261 [ref. 24106], Nakabo 2000:810 [ref. 25086], Woodland in Randall & Lim 2000:617 [ref. 25122], Hutchins 2001:33 [ref. 25847], Woodland et al. 2001:2807 [ref. 26095], Nakabo 2002:810 [ref. 26001], Choi et al. 2003:334, 609 [ref. 26218], Manilo & Bogorodsky 2003:S108 [ref. 27377], Mishra & Krishnan 2003:33 [ref. 27712], Ikejima et al. 2004:908 [ref. 27755], Sparks & Dunlap 2004:7 [ref. 27928], Heemstra & Heemstra 2004:276 [ref. 28072], Randall 2005:240 [ref. 28239], Ra et al. 2005:92 [ref. 28304], Hoese & Bray 2006:1179 [ref. 29000]). •Valid as Photoplagios elongatus (Günther 1874) -- (Sparks 2006:1 [ref. 28783], Sparks & Chakrabarty 2007:622 [ref. 29261]). •Valid as Equulites elongata (Günther 1874) -- (Kimura et al. 2008:204 [ref. 29614], Chakrabarty et al. 2010:18 [ref. 30812], Chakrabarty et al. 2010:120 [ref. 31090], Golani et al. 2011:S75 [ref. 31436]. Larson et al. 2013:126 [ref. 32988]). •Valid as Equulites elongatus (Günther 1874) -- (Bogorodsky et al. 2014:423 [ref. 33460], Suzuki & Kimura 2017:[7] [ref. 35231], Ali et al. 2018:324 [ref. 36183], Fricke et al. 2018:189 [ref. 35805], Eagderi et al. 2019:75 [ref. 37020], Fricke et al. 2019:149 [ref. 36673], Habib & Islam 2020:Supplementary table p. 22 [ref. 38321], Psomadakis et al. 2020:437 [ref. 37272], Kovačić et al. 2021:88 [ref. 39014], Balisco et al. 2023:5 [ref. 39932], Suzuki & Kimura 2023:17 [ref. 40749]). Current status: Valid as Equulites elongatus (Günther 1874). Leiognathidae: Gazzinae. Distribution: Indo-West Pacific: Myanmar, Indonesia (Ambon and north Sulawesi), and northern Australia. IUCN (2020): Not assessed. Habitat: marine.

elongatus, Leiognathus Smith [H. M.] & Pope [T. E. B.] 1906:466, Fig. 2 [Proceedings of the United States National Museum v. 31 (no. 1489); ref. 4058] Kagoshima, Japan. Holotype (unique): USNM 55613. Subjectively invalid; secondarily preoccupied in Leiognathus by Equula elongata Günther 1874, replaced by Macilentichthys popei Whitley 1932. •Synonym of Leiognathus elongatus (Günther 1874) -- (Dunlap & McFall-Ngai 1984:886 [ref. 21845], Jones 1985:581 [ref. 21842]). •Synonym of Equulites elongatus (Günther 1874) -- (Kimura 2009:128 [ref. 30426]). •Synonym of Equulites popei (Whitley 1932) -- (Suzuki & Kimura 2023:17 [ref. 40749]). 2°Homonym. Current status: Synonym of Equulites popei (Whitley 1932). Leiognathidae: Gazzinae. Habitat: marine.

ensifera, Equula Cuvier [G.] 1829:212 [Le Règne Animal (Edition 2) v. 2; ref. 995] Indian seas [Tharangambadi]. Holotype (unique): ZMB 8756 (dry). Type catalog: Bauchot & Desoutter 1989:34 [ref. 15502]. Available from footnote as, "Eq. ensifera, Nob., ou Scomber edentulus, Bl., 428, ou Leyognathe argenté, Lacép." Replacement name for Scomber edetulus Block 1795. Also described in Valenciennes in Cuvier & Valenciennes 1835:66 [ref. 1004]. •Synonym of Leiognathus equulus (Forsskål 1775) -- (Fricke 1999:260 [ref. 24106]). •Synonym of Leiognathus equula (Forsskål 1775) -- (Kottelat 2013:339 [ref. 32989] with question, Chakrabarty & Sparks 2022:425 [ref. 39869]). Current status: Synonym of Leiognathus equula (Forsskål 1775). Leiognathidae: Leiognathinae. Habitat: freshwater, brackish, marine.

