Species that contain: Leuciscus and rutilus [ 43 ] records

affinis, Leuciscus Valenciennes [A.] in Cuvier & Valenciennes 1844:150 [Histoire naturelle des poissons v. 17; ref. 1010] Gand [=Ghent], Belgium. Holotype (unique): MNHN 0000-3354. Type catalog: Bertin & Estève 1948:57-58 [ref. 19574], Almaça 1969:1119 [ref. 20712]. •Based on a hybrid between Rutilus rutilus x Scardinius erythrophthalmus -- (Kottelat 1997:184 [ref. 22952]). Hybrid. Leuciscidae: Leuciscinae. Habitat: freshwater.

alburnoides, Leuciscus Steindachner [F.] 1866:263 [3], Pl. 1 (fig. 3) [Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften v. 54 (1. Abth.); ref. 18777] Stream near Merida, Spain. Lectotype: NMW 49749:2. Paralectotypes: NMW 49749:1, 3-15 (13), 49778 (12). Species not listed in earlier abstract (Steindachner 1866 [ref. 21716]). Also in Steindachner 1866:5 [ref. 18757]. Lectotype selected by Bogutskaya 2000:20 [ref. 24891]. •Valid as Rutilus alburnoides (Steindachner 1866) -- (Kottelat 1997:76 [ref. 22952]). •See Doadrio & Carmona 2006:200 [ref. 28975]. •Valid as Iberocypris alburnoides (Steindachner 1866) -- (Kottelat & Freyhof 2007:199 [ref. 29996]). •Valid as Leuciscus alburnoides Steindachner 1866 -- (Collares-Pereira et al. 1999:466 [ref. 39857]). •Valid as Tropidophoxinellus alburnoides (Steindachner 1866) -- (Bogutskaya 2000:20 [ref. 24891], Bogutskaya et al. 2000:62 [ref. 24445], Bogutskaya & Coad 2009:171 [ref. 30270]). •Valid as Squalius alburnoides (Steindachner 1866) -- (Alves et al. 2002:649 [ref. 28744], Collares-Pereira & Coelho 2010:707 [ref. 31439]). Current status: Valid as Squalius alburnoides (Steindachner 1866). Leuciscidae: Leuciscinae. Distribution: Southwestern Europe: Portugal and Spain. Habitat: freshwater.

altus, Leuciscus Valenciennes [A.] in Cuvier & Valenciennes 1844:237 [Histoire naturelle des poissons v. 17; ref. 1010] Lago Maggiore, Italy. Holotype (unique): whereabouts unknown. Based on a figure from Bonaparte. •Synonym of Rutilus aula (Bonaparte 1841) -- (Kottelat 1997:76 [ref. 22952]). •Synonym of Leucos aula (Bonaparte 1841) Current status: Synonym of Leucos aula (Bonaparte 1841). Leuciscidae: Leuciscinae. Habitat: freshwater.

arcasii, Leucos Steindachner [F.] 1866:14 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftlichen Classe v. 3; ref. 21709] Ebro River at Logroño, La Rioja, and Cailes [Queiles] River at Tudela, Navarra, Spain. Syntypes: (many) NMW 50639-49 (3, 2, 3, 4, 3, 30, 2, 5, 6, 20, 9), 50650-51 (3, 27), 49760 (4), 49752-777 (75 spec., 26 lots), 60743 (1), 60485 (1). Species first described in the abstract above, later described in more detail in Steindachner 1866 [ref. 14058], p. 199 [p. 2 of separate]; also appeared in Steindachner 1866:14 [p. 9 of separate] [ref. 18776] as Leuciscus (Leucos) arcasii and in Steindachner 1866:311 [ref. 21711] and Steindachner 1866:5 [ref. 18757] as Leuciscus arcasii. Misspelled arca by Seeley 1886:179 [ref. 30373] in genus Leuciscus. •Valid as Rutilus arcasii (Steindachner 1866) -- (Kottelat 1997:76 [ref. 22952]). •See Ketmaier et al. 1998:47 [ref. 23809]. •Valid as Chondrostoma arcasii (Steindachner 1866) -- (Doadrio & Carmona 2003:30 [ref. 28132], Coelho et al. 2005:202 [ref. 28430], Robalo et al. 2005:24 [ref. 28431], Gante et al. 2007:26 [ref. 29300], Gante et al. 2010:968 [ref. 35176]). •Valid as Achondrostoma arcasii (Steindachner 1866) -- (Robalo et al. 2007:370 [ref. 28878], Doadrio & Elvira 2007:296 [ref. 29510], Kottelat & Freyhof 2007:156 [ref. 29996], Perea et al. 2010:14 [ref. 31802], Doadrio et al. 2023:7 [ref. 40184]). Current status: Valid as Achondrostoma arcasii (Steindachner 1866). Leuciscidae: Leuciscinae. Distribution: Southwest Europe: Ebro and Levantine (Mijares, Turia and Palancia) drainages, Spain. Habitat: freshwater.

