Institute for Biodiversity Science and Sustainability
Online Version, Updated 6 November 2023

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Catalog of Fishes

Species that contain: Lepadogaster and lepadogaster and lepadogaster [ 54 ] records

acutus, Lepadogaster Canestrini [G.] 1864:187 [Archivio per la Zoologia, l'Anatomia e la Fisiologia v. 3 (no. 1); ref. 714] Mediterranean Sea. No types known. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222], Briggs in Whitehead et al. 1986:1357 [ref. 13677], Hofrichter 1995:112 [ref. 30502]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

adherens, Piescephalus Rafinesque [C. S.] 1810:63 [Caratteri di alcuni nuovi generi e nuove specie di animali e piante della Sicilia; ref. 3594] Sicily, Italy, Mediterranean Sea. No types known. Also appeared in Rafinesque 1810:41 [ref. 3595]. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222], Hofrichter 1995:112 [ref. 30502], Henriques et al. 2002:338 [ref. 26172], Parenti 2019:103 [ref. 37117]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

adriatica, Lepadogaster Hankó [B.] 1920-21:177 [Annales Musei Nationalis Hungarici, Budapest v. 18 (for 1920-1921); ref. 15345] Northern Adriatic Sea (Mediterranean Sea). Syntypes: (26) Hungarian Natl. Mus. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637] with question, Briggs 1973:654 [ref. 7222], Hofrichter 1995:112 [ref. 30502]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

bacescui, Lepadogaster microcephalus Murgoci [A. A.] 1940:384 [Comptes Rendus des Séances de l'Institut des Sciences de Roumanie, ancienne Académie des Sciences de Roumanie, Bucuresti. v. 4 (nos 5-6); ref. 17803] Cap Caliacra [southern Dobruja Region, Bulgaria, Black Sea], depth 1-5 meters. Holotype: MGAB 54. •Synonym of Apletodon microcephalus (Brook 1890), but a valid subspecies bacescui -- (Briggs 1973:651 [ref. 7222]). •Synonym of Apletodon dentatus (Facciolà 1887), but a valid subspecies -- (Briggs in Whitehead et al. 1986:1352 [ref. 13677], Gus'kov et al. 2022:72 [ref. 39564]). •Synonym of Apletodon dentatus (Facciolà 1887) -- (Fricke et al. 2010:96 [ref. 30711], Parin et al. 2014:471 [ref. 33547]). Current status: Synonym of Apletodon dentatus (Facciolà 1887). Gobiesocidae: Lepadogastrinae. Distribution: Black Sea (if valid). Habitat: marine.

balbis, Lepadogaster Risso [A.] 1810:73, Pl. 4 (fig. 9) [Ichthyologie de Nice; ref. 3755] Villefranche-sur-Mer, France, northwestern Mediterranean Sea. No types known. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222]). •Synonym of Lepadogaster purpurea (Bonnaterre 1788) -- (Henriques et al. 2002:338 [ref. 26172]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

biciliatus, Lepadogaster Risso [A.] 1827:272 [Histoire naturelle des principales productions de l'Europe méridionale v. 3; ref. 3757] Nice, France, northwestern Mediterranean Sea. No types known. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222]). •Synonym of Lepadogaster purpurea (Bonnaterre 1788) -- (Henriques et al. 2002:338 [ref. 26172]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

brownii, Lepadogaster Risso [A.] 1827:272 [Histoire naturelle des principales productions de l'Europe méridionale v. 3; ref. 3757] Nice, France, northwestern Mediterranean Sea. No types known. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222], Hofrichter 1995:112 [ref. 30502], Bilecenoğlu et al. 2002:57 [ref. 26753], Lipej & Dulčić 2010:33 [ref. 36649]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

candolii, Lepadogaster Risso [A.] 1810:76 [Ichthyologie de Nice; ref. 3755] St. Hospice, France, northwestern Mediterranean Sea. Syntypes: MZUF 5573-74 (2). Type catalog: Vanni 1991:223 [ref. 20099]. Spelled candolii in text and index; name should not be changed. Spelled decandollii by Risso 1827:275 [ref. 3757]. •Valid as Mirbelia candolei [sic] (Risso 1810) "for the time being" -- (Almada et al. 2008:1154 [ref. 29550] as candollei). •Valid as Mirbelia candolii (Risso 1810) -- (Carballo et al. 2021:90 [ref. 39250]). •Valid as Lepadogaster candolii Risso 1810 -- (Briggs 1955:38 [ref. 637], Briggs 1973:655 [ref. 7222], Briggs in Whitehead et al. 1986:1356 [ref. 13677], Briggs 1990:476 [ref. 15498], Hofrichter 1995:124 [ref. 30502] as candollei, Bilecenoğlu et al. 2002:56 [ref. 26753] as candollei, Golani 2005:27 [ref. 37112] as candollei, Fricke et al. 2007:100 [ref. 29533], Vasil'eva 2007:135 [ref. 30517], Ninua & Japoshvili 2008:174 [ref. 31251] as candolii, Lipej & Dulčić 2010:33 [ref. 36649] as candollei, Parin et al. 2015:472 [ref. 33547], Fricke et al. in Carpenter & De Angelis 2016:2809 [ref. 34618], Artüz & Fricke 2019:554 [ref. 36612], Carneiro et al. 2019:205 [ref. 37250], Trkov et al. 2019:67 [ref. 36880], Bariche & Fricke 2020:48 [ref. 37515], Kovačić et al. 2020:31 [ref. 37519], Abliazov et al. 2021:11 [ref. 38576], Bañón & Maño 2021:63 [ref. 38883], Golani 2021:30 [ref. 38303] as candollei, Kovačić et al. 2021:55 [ref. 39014]). Current status: Valid as Lepadogaster candolii Risso 1810. Gobiesocidae: Lepadogastrinae. Distribution: Mediterranean Sea; Sea of Marmara; Black Sea; eastern Atlantic: British Isles south to Portugal, including Madeira and Canary Islands. Habitat: brackish, marine.