equula, Scomber Forsskål [P. S.] in Niebuhr 1775:58, xii [Descriptiones animalium (Forsskål); ref. 1351] [Lohajæ] Al-Luhayya, Yemen, Red Sea. Lectotype: ZMUC P48219 (dry skin). Paralectotypes: ZMUC P48220 (1, dry skin). Type catalog: Nielsen 1974:64 [ref. 9588], Fricke 2008:34 [ref. 30182]. Lectotype selected by Klausewitz & Nielsen 1965:23 [ref. 20847]; lectotype figured in Klausewitz & Nielsen 1965:Pl. 28 [ref. 20847] and in Sparks & Dunlap 2004:15 [ref. 27928]. Authorship according to Fricke 2008:34 [ref. 30182]. Misspelled Caesio aequulus by Sonnini 1803:36 (vol. 8) [ref. 30473]. •Valid as Leiognathus equulus (Forsskål 1775) -- (Dor 1984:135 [ref. 29757], Jones 1985:584 [ref. 21842], Yu & Zhou in Kuang et al. 1986:233 [ref. 26639], Allen & Swainston 1988:76 [ref. 25477], Ataur Rahman 1989:335 [ref. 24860], Allen 1991:135 [ref. 21090], Talwar & Jhingran 1991:831 [ref. 20764], Kottelat et al. 1993:111 [ref. 23448], Goren & Dor 1994:35 [ref. 25356], Randall 1995:192 [ref. 22896], Rainboth 1996:184 [ref. 22772], Mohsin & Ambak 1996:347 [ref. 27969], Reinthal & Stiassny 1997:354 [ref. 22780], Allen 1997:118 [ref. 23977], Chen et al. 1997:76 [ref. 26476], Larson & Williams 1997:359 [ref. 23967], Carpenter et al. 1997:169 [ref. 23922], Kuiter 1997:164 [ref. 25488], Myers 1999:143 [ref. 23965], Fricke 1999:260 [ref. 24106], Mishra & Srinivasan 1999:246 [ref. 24754], Nakabo 2000:811 [ref. 25086], Woodland in Randall & Lim 2000:617 [ref. 25122], Laboute & Grandperrin 2000:344 [ref. 25191], Bijukumar & Sushama 2000:186 [ref. 25703], Iwatsuki et al. 2000:100 [ref. 26368], Kimura & Peristiwady in Matsuura & Peristiwady 2000:221 [ref. 27511], Sakai et al. 2001:97 [ref. 25693], Hutchins 2001:33 [ref. 25847], Woodland et al. 2001:2808 [ref. 26095], Allen et al. 2002:340 [ref. 25930], Nakabo 2002:811 [ref. 26001], Manilo & Bogorodsky 2003:S108 [ref. 27377], Myers & Donaldson 2003:626 [ref. 27495], Khan 2003:9 [ref. 27710] as equuals, Ataur Rahman 2003:365 [ref. 31338], Ikejima et al. 2004:908 [ref. 27755], Sparks & Dunlap 2004:7 [ref. 27928], Heemstra et al. 2004:3322 [ref. 27945], Heemstra & Heemstra 2004:275 [ref. 28072], Randall 2005:239 [ref. 28239], Hoese & Bray 2006:1179 [ref. 29000], Chakrabarty et al. 2008:156 [ref. 32414], Chakrabarty et al. 2009:155 [ref. 30295], Kimura 2009:131 [ref. 30426], Fricke et al. 2009:59 [ref. 30213], Chakrabarty et al. 2010:114 [ref. 31090], Abraham et al. 2011:6 [ref. 31311], Allen & Erdmann 2012:442 [ref. 31980], Larson et al. 2013:127 [ref. 32988], Yoshida et al. 2013:127 [ref. 34464], Fricke et al. 2014:84 [ref. 33932], Psomadakis et al. 2015:235 [ref. 34104], Motomura et al. 2017:124 [ref. 35490], Ali et al. 2018:325 [ref. 36183], Golani & Fricke 2018:88 [ref. 36273], Jawad et al. 2018:106 [ref. 36262], Kimura in Kimura et al. 2018:146 [ref. 36460], Suresh et al. 2018:222 [ref. 36495], Eagderi et al. 2019:76 [ref. 37020], Taki et al. 2021:389 [ref. 39830]). •Valid as Leiognathus equula (Forsskål 1775) -- (Uyeno & Yabumoto in Masuda et al. 1984:159 [ref. 6441], Smith 1986:621 [ref. 5712], Kottelat 2013:339 [ref. 32989], Fricke et al. 2018:191 [ref. 35805], Arndt & Fricke 2019:20 [ref. 36883], Zajonz et al. 2019:76 [ref. 36871], Habib & Islam 2020:Supplementary table p. 22 [ref. 38321], Psomadakis et al. 2020:440 [ref. 37272], Giat in Motomura et al. 2021:31 [ref. 38121], Sharifuzzaman et al. 2021:[3] [ref. 38872], Chakrabarty & Sparks 2022:425 [ref. 39869], Zajonz et al. 2022:132 [ref. 40522], Jamandre 2023:158 [ref. 40455], Vilasri et al. 2023:86 [ref. 40764]). Current status: Valid as Leiognathus equula (Forsskål 1775). Leiognathidae: Leiognathinae. Distribution: Red Sea; Indo-West Pacific: Eastern Cape and KwaZulu-Natal (South Africa), East Africa, Persian Gulf, Socotra (Yemen), Seychelles, Comoros, Madagascar and Mascarenes (La Réunion, Mauritius, Rodrigues) east to eastern Caroline Islands (Federated States of Micronesia), north to Ryukyu Islands (Japan), south to Western Australia, Queensland (Australia), New Caledonia and Fiji. Habitat: freshwater, brackish, marine.

filigera, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:92, Pl. 284 [Histoire naturelle des poissons v. 10; ref. 1004] Molucca Islands; Trincomalee, Sri Lanka; Malabar; Seychelles. Syntypes: MNHN 0000-3092 (1) Malabar, MNHN A-6750 (3) Seychelles, MNHN A-6751 (4) Seychelles, MNHN A-6752 (3) Trinquemali, A-6755 (1) Seychelles; Moluccas specimen lost. Type catalog: Bauchot & Desoutter 1989:20 [ref. 15502]. •Synonym of Leiognathus fasciatus (Lacepède 1803) -- (Dor 1984:135 [ref. 29757], Fricke 1999:261 [ref. 24106]). •Synonym of Aurigequula fasciata (Lacepède 1803) -- (Chakrabarty & Sparks 2022:416 [ref. 39869]). Current status: Synonym of Aurigequula fasciata (Lacepède 1803). Leiognathidae: Leiognathinae. Habitat: brackish, marine.