auratus, Leuciscus rutilus var. Yakovlev [V. E.] 1873:334 [Priroda (Moscow) v. 1, book 2, pt 2; ref. 19640] Volga River Delta, Russia. No types known. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

bolmensis, Leuciscus rutilus forma Malm [A. W.] 1877:560 [ ref. 2881] Lake Bolmen, Småland, Sweden. Syntypes: whereabouts unknown. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

caspicus, Leuciscus rutilus var. Yakovlev [V. E.] 1870:103 [3], Fig. 2 [On the new and little-known species of fish found in the mouths of the Volga; ref. 19610] Volga River Delta, Russia. Syntypes: whereabouts unknown. On p. 3 of separate. Also described in Yakovlev 1873:334 [ref. 19640]. Author also seen as Jakovlev. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Coad 1995:20 [ref. 23608], Kottelat 1997:81 [ref. 22952]). •Synonym of Rutilus rutilus (Linnaeus 1758), but a valid subspecies as described -- (Kazanskii 1928:8 [ref. 14268], Berg 1949:513 [ref. 20661], Mitrofanov et al. 1987:50 [ref. 37731], Bogutskaya 1997:179 [ref. 23566], Reshetnikov et al. 1997:741 [ref. 24702], Bogutskaya 1998:70 [ref. 23589], Bogutskaya & Naseka 2004:87 [ref. 28183], Bogutskaya & Iliadou 2006:294 [ref. 28494], Fricke et al. 2007:46 [ref. 29533]). •Synonym of Rutilus lacustris (Pallas 1814) -- (Levin et al. 2017:44 [ref. 35061], Esmaeili et al. 2018:36 [ref. 36089], Dyldin et al. 2020:87 [ref. 37569], Jouladeh-Roudbar et al. 2020:144 [ref. 37983]). •Valid as Rutilus caspicus (Yakovlev 1870) -- (Kottelat & Freyhof 2007:239 [ref. 29996], Esmaeili et al. 2010:371 [ref. 33391], Jouladeh-Roudbar et al. 2015:879 [ref. 34062], Esmaeili et al. 2017:56 [ref. 35254], Dyldin et al. 2020:81 [ref. 37569]). Current status: Valid as Rutilus caspicus (Yakovlev 1870). Leuciscidae: Leuciscinae. Distribution: Eurasia: Caspian Sea drainages. Habitat: freshwater.

caspius, Leuciscus frisii Lönnberg [E.] 1900:16 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 26, afd. 4 (no. 8); ref. 17561] Volga River delta, Russia. Holotype (unique): NRM. Described as a "race or subspecies." •Questionably in the synonymy of Rutilus frisii (Nordmann 1840) -- (Kottelat 1997:78 [ref. 22952]). •Synonym of Rutilus frisii (Nordmann 1840), but a valid subspecies as described -- (Bogutskaya 1997:178 [ref. 23566]). •Synonym of Rutilus frisii (Nordmann 1840) -- (Bogutskaya 1998:69 [ref. 23589], Fricke et al. 2007:46 [ref. 29533], Jouladeh-Roudbar et al. 2020:143 [ref. 37983]) Current status: Synonym of Rutilus frisii (Nordmann 1840). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