cephalus, Lepadogaster Thompson [W.] 1839:34 [Annals of Natural History, or Magazine of Zoology, Botany, and Geology v. 3 (no. 14) (art. 6); ref. 17907] Roundstone Bay, Connemara, western Ireland. Holotype (unique): whereabouts unknown. •Synonym of Lepadogaster candolii Risso 1810 -- (Briggs 1955:38 [ref. 637] as candollei, Briggs 1973:655 [ref. 7222] as candollei, Briggs 1990:477 [ref. 15498] as candollei, Hofrichter 1995:125 [ref. 30502] as candollei). Current status: Synonym of Lepadogaster candolii Risso 1810. Gobiesocidae: Lepadogastrinae. Habitat: marine.

ciliatus, Lepadogaster Risso [A.] 1820:248 [Journal de Physique, de Chimie et d'Histoire Naturelle v. 91; ref. 18636] Nice, France, northwestern Mediterranean Sea. No types known. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222], Hofrichter 1995:112 [ref. 30502]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

cornubicus, Cyclopterus Shaw [G.] 1804:397 [General zoology or systematic natural history; ref. 4015] Cornwall coast and Sound of Jura. No types known. Based on literature sources. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:36 [ref. 637], Briggs 1973:654 [ref. 7222], Briggs in Whitehead et al. 1986:1357 [ref. 13677], Briggs 1990:476 [ref. 15498]). •Synonym of Lepadogaster purpurea (Bonnaterre 1788) -- (Henriques et al. 2002:338 [ref. 26172]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

cornubiensis, Lepadogaster Fleming [J.] 1828:189 [A history of British animals; ref. 1339] Southern coast of England and the Hebrides. No types known. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:36 [ref. 637], Briggs 1990:476 [ref. 15498]). •Synonym of Lepadogaster purpurea (Bonnaterre 1788) -- (Henriques et al. 2002:338 [ref. 26172]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

dentatus, Lepadogaster Facciolà [L.] 1887:165, Pl. 3 (fig. 2) [Naturalista Siciliano [Il Naturalista Siciliano, Giornale di scienze naturali] v. 6; ref. 16780] Sicily, Italy, Mediterranean Sea. No types known. Technically a junior synonym of Lepadogaster lineatus Guichenot 1850, Lepadogaster maculatus Guichenot 1850 and Lepadogaster punctatus Guichenot 1850, but treated as valid by prevailing usage; here treated as a nomen protectum. •Synonym of Diplecogaster bimaculatus (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637]). •Valid as Apletodon dentatus (Facciolà 1887), but a valid subspecies dentatus (Facciolà 1887) -- (Bañón & Maño 2021:63 [ref. 38883]). •Valid as Apletodon dentatus (Facciolà 1887) -- (Briggs in Whitehead et al. 1986:1352 [ref. 13677], Hofrichter 1995:145 [ref. 30502], Hofrichter & Patzner 1997:20 [ref. 23741], Bilecenoğlu et al. 2002:56 [ref. 26753], Fricke 2007:69 [ref. 29022], Fricke et al. 2007:100 [ref. 29533], Vasil'eva 2007:134 [ref. 30517], Fricke et al. 2010:96 [ref. 30711], Parin et al. 2014:471 [ref. 33547], Fricke et al. in Carpenter & De Angelis 2016:2809 [ref. 34618], Fricke & Wirtz 2018:6 [ref. 36380], Artüz & Fricke 2019:554 [ref. 36612], Carneiro et al. 2019:204 [ref. 37250], Kovačić et al. 2020:30 [ref. 37519], Kovačić et al. 2021:55 [ref. 39014], Sahraoui & Refes in Gerovasileiou et al. 2020:343 [ref. 37890]). Nomen Protectum. Current status: Valid as Apletodon dentatus (Facciolà 1887). Gobiesocidae: Lepadogastrinae. Distribution: Mediterranean Sea; Sea of Marmara; northeastern Atlantic: British Isles south to Portugal, including Madeira and Canary Islands. Habitat: marine.