gerreoides, Equula Bleeker [P.] 1851:371 [Natuurkundig Tijdschrift voor Nederlandsch Indië v. 1 (no. 4); ref. 326] Jakarta, Indonesia. Neotype: RMNH 35540 [ex. RMNH 6140]. Possible types: BMNH 1880.4.21.141-142 (2) but not labeled types. Also in Bleeker 1852:81 [ref. 332]. Neotype designated by Kimura et al. 2008:31 [ref. 29525]. •Synonym of Leiognathus daura (Cuvier 1829) -- (Woodland et al. 2001:2804 [ref. 26095]). •Valid as Nuchequula gerreoides (Bleeker 1851) -- (Kimura et al. 2008:31 [ref. 29525], Kimura 2009:133 [ref. 30426], Larson et al. 2013:128 [ref. 32988], Kottelat 2013:340 [ref. 32989], Kimura in Kimura et al. 2015:48 [ref. 34319], Psomadakis et al. 2015:236 [ref. 34104], Kimura in Kimura et al. 2018:146 [ref. 36460], Eagderi et al. 2019:76 [ref. 37020], Fricke et al. 2019:150 [ref. 36673], Habib & Islam 2020:Supplementary table p. 22 [ref. 38321], Psomadakis et al. 2020:441 [ref. 37272], Sharifuzzaman et al. 2021:[3] [ref. 38872], Taki et al. 2021:389 [ref. 39830], Chakrabarty & Sparks 2022:427 [ref. 39869]). Current status: Valid as Nuchequula gerreoides (Bleeker 1851). Leiognathidae: Gazzinae. Distribution: Indo-West Pacific: Persian Gulf east to Philippines and New Ireland (Papua New Guinea), north to Taiwan, south to northern Australia. Habitat: marine.

gomorah, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:80 [Histoire naturelle des poissons v. 10; ref. 1004] Puducherry, India. Lectotype: MNHN A-6724. Paralectotypes: MNHN A-6725 (1) Malabar, A-6726 (1) Malabar, 0000-0581 (1) Red Sea. Type catalog: Bauchot & Desoutter 1989:20 [ref. 15502]. Lectotype selected by Kimura et al. 2005:285 [ref. 28424]. •Synonym of Leiognathus splendens (Cuvier 1829) -- (Dor 1984:136 [ref. 29757], Kimura et al. 2005:285 [ref. 28424]). •Synonym of Eubleekeria splendens (Cuvier 1829) -- (Kottelat 2013:338 [ref. 32989], Chakrabarty & Sparks 2022:421 [ref. 39869]). Current status: Synonym of Eubleekeria splendens (Cuvier 1829). Leiognathidae: Gazzinae. Habitat: brackish, marine.

hastatus, Liognathus Ogilby [J. D.] 1912:58 [Memoirs of the Queensland Museum v. 1; ref. 14855] Cape York, Queensland, Australia. Holotype (unique): QM I.878. Replacement name for Equula longispina De Vis 1884, preoccupied by Equula longispinis Valenciennes 1835. •Synonym of Leiognathus leuciscus (Günther 1860) -- (Hoese & Bray 2006:1180 [ref. 29000]). •Synonym of Leiognathus moretoniensis Ogilby 1912 -- (Woodland et al. 2001:2812 [ref. 26095]). •Synonym of Equulites leuciscus (Günther 1860) -- (Suzuki & Kimura 2023:30 [ref. 40749]). Replacement Name. Current status: Synonym of Equulites leuciscus (Günther 1860). Leiognathidae: Gazzinae. Habitat: marine.

klunzingeri, Equula Steindachner [F.] 1898:199 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe v. 35 (no. 19) (for 14July 1898); ref. 20495] Al Ghulayfiqah, Yemen [Gulf of Suez, Egypt, Red Sea]. Lectotype: NMW 76009-6. Paralectotypes: (several) NMW 22366 (1), 59908-09 (21, 8), 59904 (1), 68276-81 (8, 26, 9, 4, 2, 13), 76008-09 (1, 7). Later illustrated and described in more detail in Steindachner 1898:782, pl. 1 (figs. 2-2a) [ref. 14693]. Lectotype selected by Suzuki & Kimura 2023:20 [ref. 40749]. •Valid as Leiognathus klunzingeri (Steindachner 1898) -- (Tortonese 1973:390 [ref. 7192], Dor 1984:135 [ref. 29757], Roux in Whitehead et al. 1986:856 [ref. 13676], Goren & Dor 1994:35 [ref. 25356], Golani 1998:97 [ref. 27567], Bilecenoğlu et al. 2002:88 [ref. 26753], Dulĉić & Pallaoro 2002:523 [ref. 26144], Heemstra & Heemstra 2004:274 [ref. 28072], Golani 2005:41 [ref. 37112], Fricke et al. 2007:85 [ref. 29533], Lipej & Dulčić 2010:46 [ref. 36649]). •Valid, tentatively as Photoplagios klunzingeri (Steindachner 1898) -- (Sparks et al. 2005:315 [ref. 28434]). •Valid as Photoplagios klunzingeri (Steindachner 1898) -- (Sparks 2006:1 [ref. 28783], Sparks & Chakrabarty 2007:622 [ref. 29261]). •Valid as Equulites klunzingeri (Steindachner 1898) -- (Kimura et al. 2008:204 [ref. 29614], Chakrabarty et al. 2010:18 [ref. 30812], Chakrabarty & Sparks 2008:6 [ref. 29788], Golani et al. 2011:S75 [ref. 31436], Bogorodsky et al. 2014:423 [ref. 33460], Suzuki & Kimura 2017:[11] [ref. 35231], Bogorodsky & Randall 2018:[21] 259 [ref. 36492], Golani & Fricke 2018:88 [ref. 36273], Artüz & Fricke 2019:556 [ref. 36612], Khajavi et al. 2019:537 [ref. 37194], Bariche & Fricke 2020:130 [ref. 37515], Kovačić et al. 2020:33 [ref. 37519], Golani 2021:51 [ref. 38303], Kovačić et al. 2021:60 [ref. 39014], Chakrabarty & Sparks 2022:419 [ref. 39869]). •Synonym of Equulites berbis (Valenciennes 1835) -- (Suzuki & Kimura 2023:19 [ref. 40749]). Current status: Synonym of Equulites berbis (Valenciennes 1835). Leiognathidae: Gazzinae. Habitat: marine.