communis, Leuciscus rutilus var. Yakovlev [V. E.] 1873:334 [Priroda (Moscow) v. 1, book 2, pt 2; ref. 19640] Volga River Delta, Russia. No types known. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

communis, Leuciscus rutilus var. Rossikov [K. M.] 1895:No. 53 [Kavkazskoe Sel'skoe Khozyajstvo 54-56; ref. 23013] Terek River, Mosdok. No types known. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]) Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

daugawensis, Leuciscus rutilus var. Dybowski [B. N.] (ex Kessler) 1862:228, 237, Pl. 6 [Archiv für die Naturkunde Liv-, Ehst- und Kurlands. Zweiter Serie. Biologische Naturkunde. Dorpat v. 3; ref. 1168] Düna River, Riga, Latvia. No types at ZIN. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

dauricus, Leuciscus Valenciennes [A.] in Cuvier & Valenciennes 1844:149 [Histoire naturelle des poissons v. 17; ref. 1010] Waters of Daourie [Transbaikalia], Russia. On Cyprinus rutilus of Pallas. •Synonym of Phoxinus percnurus (Pallas 1814) -- (Berg 1949:574 [ref. 20661]). •Synonym of Eupallasella percnurus (Pallas 1814) -- (Kottelat 2006:31 [ref. 28831]). •Synonym of Rhynchocypris percnurus (Pallas 1814) -- (Bogutskaya et al. 2008:333 [ref. 30085]). Current status: Synonym of Rhynchocypris percnurus (Pallas 1814). Leuciscidae: Pseudaspininae. Habitat: freshwater.

dojranensis, Leuciscus pigus Karaman [S.] 1928:153, Fig. 2a [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 6; ref. 15903] Lake Dojran, Republic of Macedonia. Syntypes: (several) IEBMT. Also as new in Karaman 1929:172 [ref. 15904]. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). •Synonym of Rutilus rutilus (Linnaeus 1758), but a possible valid subspecies -- (Bogutskaya 1997:179 [ref. 23566]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

fluviatilis, Leuciscus rutilus var. Yakovlev [V. E.] 1873:334, Fig. 3 [Priroda (Moscow) v. 1, book 2, pt 2; ref. 19640] Volga River Delta, Russia. No types known. •Synonym of Rutilus rutilus (Linnaeus 1758), but a valid subspecies as described -- (Berg 1949:497 [ref. 20661], Mitrofanov et al. 1987:9 [ref. 37731], Bogutskaya 1997:179 [ref. 23566]). •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952], Fricke et al. 2007:46 [ref. 29533]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

frisii, Leuciscus Nordmann [A. von] 1840:487 [Voyage dans la Russie méridionale et la Crimée; ref. 3209] Market in Odessa, Danube, Bug, Dniester, and Dnieper rivers. Syntypes: ?NMW 50456 (2). Incorrectly spelled frizii by Slastenenko 1955-56:131 [ref. 6450]. •Valid as Rutilus frisii (Nordmann 1840) -- (Berg 1949:532 [ref. 20661], Mitrofanov et al. 1987:72 [ref. 37731], Coad 1995:19 [ref. 23608], Coad 1996:84 [ref. 25304], Bogutskaya 1997:178 [ref. 23566], Kottelat 1997:77 [ref. 22952], Reshetnikov et al. 1997:741 [ref. 24702], Bogutskaya 1998:69 [ref. 23589], Coad 1998:102 [ref. 23869], Bogutskaya et al. 2001:43 [ref. 26178], Bogutskaya & Naseka 2004:87 [ref. 28183], Vassilev & Pehlivanov 2005:170 [ref. 28369], Bogutskaya & Iliadou 2006:294 [ref. 28494], Fricke et al. 2007:46 [ref. 29533], Kottelat & Freyhof 2007:240 [ref. 29996], Ninua & Japoshvili 2008:167 [ref. 31251], Marić 2010:157 [ref. 31515], Perea et al. 2010:15 [ref. 31802], Parin et al. 2014:91 [ref. 33547] as subspecies, Çiçek et al. 2015:147 [ref. 35151], Levin et al. 2017:44 [ref. 35061], Özuluğ & Saç 2019:30 [ref. 36599], Çiçek et al. 2020:252 [ref. 37644], Dyldin et al. 2020:81, 105 [ref. 37569], Jouladeh-Roudbar et al. 2020:143 [ref. 37983], Kuljanishvili et al. 2020:[5] [ref. 37535], Çiçek et al. 2023:408 [ref. 40625], Dyldin et al. 2023:[34] [ref. 40328], Çiçek et al. 2024:251 [ref. 40789]). Current status: Valid as Rutilus frisii (Nordmann 1840). Leuciscidae: Leuciscinae. Distribution: Eurasia: Black, Azov and Caspian Sea basins. Habitat: freshwater, brackish.