desfontanii, Lepadogaster Risso [A.] 1827:275, Pl. 14 (fig. 39) [Histoire naturelle des principales productions de l'Europe méridionale v. 3; ref. 3757] Nice, France, northwestern Mediterranean Sea. Syntypes: MZUF 5584-0093 [ex MNHN] (10). Type catalog: Vanni 1991:223 [ref. 20099], Nocita & Vanni 1998:69 [ref. 36144]. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Briggs in Whitehead et al. 1986:1354 [ref. 13677], Hofrichter 1995:138 [ref. 30502], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

gouani, Lepadogaster Walbaum [J. J.] 1792:693 [Petri Artedi sueci genera piscium Part 3; ref. 4572] No locality stated. No types known. Originally goüani, but the 'ü' should be treated as a vocal separate from the 'o', spoken 'go-u-an. Name is correctly gouani (R. Fricke, pers. comm., 22 Mar. 2008). •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Lipej & Dulčić 2010:33 [ref. 36649] as gouanii, Parin et al. 2015:472 [ref. 33547]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

gouanii, Lepadogaster Risso [A.] 1810:72 [Ichthyologie de Nice; ref. 3755] Nice, France, northwestern Mediterranean Sea. No types known. Spelled govani in Risso 1827:271 [ref. 3757]. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222], Briggs in Whitehead et al. 1986:1357 [ref. 13677], Hofrichter 1995:112 [ref. 30502], Parin et al. 2015:472 [ref. 33547]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

jussieui, Lepadogaster Risso [A.] 1827:273 [Histoire naturelle des principales productions de l'Europe méridionale v. 3; ref. 3757] Villefranche, France, northwestern Mediterranean Sea. No types known. •Synonym of Lepadogaster candolii Risso 1810 -- (Briggs 1955:38 [ref. 637] as candollei, Briggs 1973:655 [ref. 7222] as candollei, Hofrichter 1995:124 [ref. 30502] as candollei). Current status: Synonym of Lepadogaster candolii Risso 1810. Gobiesocidae: Lepadogastrinae. Habitat: marine.

latirostris, Lepadogaster Costa [O. G.] 1840:Lepadogaster p. 4 [Fauna del regno di Napoli Part 1; ref. 976] Naples, Italy, Mediterranean Sea. Published 3 Jan 1840. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

lepadogaster, Cyclopterus Walbaum [J. J.] 1792:490 [Petri Artedi sueci genera piscium Part 3; ref. 4572] No locality stated. No types known. Perhaps independent of Cyclopterus lepadogaster [lepodogaster] Bonnaterre 1788 and therefore available. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Hofrichter 1995:112 [ref. 30502]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

lepodogaster, Cyclopterus Bonnaterre [J. P.] (ex Goüan) 1788:29, Pl. 86 (fig. 356) [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] Montpellier, Languedoc-Roussillon Region, France, western Mediterranean Sea. No types known. Originally spelled lepodogaster (spelling error), but treated (perhaps incorrectly) by authors as lepadogaster; continuation of current usage is suggested. Based on Lepadogaster of Goüan 1770:106 [ref. 1863]. Misspelled lepidogaster by Cabrera y Corro, Pérez & Hänseler in Graells 1887:181 [ref. 30438]. •Valid as Lepadogaster lepadogaster subspecies lepadogaster (Bonnaterre 1788) -- (Bilecenoğlu et al. 2002:57 [ref. 26753], Golani 2005:27 [ref. 37112], Golani 2021:30 [ref. 38303]). •Valid as Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222], Briggs in Whitehead et al. 1986:1357 [ref. 13677], Briggs 1990:475 [ref. 15498], Hofrichter 1995:111 [ref. 30502], Henriques et al. 2002:334 [ref. 26172], Almada et al. 2008:1154 [ref. 29550], Fricke et al. 2007:100 [ref. 29533], Vasil'eva 2007:136 [ref. 30517], Ninua & Japoshvili 2008:174 [ref. 31251], Lipej & Dulčić 2010:33 [ref. 36649], Parin et al. 2015:472 [ref. 33547], Fricke et al. in Carpenter & De Angelis 2016:2809 [ref. 34618], Artüz & Fricke 2019:554 [ref. 36612], Carneiro et al. 2019:205 [ref. 37250], Trkov et al. 2019:67 [ref. 36880], Bariche & Fricke 2020:49 [ref. 37515], Kovačić et al. 2020:31 [ref. 37519], Sahraoui & Refes in Gerovasileiou et al. 2020:343 [ref. 37890], Abliazov et al. 2021:11 [ref. 38576], Bañón & Maño 2021:62 [ref. 38883], Carballo et al. 2021:86 [ref. 39250], Kovačić et al. 2021:55 [ref. 39014], Ragkousis et al. 2021:12 [ref. 38358] as cf. lepadogaster). Current status: Valid as Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Distribution: Mediterranean Sea; Sea of Marmara; Black Sea; eastern Atlantic: Galicia (Spain) south to Portugal, including Madeira and Canary Islands. Habitat: brackish, marine.