leuciscus, Equula Günther [A.] 1860:503 [Catalogue of the fishes in the British Museum v. 2; ref. 1963] Ambon Island, Molucca Islands, Indonesia. Holotype (unique): BMNH 1858.4.21.243. •Valid as Leiognathus leuciscus (Günther 1860) -- (Uyeno & Yabumoto in Masuda et al. 1984:158 [ref. 6441], Jones 1985:590 [ref. 21842], Allen & Swainston 1988:76 [ref. 25477], Krishnan & Mishra 1994:289 [ref. 24014], Mohsin & Ambak 1996:348 [ref. 27969], Allen 1997:118 [ref. 23977], Nakabo 2000:812 [ref. 25086], Woodland in Randall & Lim 2000:617 [ref. 25122], Hutchins 2001:33 [ref. 25847], Woodland et al. 2001:2810 [ref. 26095], Nakabo 2002:812 [ref. 26001], Manilo & Bogorodsky 2003:S108 [ref. 27377], Kimura et al. in Kimura & Matsuura 2003:83 [ref. 27510], Adrim et al. 2004:121 [ref. 27715], Sparks & Dunlap 2004:7 [ref. 27928], Hoese & Bray 2006:1180 [ref. 29000]). •Valid as Photoplagios leuciscus (Günther 1860) -- (Sparks et al. 2005:315 [ref. 28434], Sparks 2006:1 [ref. 28783], Sparks & Chakrabarty 2007:622 [ref. 29261], Yamanoue & Toda 2008:205 [ref. 29613]). •Valid as Equulites leuciscus (Günther 1860) -- (Kimura et al. 2008:204 [ref. 29614], Chakrabarty & Sparks 2008:6 [ref. 29788], Kimura 2009:129 [ref. 30426], Chakrabarty et al. 2010:15 [ref. 30812], Chakrabarty et al. 2010:111 [ref. 31090], Abraham et al. 2011:13 [ref. 31311], Allen & Erdmann 2012:441 [ref. 31980], Larson et al. 2013:126 [ref. 32988], Kimura in Kimura et al. 2015:46 [ref. 34319], Psomadakis et al. 2015:234 [ref. 34104], Suzuki & Kimura 2017:[11] [ref. 35231], Fricke et al. 2018:189 [ref. 35805], Sparks & Chakrabarty 2019:9 [ref. 36798], Psomadakis et al. 2020:438 [ref. 37272], Sharifuzzaman et al. 2021:[3] [ref. 38872], Chakrabarty & Sparks 2022:420 [ref. 39869], Suzuki & Kimura 2023:29 [ref. 40749]). Current status: Valid as Equulites leuciscus (Günther 1860). Leiognathidae: Gazzinae. Distribution: Indo-West Pacific: East Africa, Oman, Seychelles and Madagascar east to Philippines and Vanuatu, north to Taiwan and China, south to northern Australia and New Caledonia. Habitat: marine.

lineolata, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:86 [Histoire naturelle des poissons v. 10; ref. 1004] Java, Indonesia. "Syntypes": MNHN A-6738 (largest, 46 mm), MNHN A-6738 (smallest), A-6739 (1, poor condition), 1988-0327 [ex MNHN A-6739] (1, poor condition), 1988-0328 [ex MNHN A-6739] (2); Antjer specimens not found. Type catalog: Bauchot & Desoutter 1989:20 [ref. 15502]. Jones 1985:575 [ref. 21842] designated two specimens as lectotypes; Bauchot & Desoutter 1989:22 restricted the designation to one specimen so they designated the lectotype, but see Sparks 2006:15 [ref. 28783]. See also Suzuki & Kimura 2023:44 [ref. 40749]. •Synonym of Leiognathus bindus (Valenciennes 1835) -- (Jones 1985:573 [ref. 21842]). •Valid as Leiognathus lineolatus (Valenciennes 1835) -- (Uyeno & Yabumoto in Masuda et al. 1984:158 [ref. 6441], Ye in Pan et al. 1991:393 [ref. 23876], Krishnan & Mishra 1994:290 [ref. 24014], Goren & Dor 1994:36 [ref. 25356], Heemstra in Smith & Heemstra 1995:xii [ref. 21953], Chen et al. 1997:77 [ref. 26476], Iwatsuki et al. 2000:100 [ref. 26368], Choi et al. 2003:334, 609 [ref. 26218], Ra et al. 2005:93 [ref. 28304]). •See Randall 1995:193 [ref. 22896]. •Valid as Photoplagios lineolatus (Valenciennes 1835) -- (Sparks et al. 2005:315 [ref. 28434], Sparks 2006:15 [ref. 28783] but status uncertain, Sparks 2006:1, 12 [ref. 28783]). •In genus Equulites Fowler 1904 -- (Kimura et al. 2008:204-205 [ref. 29614]). •Valid as Equulites lineolatus (Valenciennes 1835) -- (Ali et al. 2018:324 [ref. 36183], Fricke et al. 2018:190 [ref. 35805], Eagderi et al. 2019:76 [ref. 37020], Fricke et al. 2019:149 [ref. 36673], Habib & Islam 2020:Supplementary table p. 22 [ref. 38321]). •Nomen Dubium -- (Suzuki & Kimura 2023:44 [ref. 40749]). Nomen Dubium. Current status: Synonym of Equulites. Leiognathidae: Gazzinae. Habitat: marine.