heckelii, Leuciscus Nordmann [A. von] 1840:491, Pl. 23 (fig. 1) [Voyage dans la Russie méridionale et la Crimée; ref. 3209] Crimea, Ukraine, Black Sea. No types known. Syntypes: ZIN 3008-09 (3). •Synonym of Rutilus rutilus (Linnaeus 1758), but a valid subspecies -- (Berg 1949:500 [ref. 20661], Bogutskaya 1997:179 [ref. 23566], Michajlova 1989:25 [ref. 20062] as heckeli). •Questionably valid as Rutilus heckelii (Nordmann 1840) -- (Kottelat 1997:78 [ref. 22952]). •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Bogutskaya 1998:70 [ref. 23589] as heckeli, Parin et al. 2014:92 [ref. 33547]). •Synonym of Rutilus lacustris (Pallas 1814) -- (Levin et al. 2017:44 [ref. 35061], Dyldin et al. 2020:87 [ref. 37569], Jouladeh-Roudbar et al. 2020:144 [ref. 37983]). •Valid as Rutilus heckelii (Nordmann 1840) -- (Fricke et al. 2007:46 [ref. 29533], Kottelat & Freyhof 2007:241 [ref. 29996], Marić 2010:157 [ref. 31515] as heckeli, Perea et al. 2010:15 [ref. 31802], Dyldin et al. 2020:81 [ref. 37569]). Current status: Valid as Rutilus heckelii (Nordmann 1840). Leuciscidae: Leuciscinae. Distribution: Eurasia: Black Sea basins. Habitat: freshwater, brackish.

jurinii, Leuciscus Dybowski [B. N.] 1862:230 [Archiv für die Naturkunde Liv-, Ehst- und Kurlands. Zweiter Serie. Biologische Naturkunde. Dorpat v. 3; ref. 1168] Lake Geneva, Switzerland and France. See Kottelat 1997:81 [ref. 22952] for types. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

kutum, Leuciscus frisii var. Kamensky [S. N.] 1901:23 [German p. 154], Pl. 12 [Die Cypriniden der Kaukasusländer und ihrer angrenzenden Meere. 2.; ref. 14761] Southern Caspian Sea and tributaries. Syntypes: ?ZMT, none at ZIN. •Synonym of Rutilus frisii (Nordmann 1840), but a valid subspecies as described -- (Kazanskii 1928:13 [ref. 14268], Berg 1949:534 [ref. 20661], Mitrofanov et al. 1987:72 [ref. 37731], Reshetnikov et al. 1997:741 [ref. 24702]). •Valid as Rutilus frisii (Nordmann 1840), but a valid subspecies as described -- (Coad 1995:20 [ref. 23608], Parin et al. 2014:91 [ref. 33547]). •Synonym of Rutilus frisii (Nordmann 1840), subspecies caspius (Lönnberg 1900) -- (Bogutskaya 1997:178 [ref. 23566]). •Questionably in the synonymy of Rutilus frisii (Nordmann 1840) -- (Kottelat 1997:78 [ref. 22952]). •Synonym of Rutilus frisii (Nordmann 1840) -- (Bogutskaya 1998:69 [ref. 23589]). •Valid as Rutilus kutum (Kamensky 1901) -- (Bogutskaya & Iliadou 2006:294 [ref. 28494], Fricke et al. 2007:46 [ref. 29533], Esmaeili et al. 2010:371 [ref. 33391], Jouladeh-Roudbar et al. 2015:879 [ref. 34062], Esmaeili et al. 2017:56 [ref. 35254], Esmaeili et al. 2018:36 [ref. 36089], Dyldin et al. 2020:81, 105 [ref. 37569], Kaya et al. 2020:328 [ref. 37582], Dyldin et al. 2023:[34] [ref. 40328]). Current status: Valid as Rutilus kutum (Kamensky 1901). Leuciscidae: Leuciscinae. Distribution: Eurasia: Caspian Sea basin. Habitat: freshwater.