lineatus, Lepadogaster Guichenot [A.] 1850:110, Pl. 6 (fig. 3) [Histoire naturelle des reptiles et poissons; ref. 1940] Algeria, western Mediterranean Sea. Holotype (unique): MNHN A-4963. Technically a senior synonym of Lepadogaster dentatus Facciolà 1887, but not used as valid; therefore here treated as a nomen oblitum. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222]). •Synonym of Apletodon dentatus (Facciolà 1887) -- (Hofrichter 1995:145 [ref. 30502]). Nomen Oblitum. Current status: Synonym of Apletodon dentatus (Facciolà 1887). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

listellus, Lepadogaster Nardo [G. D.] (ex Chiereghini) 1847:col. 113 [Sinonimia moderna delle specie registrate nell' opera intitolata: ...; ref. 17994] Italy. No types known. Available as above with brief description. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Hofrichter 1995:138 [ref. 30502], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

maculatus, Lepadogaster Guichenot [A.] 1850:109, Pl. 6 (fig. 4) [Histoire naturelle des reptiles et poissons; ref. 1940] Algeria, western Mediterranean Sea. Holotype: MNHN A-4961. Technically a senior synonym of Lepadogaster dentatus Facciolà 1887, but not used as valid; therefore here treated as a nomen oblitum. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222]). •Synonym of Apletodon dentatus (Facciolà 1887) -- (Hofrichter 1995:145 [ref. 30502]). Nomen Oblitum. Current status: Synonym of Apletodon dentatus (Facciolà 1887). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

maeandricus, Lepadogaster Girard [C. F.] 1858:130 [General report upon zoology of the several Pacific railroad routes, 1857; ref. 4911] San Luis Obispo, California, U.S.A. Syntypes: ?USNM 516 to 518 (1, 1, 1). Replacement name for Lepadogaster reticulatus Girard 1854, preoccupied by Lepadogaster reticulatus Risso 1810. •Valid as Gobiesox maeandricus (Girard 1858) -- (Briggs 1955:101 [ref. 637], Eschmeyer & Herald 1983:109 [ref. 9277], Matarese et al. 1989:230 [ref. 26885], McAllister 1990:228 [ref. 14674], Mecklenburg et al. 2002:798 [ref. 25968], Nelson et al. 2004:167 [ref. 27807], Love et al. 2005:159 [ref. 37547], Page et al. 2013:173 [ref. 32708], Pietsch & Orr 2015:66 [ref. 34694], Kells et al. 2016:300 [ref. 35888], Hastings & Conway 2017:396 [ref. 35062] as meandricus, Conway et al. 2017:[9] [ref. 35287], Burton & Lea 2019:90 [ref. 37205], Love et al. 2021:201 [ref. 39279]). Replacement Name. Current status: Valid as Gobiesox maeandricus (Girard 1858). Gobiesocidae: Gobiesocinae. Distribution: Eastern Pacific: southeastern Alaska (U.S.A.) south to Pacific coast of central Baja California (Mexico), including Guadalupe Island (Mexico). Habitat: marine.

microcephalus, Lepadogaster Brook [G.] 1890:166, Pl. 7 (figs. 1-4) [Proceedings of the Royal Physical Society of Edinburgh v. 10; ref. 17286] West coast of Scotland. No types known. •Valid as Apletodon microcephalus (Brook 1890) -- (Briggs 1955:26 [ref. 637], Briggs 1973:651 [ref. 7222]). •Synonym of Apletodon dentatus (Facciolà 1887) -- (Briggs in Whitehead et al. 1986:1352 [ref. 13677], Hofrichter 1995:159 [ref. 30502], Bilecenoğlu et al. 2002:56 [ref. 26753], Fricke et al. 2010:96 [ref. 30711]). Current status: Synonym of Apletodon dentatus (Facciolà 1887). Gobiesocidae: Lepadogastrinae. Habitat: marine.

minimus, Lepadogaster Döderlein [L.] in Steindachner & Döderlein 1887:270 [14] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 53 (1. abth.); ref. 4249] Sagami Bay, Kanagawa Prefecture, Japan, depth 100-150 fathoms [circa 180-270 meters]. Lectotype: NMW 76955. Paralectotype: NMW 999353 (1) [ex NMW 76955]. Lectotype selected by Fujiwara & Motomura 2019:65 [ref. 36982]. •Valid as Aspasma minima (Döderlein 1887) -- (Briggs 1955:135 [ref. 637], Yoshino in Masuda et al. 1984:341 [ref. 6441], Lindberg et al. 1997:192 [ref. 23547], Hutchins in Randall & Lim 2000:598 [ref. 25122], Nakabo 2000:1122 [ref. 25182], Nakabo 2002:1122 [ref. 26193], Kwun et al. 2017:143 [ref. 37287]). •Valid as Kopua minima (Döderlein 1887) -- (Fujiwara & Motomura 2019:65 [ref. 36982], Fujiwara & Wada 2023:8 [ref. 40835]). Current status: Valid as Kopua minima (Döderlein 1887). Gobiesocidae: Protogobiesocinae. Distribution: Northwestern Pacific: East China Sea to Korea and Japan. Habitat: marine.