longispina, Equula De Vis [C. W.] 1884:542 [Proceedings of the Linnean Society of New South Wales v. 9 (pt 3); ref. 4899] Cape York, Queensland, Australia. Holotype (unique): QM I.878 [not AMS I.465]. Apparently described independently from (and preoccupied by) Equula longispinis Valenciennes 1835, replaced by Leiognathus hastatus Ogilby 1912. Objectively invalid. The holotype is QM I.878 and not AMS I.465 as given by Whitley 1932:107 [ref. 4675]. •Synonym of Leiognathus leuciscus (Günther 1860) -- (Jones 1985:590 [ref. 21842]). •Synonym of Equulites leuciscus (Günther 1860) -- (Suzuki & Kimura 2023:29 [ref. 40749]). 1°Homonym. Current status: Synonym of Equulites leuciscus (Günther 1860). Leiognathidae: Gazzinae. Habitat: marine.

longispinis, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:94 [Histoire naturelle des poissons v. 10; ref. 1004] Waigeo, Papua Barat, Indonesia. Lectotype: MNHN A-0579. Type catalog: Bauchot & Desoutter 1989:21 [ref. 15502]. Lectotype designated by Kottelat (2013:239 [ref. 32989]). The species epithet is trated as a noun in apposition. •Valid as Leiognathus longispinis (Valenciennes 1835) -- (Woodland in Randall & Lim 2000:617 [ref. 25122], Woodland et al. 2001:2811 [ref. 26095], Manilo & Bogorodsky 2003:S108 [ref. 27377], Sparks 2006:539 [ref. 28801], Fricke et al. 2009:59 [ref. 30213]). •Valid as Equula longispinis (Valenciennes 1835) -- (Chakrabarty & Sparks 2008:5 [ref. 29788]). •Valid as Leiognathus longipinnis (sic) (Valenciennes 1835) -- (Kimura 2009:132 [ref. 30426]). •Valid as Aurigequula longispina (Valenciennes 1835) -- (Chakrabarty et al. 2009:149 [ref. 30295], Abraham et al. 2011:14 [ref. 31311], Psomadakis et al. 2015:233 [ref. 34104], Fricke et al. 2019:149 [ref. 36673], Psomadakis et al. 2020:436 [ref. 37272], Sharifuzzaman et al. 2021:[3] [ref. 38872], Chakrabarty & Sparks 2022:416 [ref. 39869]). •Valid as Aurigequula longispinis (Valenciennes 1835) -- (Chakrabarty et al. 2008:149 [ref. 32414], Larson et al. 2013:123 [ref. 32988] as longispinus, Vilasri et al. 2023:84 [ref. 40764]). Current status: Valid as Aurigequula longispinis (Valenciennes 1835). Leiognathidae: Leiognathinae. Distribution: Indo-West Pacific: South Africa, Seychelles, Madagascar, western Mascarenes (La Réunion) and India east to Palawan (Philippines) and New Ireland (Papua New Guinea), north to Gulf of Thailand, south to Onslow (Western Australia). Habitat: marine.

minimus, Scomber equula Forsskål [P. S.] in Niebuhr 1775:58, xii [Descriptiones animalium (Forsskål); ref. 1351] Jeddah, Saudi Arabia, Red Sea; Al-Luhayya, Yemen, Red Sea. Named on p. xii, tied to description on p. 58 through common name. Authorship according to Fricke 2008:34 [ref. 30182]. Nomen oblitum according to Fricke 2008:34 [ref. 30182]. •In the synonymy of Leiognathus berbis (Valenciennes 1835) -- (Dor 1984:134 [ref. 29757], Golani & Fricke 2018:88 [ref. 36273]). Nomen Oblitum. Current status: Synonym of Leiognathus berbis (Valenciennes 1835). Leiognathidae: Leiognathinae. Habitat: brackish, marine.

novaehollandiae, Equula Steindachner [F.] 1879:31 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe v. 16 (no. 4); ref. 20480] Cleveland Bay, Townsville, Queensland, Australia, 19°16'S, 146°48'E. Holotype (unique): SMNS 2285 (discarded in 1958). Type catalog: Fricke 1995:16 [ref. 28100], Fricke 2005:46 [ref. 29864]. Originally as novae Hollandiae, correction mandatory; later described in more detail in Steindachner 1879:11 [ref. 4228]. •Synonym of Leiognathus leuciscus (Günther 1860) -- (Jones 1985:590 [ref. 21842], Woodland et al. 2001:2810 [ref. 26095]). •Synonym of Equulites leuciscus (Günther 1860) -- (Chakrabarty & Sparks 2022:420 [ref. 39869]). •Nomen Dubium -- (Suzuki & Kimura 2023:32 [ref. 40749]). Nomen Dubium. Current status: Synonym of Equulites. Leiognathidae: Gazzinae. Habitat: marine.

novemaculeata, Equula splendens var. Klunzinger [C. B.] 1880:379 [55] [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 80 (1. Abth.) (nos 3-4); ref. 2624] Endeavour River, Queensland, Australia. Holotype (unique): SMNS 2448. Type catalog: Fricke 1992:12 [ref. 20296] and Fricke 2005:47 [ref. 29864] as novemaculata. On p. 55 of separate. •Synonym of Leiognathus splendens (Cuvier 1829) -- (Hoese & Bray 2006:1181 [ref. 29000]). •Synonym of Eubleekeria splendens (Cuvier 1829) -- (Kottelat 2013:338 [ref. 32989], Chakrabarty & Sparks 2022:421 [ref. 39869]). Current status: Synonym of Eubleekeria splendens (Cuvier 1829). Leiognathidae: Gazzinae. Habitat: brackish, marine.

nuchalis, Equula Temminck [C. J.] & Schlegel [H.] 1845:126, Pl. 67 (fig. 1) [Fauna Japonica Parts 7-9; ref. 4373] Nagasaki, Japan. Holotype: RMNH 1287. Type catalog: Boeseman 1947:110 [ref. 12876]. •Valid as Leiognathus nuchalis (Temminck & Schlegel 1845) -- (Uyeno & Yabumoto in Masuda et al. 1984:158 [ref. 6441], Kawanabe & Mizuno 1989:519 [ref. 25598], Kim 1997:419 [ref. 24032], Sato et al. 1998:40 [ref. 24972], Mizuno et al. 1999:9 [ref. 25311], Nakabo 2000:810 [ref. 25086], Woodland in Randall & Lim 2000:617 [ref. 25122], Sadovy & Cornish 2000:111 [ref. 25480], Kim & Park 2002:344 [ref. 25913], Woodland et al. 2001:2797 [ref. 26095], Nakabo 2002:810 [ref. 26001], Choi et al. 2003:334, 609 [ref. 26218], Ikejima et al. 2004:908 [ref. 27755], Sparks & Dunlap 2004:7 [ref. 27928], Ra et al. 2005:94 [ref. 28304]). •Valid as Nuchequula nuchalis (Temminck & Schlegel 1845) -- (Chakrabarty & Sparks 2007:18 [ref. 29266], Kimura et al. 2008:39 [ref. 29525], Chakrabarty et al. 2010:116 [ref. 31090], Abraham et al. 2011:6 [ref. 31311], Suzuki & Kimura in Kimura et al. 2018:147 [ref. 36460], Kim et al. 2020:129 [ref. 39118], Sonoyama et al. 2020:66 [ref. 37637]). Current status: Valid as Nuchequula nuchalis (Temminck & Schlegel 1845). Leiognathidae: Gazzinae. Distribution: Northwestern Pacific: northern Vietnam, China and Taiwan to Korea and Japan. Habitat: marine.

oblonga, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:85 [Histoire naturelle des poissons v. 10; ref. 1004] Timor Island, southern Malay Archipelago. Lectotype: MNHN A-6754. Paralectotype: A-6756 (1) Malabar. Type catalog: Bauchot & Desoutter 1989:21 [ref. 15502] with holotype (A-6754) and paratype. Sparks & Chakrabarty 2019:9 [ref. 36798] report MNHN A-6754 as holotype. Lectotype selected by Suzuki & Kimura 2023:36 [ref. 40749]. •Synonym of Leiognathus berbis (Valenciennes 1835) -- (Dor 1984:136 [ref. 29757]). •Valid as Leiognathus oblongus (Valenciennes 1835) -- (Randall 1995:193 [ref. 22896], Carpenter et al. 1997:170 [ref. 23922], Kimura & Peristiwady in Matsuura & Peristiwady 2000:223 [ref. 27511] with author as Lacepède 1803, Manilo & Bogorodsky 2003:S108 [ref. 27377], Chakrabarty et al. 2010:21 [ref. 30812]). •Valid as Equulites oblongus (Valenciennes 1835) -- (Kimura 2009:129 [ref. 30426], Yoshida et al. 2013:124 [ref. 34464], Kimura in Kimura et al. 2015:47 [ref. 34319], Psomadakis et al. 2015:234 [ref. 34104], Suzuki & Kimura 2017:[12] [ref. 35231], Motomura et al. 2017:123 [ref. 35490], Kimura in Kimura et al. 2018:145 [ref. 36460], Eagderi et al. 2019:75 [ref. 37020], Sparks & Chakrabarty 2019:9 [ref. 36798], Psomadakis et al. 2020:438 [ref. 37272], Giat in Motomura et al. 2021:31 [ref. 38121], Suzuki & Kimura 2023:36 [ref. 40749], Vilasri et al. 2023:84 [ref. 40764]). Current status: Valid as Equulites oblongus (Valenciennes 1835). Leiognathidae: Leiognathinae. Distribution: Indo-West Pacific: India, Sri Lanka, Malaysia, Thailand, Philippines, eastern Indonesia and Kagoshima, Japan. Habitat: marine.

obscura, Leiognathus Seale [A.] 1901:74 [Occasional Papers of the Bernice Pauahi Bishop Museum of Polynesian Ethnology and Natural History v. 1 (no. 3); ref. 14357] Agaña, Guam, Mariana Islands, western Pacific. Holotype: BPBM 84. Paratypes: ANSP 77756 (1), BPBM 85 (1). Type catalog: Böhlke 1984:119 [ref. 13621]. •Synonym of Leiognathus equula (Forsskål 1775) -- (Myers & Donaldson 2003:626 [ref. 27495] as equulus). Current status: Synonym of Leiognathus equula (Forsskål 1775). Leiognathidae: Leiognathinae. Habitat: freshwater, brackish, marine.

ovalis, Equula De Vis [C. W.] 1884:543 [Proceedings of the Linnean Society of New South Wales v. 9 (pt 3); ref. 4899] Cape York, Queensland, Australia. Lectotype: QM I.1703. Paralectotypes: AMS IA.4900 (1); QM I.7802 (missing), I.9810 (8). Lectotype selected by Whitley 1932:110 [ref. 4675]. •Synonym of Leiognathus splendens (Cuvier 1829) -- (Jones 1985:599 [ref. 21842], Kimura et al. 2005:285 [ref. 28424], Hoese & Bray 2006:1181 [ref. 29000]). •Synonym of Eubleekeria splendens (Cuvier 1829) -- (Kottelat 2013:338 [ref. 32989], Chakrabarty & Sparks 2022:421 [ref. 39869]). Current status: Synonym of Eubleekeria splendens (Cuvier 1829). Leiognathidae: Gazzinae. Habitat: brackish, marine.

parviceps, Equula Valenciennes [A.] in Cuvier & Valenciennes 1835:87 [Histoire naturelle des poissons v. 10; ref. 1004] Mauritius, Mascarenes, southwestern Indian Ocean. Syntypes: MNHN A-0580 (2), A-0581 (2). Type catalog: Bauchot & Desoutter 1989:21 [ref. 15502]. •Valid as Leiognathus parviceps (Valenciennes 1835) -- (Fricke 1999:261 [ref. 24106]). •Valid as ?Photoplagios parviceps (Valenciennes 1835) -- (Sparks & Chakrabarty 2007:628 [ref. 29261]). •Possibly in genus Equulites. Current status: Valid as Leiognathus parviceps (Valenciennes 1835). Leiognathidae: Leiognathinae. Distribution: Southwestern Indian Ocean: Mauritius (Mascarenes endemic); other records probably based on misidentifications. Habitat: marine.