mirbeli, Lepadogaster Risso [A.] 1820:249 [Journal de Physique, de Chimie et d'Histoire Naturelle v. 91; ref. 18636] Nice, France, northwestern Mediterranean Sea. No types known. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Hofrichter 1995:138 [ref. 30502], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

natator, Lepadogaster Risso [A.] 1820:248 [Journal de Physique, de Chimie et d'Histoire Naturelle v. 91; ref. 18636] Nice, France, northwestern Mediterranean Sea. No types known. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:34 [ref. 637], Briggs 1973:654 [ref. 7222], Hofrichter 1995:112 [ref. 30502]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

norvegicus, Lepadogaster Düben [M. W. von] 1845:112 [Öfversigt af Kongliga Vetenskaps-Akademiens förhandlingar, Kungliga Svenska Vetenskapsakademien. v. 1 (1844); ref. 17421] Norway. No types known. Reidentified as Lepadogaster bimaculatus Pennant in Düben & Koren 1846:11 [ref. 21662]. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Hofrichter 1995:138 [ref. 30502], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

ocellatus, Cyclopterus Donovan [E.] 1806:6 unnum. pp., Pl. 76 [The natural history of British fishes, including scientific and general descriptions of the most interesting species, and an extensive selection of accurately finished coloured plates.; ref. 17408] England. No types known. New name for some previous species, especially Cyclopterus cornubicus Shaw. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:36 [ref. 637], Briggs 1973:654 [ref. 7222], Briggs 1990:476 [ref. 15498]). •Synonym of Lepadogaster purpurea (Bonnaterre 1788) -- (Henriques et al. 2002:338 [ref. 26172]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

ocellatus, Lepadogaster Risso [A.] 1810:74 [Ichthyologie de Nice; ref. 3755] Villefranche-sur-Mer, France, northwestern Mediterranean Sea. No types known. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Hofrichter 1995:138 [ref. 30502], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

olivaceus, Lepadogaster Risso [A.] 1810:75 [Ichthyologie de Nice; ref. 3755] Chapelle de Saint-Hospice, Saint-Jean-Cap-Ferrat, France, northwestern Mediterranean Sea. No types known. •Synonym of Lepadogaster candolii Risso 1810 -- (Briggs 1955:38 [ref. 637] as candollei, Briggs 1973:655 [ref. 7222] as candollei, Briggs in Whitehead et al. 1986:1356 [ref. 13677] as candollei, Briggs 1990:477 [ref. 15498] as candollei, Hofrichter 1995:124 [ref. 30502] as candollei). Current status: Synonym of Lepadogaster candolii Risso 1810. Gobiesocidae: Lepadogastrinae. Habitat: marine.

ottaviani, Lepadogaster Cocco [A.] 1840:243 [Il Maurolico, Giornale di Scienze, Lettere e Arti v. 4 (fasc. 6); ref. 23130] Messina, Sicily, Italy, Straits of Messina, Mediterranean Sea. No types known. Briggs 1955:38 [ref. 637] had Lepadogaster ottaviani Cocco 1833 [wrong date and ref.]. •Synonym of Lepadogaster candolii Risso 1810 -- (Briggs 1955:38 [ref. 637] as candollei, Briggs 1973:655 [ref. 7222] as candollei, Briggs 1990:477 [ref. 15498] as candollei, Hofrichter 1995:125 [ref. 30502] as candollei). Current status: Synonym of Lepadogaster candolii Risso 1810. Gobiesocidae: Lepadogastrinae. Habitat: marine.

pellegrini, Lepadogaster (Mirbelia) Chabanaud [P.] 1925:283 [Bulletin du Muséum National d'Histoire Naturelle (Série 1) v. 31 (no. 4); ref. 15533] Northwestern Africa, eastern Atlantic. Syntypes: MNHN 1907-0257 (1) Senegal, 1925-0239 and 0240 (2), 1925-0241 (2) Cape Blanc. •Valid as Apletodon pellegrini (Chabanaud 1925) -- (Briggs 1955:27 [ref. 637], Briggs 1973:651 [ref. 7222], Briggs in Whitehead et al. 1986:1353 [ref. 13677], Briggs 1986:379 [ref. 5643], Briggs 1990:474 [ref. 15498], Hofrichter 1995:170 [ref. 30502], Santos et al. 1997:102 [ref. 23531], Hofrichter & Patzner 1997:20 [ref. 23741], Arruda 1997:124 [ref. 24952], Heemstra & Heemstra 2004:124 [ref. 28072], Fricke 2007:69 [ref. 29022], Fricke et al. 2010:96 [ref. 30711], Wirtz et al. 2013:132 [ref. 32972] but see barbatus Fricke et al. 2012, Fricke et al. in Carpenter & De Angelis 2016:2809 [ref. 34618], Fricke & Wirtz 2018:6 [ref. 36380], Carneiro et al. 2019:205 [ref. 37250], Carballo et al. 2021:80 [ref. 39250], Briggs 2022:437 [ref. 39965], Fermon et al. 2022:273 [ref. 39448] possible). Current status: Valid as Apletodon pellegrini (Chabanaud 1925). Gobiesocidae: Lepadogastrinae. Distribution: Eastern Atlantic: Portugal south to Cape of Good Hope (South Africa), including Azores (Portugal), Madeira (Portugal) and Canary Islands (Spain); southwestern Indian Ocean: Western Cape and eastern Cape (South Africa). Habitat: marine.