popei, Macilentichthys Whitley [G. P.] 1932:114 [Memoirs of the Queensland Museum v. 10 (pt 2); ref. 4675] Kagoshima, Japan. Holotype (unique): USNM 55613. Replacement name for Leiognathus elongatus Smith & Pope 1906, preoccupied in Leiognathus by Equula elongata Günther 1874. •Synonym of Leiognathus elongatus (Günther 1874) -- (Dunlap & McFall-Ngai 1984:888 [ref. 21845], Jones 1985:581 [ref. 21842]). •Synonym of Photoplagios elongatus (Günther 1874) -- (Sparks 2006:1 [ref. 28783]). •Synonym of Equulites elongatus (Günther 1874). •Valid as Equulites popei (Whitley 1932) -- (Suzuki & Kimura 2017:[10] [ref. 35231], Bogorodsky & Randall 2018:[21] 259 [ref. 36492], Golani & Fricke 2018:88 [ref. 36273], Ergüden et al. 2019:372 [ref. 37185], Mavruk et al. 2019:261 [ref. 37240], Bariche & Fricke 2020:131 [ref. 37515], Ibrahim et al. 2020:[2] [ref. 37591], Golani 2021:51 [ref. 38303], Kovačić et al. 2021:60 [ref. 39014] as Equuilites popei, Suzuki & Kimura 2023:17 [ref. 40749]). Replacement Name. Current status: Valid as Equulites popei (Whitley 1932). Leiognathidae: Gazzinae. Distribution: Red Sea; Indo-West Pacific: southern Mozambique, Oman, east to Philippines, north to southern Japan (Okinawa Island); Mediterranean Sea (Red Sea immigrant). Habitat: marine.

rivulata, Equula Temminck [C. J.] & Schlegel [H.] 1845:126, Pl. 67 (fig. 2) [Fauna Japonica Parts 7-9; ref. 4373] Nagasaki, Japan. Holotype: RMNH 1438. Type catalog: Boeseman 1947:110 [ref. 12876]. •Valid as Leiognathus rivulatus (Temminck & Schlegel 1845) -- (Uyeno & Yabumoto in Masuda et al. 1984:158 [ref. 6441], Nakabo 2000:812 [ref. 25086], Woodland in Randall & Lim 2000:617 [ref. 25122], Shimizu 2001:30 [ref. 25799], Nakabo 2002:812 [ref. 26001], Choi et al. 2003:334, 609 [ref. 26218], Ikejima et al. 2004:908 [ref. 27755], Sparks & Dunlap 2004:7 [ref. 27928], Ra et al. 2005:94 [ref. 28304]). •Valid as Photoplagios rivulatus (Temminck & Schlegel 1845) -- (Sparks et al. 2005:315 [ref. 28434], Sparks 2006:1 [ref. 28783], Sparks & Chakrabarty 2007:622 [ref. 29261]). •Valid as Equulites rivulatus (Temminck & Sclegel 1845) -- (Kimura et al. 2008:204 [ref. 29614], Chakrabarty et al. 2010:18 [ref. 30812], Suzuki & Kimura 2017:[12] [ref. 35231], Kim et al. 2020:129 [ref. 39118], Sonoyama et al. 2020:66 [ref. 37637], Suzuki & Kimura 2023:40 [ref. 40749]). Current status: Valid as Equulites rivulatus (Temminck & Schlegel 1845). Leiognathidae: Gazzinae. Distribution: Western Pacific: China, Taiwan, Korean Peninsula and Japan. Habitat: marine.

serrulifera, Equula Richardson [J.] 1848:137, Pl. 59 (fig. 12-14) [Ichthyology of the voyage of H. M. S. Erebus & Terror v. 2 (2); ref. 3740] Sydney, New South Wales, Australia [likely erroneous]. Syntypes: (7 or 8) BMNH 1848.3.18.168-170 (3). •Synonym of Leiognathus fasciatus (Lacepède 1803) -- (Jones 1985:587 [ref. 21842], Hoese & Bray 2006:1180 [ref. 29000]). •Synonym of Aurigequula fasciata (Lacepède 1803) -- (Chakrabarty & Sparks 2022:416 [ref. 39869]). Current status: Synonym of Aurigequula fasciata (Lacepède 1803). Leiognathidae: Leiognathinae. Habitat: brackish, marine.

simplex, Equula De Vis [C. W.] 1884:544 [Proceedings of the Linnean Society of New South Wales v. 9 (pt 3); ref. 4899] Cape York, Queensland, Australia. Lectotype: QM I.1702. Paralectotypes: QM I.9811 [ex QM I.1702] (2). Lectotype selected by Whitley 1932:111 [ref. 4675]. •Synonym of Leiognathus splendens (Cuvier 1829) -- (Jones 1985:599 [ref. 21842], Hoese & Bray 2006:1181 [ref. 29000]). •Synonym of Eubleekeria splendens (Cuvier 1829) -- (Kottelat 2013:338 [ref. 32989], Chakrabarty & Sparks 2022:421 [ref. 39869]). Current status: Synonym of Eubleekeria splendens (Cuvier 1829). Leiognathidae: Gazzinae. Habitat: brackish, marine.