piger, Lepadogaster Nardo [G. D.] 1827:26, 35 (no. 31) [Giornale di fisica, chimica e storia naturale, medicina, ed arti. (series 2) Dec. II, v. 10; ref. 3146] Glavotok, Krk, Croatia, 45°05'44.9"N, 14°26'32.4"E [Adriatic Sea (Mediterranean Sea)]. Neotype: PMR VP3529. Also in Nardo 1827:col. 477 and 483 [ref. 19740] and Nardo 1827:105 [ref. 26236]. Neotype designated by Wagner et al. 2020:17 [ref. 37868]. •Synonym of Gouania willdenowi (Risso 1810) -- (Briggs 1955:23 [ref. 637] as wildenowi, Briggs 1973:653 [ref. 7222] as wildenowi, Briggs in Whitehead et al. 1986:1355 [ref. 13677] as wildenowi, Hofrichter 1995:103 [ref. 30502] as wildenowi). •Valid as Gouania pigra (Nardo 1827) -- (Wagner et al. 2020:14 [ref. 37868], Kovačić et al. 2021:55 [ref. 39014]). Current status: Valid as Gouania pigra (Nardo 1827). Gobiesocidae: Lepadogastrinae. Distribution: Mediterranean Sea: Adriatic Sea endemic. Habitat: marine.

pinnulatus, Lepadogaster Forster [J. R.] in Bloch & Schneider 1801:2 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Queen Charlotte Sound, Marlborough Sounds, South Island, New Zealand. No types known. Based on description in manuscript by Forster (Johann Reinhold Forster, 1729-1798, German explorer and naturalist of Cook's second voyage). •Valid as Trachelochismus pinnulatus (Forster 1801) -- (Briggs 1955:19 [ref. 637], Paulin et al. 1989:142 [ref. 24556], Paulin & Roberts 1992:46 [ref. 24625], Stewart 2015:1555 [ref. 34196], Conway et al. 2017:547 [ref. 35580]). Current status: Valid as Trachelochismus pinnulatus (Forster 1801). Gobiesocidae: Trachelochisminae. Distribution: New Zealand region. Habitat: marine.

punctatus, Lepadogaster Guichenot [A.] 1850:110, Pl. 6 (fig. 5) [Histoire naturelle des reptiles et poissons; ref. 1940] Algeria, western Mediterranean Sea. No types known. Technically a senior synonym of Lepadogaster dentatus Facciolà 1887, but not used as valid; therefore here treated as a nomen oblitum. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222]). •Synonym of Apletodon dentatus (Facciolà 1887) -- (Hofrichter 1995:145 [ref. 30502]). Nomen Oblitum. Current status: Synonym of Apletodon dentatus (Facciolà 1887). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

puniceus, Lepadogaster Richardson [J.] 1846:71, Pl. 43 (figs. 1-7) [Ichthyology of the voyage of H. M. S. Erebus & Terror v. 2 (2); ref. 3740] New Zealand. Holotype (unique): BMNH 1848.3.18.255. •Valid as Diplocrepis puniceus (Richardson 1846) -- (Briggs 1955:42 [ref. 637], Paulin et al. 1989:142 [ref. 24556], Paulin & Roberts 1992:47 [ref. 24625], Stewart 2015:1546 [ref. 34196], Conway et al. 2017:139 [ref. 35266], Conway et al. 2017:547 [ref. 35580]). Current status: Valid as Diplocrepis puniceus (Richardson 1846). Gobiesocidae: Diplocrepinae. Distribution: New Zealand. Habitat: marine.

purpureus, Cyclopterus Bonnaterre [J. P.] (ex Boriase) 1788:29 [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] Monts-Bay (Mounts Bay), Cornwall, England. No types known. Based on Borlase 1758:269, Pl. 25, fig. 68 [ref. 30228]. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788), but a valid subspecies purpurea (Bonnaterre 1788) -- (Briggs 1955:36 [ref. 637], Briggs 1973:654 [ref. 7222], Briggs in Whitehead et al. 1986:1357 [ref. 13677], Briggs 1990:475 [ref. 15498]). •Valid as Lepadogaster purpurea (Bonnaterre 1788) -- (Henriques et al. 2002:334 [ref. 26172], Carneiro et al. 2019:205 [ref. 37250], Kovačić et al. 2020:31 [ref. 37519], Bañón & Maño 2021:62 [ref. 38883], Carballo et al. 2021:88 [ref. 39250], Kovačić et al. 2021:56 [ref. 39014]). Current status: Valid as Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Distribution: Western Mediterranean Sea; eastern Atlantic: British Isles south to Senegal, including Azores, Madeira and Canary Islands. Habitat: marine.