smithursti, Equula Ramsay [E. P.] & Ogilby [J. D.] 1886:11 [Proceedings of the Linnean Society of New South Wales (Series 2) v. 1 (pt 1); ref. 14464] Hood Lagoon, Papua New Guinea. Holotype (unique): AMS B.9962. •Valid as Leiognathus smithursti (Ramsay & Ogilby 1886) -- (Kyushin et al. 1982:129 [ref. 19754], Jones 1985:597 [ref. 21842], Allen & Swainston 1988:76 [ref. 25477], Mohsin & Ambak 1996:349 [ref. 27969], Allen 1997:118 [ref. 23977], Woodland in Randall & Lim 2000:617 [ref. 25122], Kimura & Peristiwady in Matsuura & Peristiwady 2000:226 [ref. 27511], Hutchins 2001:33 [ref. 25847]). •Mention -- (Randall 2005:240 [ref. 28239]). •Synonym of Leiognathus longispinis (Valenciennes 1835) -- (Sparks 2006:539 [ref. 28801]). •Synonym of Aurigequula longispina (Valenciennes 1835) -- (Larson et al. 2013:123 [ref. 32988], Chakrabarty & Sparks 2022:416 [ref. 39869]). •Synonym of Aurigequula longispinis (Valenciennes 1835). Current status: Synonym of Aurigequula longispinis (Valenciennes 1835). Leiognathidae: Leiognathinae. Habitat: marine.

splendens, Equula Cuvier [G.] 1829:212 [Le Règne Animal (Edition 2) v. 2; ref. 995] Chennai, India [original locality: Vizagapatam, India]. Neotype: RMNH 1441. Type catalog: Bauchot & Desoutter 1989:34 [ref. 15502]. Available from footnote as, "Eq. splendens, N., Russel, 61" [Russell 1803, v. 1:48, Pl. 61]. Lectotype selected by Fricke 1999:262 [ref. 24106] as the specimen shown in Russell's Pl. 61; Kimura et al. 2005:289 [ref. 28424] designated RMNH 1441 as the neotype, contending the the figure could be either of two species. •Valid as Leiognathus splendens (Cuvier 1829) -- (Kyushin et al. 1982:130 [ref. 19754], Dor 1984:136 [ref. 29757], Uyeno & Yabumoto in Masuda et al. 1984:159 [ref. 6441], Jones 1985:599 [ref. 21842], Talwar & Jhingran 1991:832 [ref. 20764], Kottelat et al. 1993:111 [ref. 23448], Goren & Dor 1994:36 [ref. 25356], Mohsin & Ambak 1996:349 [ref. 27969], Larson & Williams 1997:359 [ref. 23967], Fricke 1999:262 [ref. 24106], Mishra & Srinivasan 1999:247 [ref. 24754], Mishra et al. 1999:86 [ref. 25315], Nakabo 2000:811 [ref. 25086], Woodland in Randall & Lim 2000:617 [ref. 25122], Bijukumar & Sushama 2000:186 [ref. 25703], Kimura & Peristiwady in Matsuura & Peristiwady 2000:227 [ref. 27511], Sakai et al. 2001:97 [ref. 25693], Hutchins 2001:33 [ref. 25847], Woodland et al. 2001:2815 [ref. 26095], Nakabo 2002:811 [ref. 26001], Manilo & Bogorodsky 2003:S108 [ref. 27377], Kimura et al. in Kimura & Matsuura 2003:84 [ref. 27510], Mishra & Krishnan 2003:33 [ref. 27712], Ikejima et al. 2004:908 [ref. 27755], Sparks & Dunlap 2004:7 [ref. 27928], Randall 2005:240 [ref. 28239], Kimura et al. 2005:285 [ref. 28424], Hoese & Bray 2006:1181 [ref. 29000]). •Valid as Eubleekeria splendens (Cuvier 1829) -- (Chakrabarty et al. 2009:152 [ref. 30295] but as Equulites splendens in key on p. 149). •Valid as Eubleekeria splendens (Cuvier 1829) -- (Kimura et al. 2008:202 [ref. 30867], Chakrabarty & Sparks 2008:6 [ref. 29788], Chakrabarty et al. 2008:152 [ref. 32414], Chakrabarty et al. 2010:112 [ref. 31090], Abraham et al. 2011:3 [ref. 31311], Larson et al. 2013:127 [ref. 32988], Kottelat 2013:338 [ref. 32989], Fricke et al. 2014:84 [ref. 33932], Miesen et al. 2016:89 [ref. 34492], Motomura et al. 2017:123 [ref. 35490], Ali et al. 2018:324 [ref. 36183], Fricke et al. 2018:190 [ref. 35805], Golani & Fricke 2018:88 [ref. 36273], Eagderi et al. 2019:76 [ref. 37020], Fricke et al. 2019:149 [ref. 36673], Habib & Islam 2020:Supplementary table p. 22 [ref. 38321], Sharifuzzaman et al. 2021:[3] [ref. 38872], Yuki et al. 2021:12 [ref. 38517], Chakrabarty & Sparks 2022:421 [ref. 39869]). Current status: Valid as Eubleekeria splendens (Cuvier 1829). Leiognathidae: Gazzinae. Distribution: Red Sea; Indo-West Pacific: East Africa, Persian Gulf, Madagascar and Mascarenes (La Réunion, Mauritius, Rodrigues) east to Philippines and Fiji, north to Tosa Bay (southern Japan), south to Kimberley (Western Australia), Queensland (Australia) and New Caledonia. Habitat: brackish, marine.

totta, Equula Cuvier [G.] 1829:212 [Le Règne Animal (Edition 2) v. 2; ref. 995] Indian seas [Vizagapatam, India]. No types known. Type catalog: Bauchot & Desoutter 1989:34 [ref. 15502]. Available from footnote as, "Eq. totta, Russel, 62." •Synonym of Leiognathus equula (Forsskål 1775) -- (Kottelat 2013:339 [ref. 32989] with question). Current status: Synonym of Leiognathus equula (Forsskål 1775). Leiognathidae: Leiognathinae. Habitat: freshwater, brackish, marine.