rafinesqui, Lepadogaster Costa [O. G.] 1840:Lepadogaster p. 5, Pl. 24 (figs. 1-3) [Fauna del regno di Napoli Part 1; ref. 976] [Napoli] Naples, Italy, Mediterranean Sea. No types known. Published 4 Mar 1840. Based in part on Piescephalus adherens Rafinesque and Lepadogaster candolii Risso. •Synonym of Lepadogaster candolii Risso 1810 -- (Briggs 1955:38 [ref. 637] as candollei, Briggs 1973:655 [ref. 7222], Briggs 1990:477 [ref. 15498] as candollei, Hofrichter 1995:125 [ref. 30502] as candollei). Current status: Synonym of Lepadogaster candolii Risso 1810. Gobiesocidae: Lepadogastrinae. Habitat: marine.

raninus, Lepadogaster Nardo [G. D.] (ex Chiereghini) 1847:col. 113 [Sinonimia moderna delle specie registrate nell' opera intitolata: ...; ref. 17994] Italy. No types known. Apparently an available name based on Cyclopterus raninus Chiereghini (manuscript), with a short description. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Hofrichter 1995:138 [ref. 30502], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

reticulatus, Lepadogaster Risso [A.] 1810:77 [Ichthyologie de Nice; ref. 3755] Nice, France, northwestern Mediterranean Sea. No types known. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Hofrichter 1995:138 [ref. 30502], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

reticulatus, Lepadogaster Girard [C. F.] 1854:155 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 5769] San Luis Obispo, California, U.S.A. No types known. Objectively invalid; preoccupied by Lepadogaster reticulatus Risso 1810, replaced by Lepadogaster maeandricus Girard 1858. •Objective synonym of Gobiesox maeandricus (Girard 1858) -- (Briggs 1955:101 [ref. 637]). 1°Homonym. Current status: Synonym of Gobiesox maeandricus (Girard 1858). Gobiesocidae: Gobiesocinae. Habitat: marine.

rostratus, Lepadogaster Bloch [M. E.] & Schneider [J. G.] 1801:1 [M. E. Blochii, Systema Ichthyologiae; ref. 471] British sea. Syntypes: ZMB 4110 (2). Type catalog: Paepke 1999:83 [ref. 24282]. As gouani on p. xxiii. •Synonym of Lepadogaster lepadogaster (Bonaterre 1788) -- (Briggs 1955:36 [ref. 637], Briggs 1973:654 [ref. 7222], Briggs 1990:475 [ref. 15498]). •Synonym of Lepadogaster purpurea (Bonnaterre 1788) -- (Henriques et al. 2002:338 [ref. 26172]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

ruber, Lepadogaster Plucàr [E.] 1846:25 [Der Fischplatz zu Triest; ref. 18559] Trieste, Italy, Gulf of Trieste, Adriatic Sea (Mediterranean Sea). No types known. •Synonym of Lepadogaster candolii Risso 1810 -- (Briggs 1955:38 [ref. 637] as candollei, Briggs 1973:655 [ref. 7222] as candollei, Briggs 1990:476 [ref. 15498] as candollei, Hofrichter 1995:125 [ref. 30502] as candollei). Current status: Synonym of Lepadogaster candolii Risso 1810. Gobiesocidae: Lepadogastrinae. Habitat: marine.

smyrnensis, Gobius Bonnaterre [J. P.] (ex Koelreuter) 1788:66 [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] İzmir, Turkey. No types known. Based in part on Koelreuter 1764:439 et seq., Pl. 9 (fig. 5) [ref. 26164]. Gobiesocidae: Lepadogastrinae. Habitat: marine.

spathulata, Cycloptera Lacepède [B. G. E.] 1832:67 [v. 69; ref. 29724] No types known. Original not seen. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Briggs 1955:36 [ref. 637], Briggs 1973:654 [ref. 7222], Briggs in Whitehead et al. 1986:1357 [ref. 13677]). •Synonym of Lepadogaster purpurea (Bonnaterre 1788) -- (Henriques et al. 2002:338 [ref. 26172]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

spatula, Cyclopterus Cloquet [H.] 1818:296 (v. 12) [Dictionnaire des sciences Naturelles; ref. 852] English seas [Cornwall, Northeastern Atlantic]. No types known. •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) subspecies perpurea (Bonnaterre 1788) -- (Briggs 1973:654 [ref. 7222], Briggs 1990:476 [ref. 15498], Hofrichter 1995:112 [ref. 30502]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

stictopteryx, Lepadogaster Holt [E. W. L.] & Byrne [L. W.] 1899:589 [Proceedings of the Zoological Society of London 1898 (pt 4) (for 15 Nov. 1898); ref. 14706] Plymouth and Loch Craignish in Argyllshire, Great Britain. Syntypes: (3, plus young, whereabouts unknown). •Synonym of Apletodon microcephalus (Brook 1890) -- (Briggs 1955:26 [ref. 637], Briggs 1973:651 [ref. 7222]). •Synonym of Apletodon dentatus (Facciolà 1887) -- (Briggs in Whitehead et al. 1986:1352 [ref. 13677], Hofrichter 1995:159 [ref. 30502], Fricke et al. 2010:96 [ref. 30711]). Current status: Synonym of Apletodon dentatus (Facciolà 1887). Gobiesocidae: Lepadogastrinae. Habitat: marine.

testar, Lepadogaster Bloch [M. E.] & Schneider [J. G.] 1801:4, Pl. 1 [M. E. Blochii, Systema Ichthyologiae; ref. 471] West Indies. No types known. •Synonym of Gobiesox nudus (Linnaeus 1758) [based on Briggs] -- (Briggs 1955:94 [ref. 637], Ferraris in Reis et al. 2003:512 [ref. 27061]). •Synonym of Gobiesox cephalus Lacepède 1800. Current status: Synonym of Gobiesox cephalus Lacepède 1800. Gobiesocidae: Gobiesocinae. Habitat: freshwater.

urifasciatus, Lepadogaster Costa [O. G.] 1840:Lepadogaster p. 9 [Fauna del regno di Napoli Part 1; ref. 976] Gulf of Salerno, Italy, Mediterranean Sea. No types known. Published 3 May 1840. •Synonym of Diplecogaster bimaculata (Bonnaterre 1788) -- (Briggs 1955:29 [ref. 637], Briggs 1973:652 [ref. 7222], Hofrichter 1995:138 [ref. 30502], Fricke et al. 2015:5 [ref. 33962]). Current status: Synonym of Diplecogaster bimaculata (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.

webbianus, Lepadogaster Valenciennes [A.] 1843:85 [Ichthyologie des îles Canaries v. 2 (pt 2); ref. 4502] Canary Islands. Holotype (unique): MNHN A-4964. •Synonym of Lepadogaster lepadogaster purpurea (Bonnaterre 1788) -- (Briggs 1973:654 [ref. 7222], Briggs 1990:476 [ref. 15498]). Current status: Synonym of Lepadogaster purpurea (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: marine.

willdenowi, Lepadogaster Risso [A.] 1810:75, Pl. 4 (fig. 10) [Ichthyologie de Nice; ref. 3755] Cagnes-sur-mer, Nice, France, 43°39'22.1"N, 7°10'25.3"E [Nice, France, northwestern Mediterranean Sea]. Neotype: PMR VP4574. Appeared in text as "4. L. Willdenow. N. L. Willdenowi" Spelled Wildenowii in index; first reviser not researched; wildenowi seems to be common in the literature. Neotype designated by Wagner et al. 2020:19 [ref. 37868]. •Valid as Gouania willdenowi (Risso 1810) -- (Briggs 1955:23 [ref. 637] as wildenowi Briggs 1973:653 [ref. 7222] as wildenowi, Hofrichter 1995:103 [ref. 30502] as wildenowi, Bilecenoğlu et al. 2002:56 [ref. 26753], Fricke et al. 2007:100 [ref. 29533], Almada et al. 2008:1154 [ref. 29550], Golani 2005:27 [ref. 37112] as wildenowi, Lipej & Dulčić 2010:33 [ref. 36649] as wildenowi, Elbaraasi et al. 2019:94 [ref. 36864], Kovačić & Schembri 2019:289 [ref. 36757] as wildenowi, Wagner et al. 2019:2 [ref. 36765], Bariche & Fricke 2020:48 [ref. 37515], Kovačić et al. 2020:31 [ref. 37519] as wildenowi, Wagner et al. 2020:17 [ref. 37868], Golani 2021:30 [ref. 38303], Kovačić et al. 2021:55 [ref. 39014]). Current status: Valid as Gouania willdenowi (Risso 1810). Gobiesocidae: Lepadogastrinae. Distribution: Western Mediterranean Sea: east to Messina (Sicily, Italy). In eastern Mediterranean replaced by Gouania hofrichteri Wagner, Kovačić & Koblmüller 2020. Habitat: marine.

zebrinus, Lepadogaster Lowe [R. T.] 1839:88 [Proceedings of the Zoological Society of London 1839 (pt 7); ref. 2829] Off Madeira, eastern Atlantic. Holotype: whereabouts unknown. •Valid as Lepadogaster zebrinus Lowe 1839 -- (Briggs 1955:36 [ref. 637], Briggs 1973:655 [ref. 7222], Briggs in Whitehead et al. 1986:1358 [ref. 13677], Briggs 1990:476 [ref. 15498]). •Valid as Lepadogaster zebrina Lowe 1839 -- (Hofrichter 1995:131 [ref. 30502]). •Synonym of Lepadogaster lepadogaster (Bonnaterre 1788) -- (Henriques et al. 2002:338 [ref. 26172], Almada et al. 2008:1154 [ref. 29550]). Current status: Synonym of Lepadogaster lepadogaster (Bonnaterre 1788). Gobiesocidae: Lepadogastrinae. Habitat: brackish, marine.


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