lascha, Leuciscus Costa [O. G.] 1838:Cyprinoides p. 19, Pl. 16 [Fauna del regno di Napoli Part 1; ref. 976] Italy. Syntypes: MNHN 0194 (1) Lake Fucino, near Naples. Type catalog: Bertin & Estève 1948:58 [ref. 19574], Almaça 1969:1118-1119 [ref. 20712]. •Synonym of Rutilus rubilio (Bonaparte 1837) -- (Bianco & Taraborelli 1985:136 [ref. 21914], Kottelat 1997:80 [ref. 22952]). •Synonym of Sarmarutilus rubilio (Bonaparte 1837). Current status: Synonym of Sarmarutilus rubilio (Bonaparte 1837). Leuciscidae: Leuciscinae. Habitat: freshwater.

lemmingii, Leuciscus Steindachner [F.] 1866:79 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftlichen Classe v. 3; ref. 21716] Guadiana River and tributaries near Merida; rivers Guadalquivir and Guadaira near Sevilla, Spain. Syntypes: NMW 52356-57 (4, 2), 52362 (2), 52366 (2), 52373 (4). Species first spelled as leminingii when described in the above abstract (apparently in error), spelling later appeared as lemmingii when later illustrated and described in more detail in Steindachner 1866:265 [p. 5 of separate], Pl. 1 [ref. 18777]. Also in Steindachner 1866:8, 9 [ref. 18757] as Chondrostoma lemmingii. •Valid as Rutilus lemmingii (Steindachner 1866) -- (Kottelat 1997:78 [ref. 22952]). •Valid as Chondrostoma lemmingii (Steindachner 1866) -- (Doadrio & Carmona 2003:30 [ref. 28132], Coelho et al. 2005:202 [ref. 28430], Gante et al. 2010:968 [ref. 35176], Çiftci et al. 2020:214 [ref. 37725]). •Valid as Iberochondrostoma lemmingii (Steindachner 1866) -- (Robalo et al. 2007:370 [ref. 28878], Doadrio & Elvira 2007:296 [ref. 29510], Kottelat & Freyhof 2007:197 [ref. 29996], Perea et al. 2010:15 [ref. 31802]). Current status: Valid as Iberochondrostoma lemmingii (Steindachner 1866). Leuciscidae: Leuciscinae. Distribution: Southwestern Europe: Ibearian Peninsula (Spain and Portugal). Habitat: freshwater.

lividus, Leuciscus Heckel [J. J.] 1843:1039 [49] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Balaton Lake, and Marizza River, Hungary. Syntypes: NMW 49978-79 (2, 3). On p. 49 of separate [ref. 2067]. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

macrolepidotus, Leuciscus Steindachner [F.] 1866:15 [10] [Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften v. 54 (1. Abth.); ref. 18776] Creeks near Alcobazar and Cintra, Portugal. Syntypes: NMW 49815 (4, now 3). Species not listed in earlier abstract (Steindachner 1966 [ref. 21715]). On. p. 10 of separate. Also in Steindachner 1866:6 [ref. 18757]. Objectively invalid; apparently preoccupied by Leuciscus macrolepidotus Ayres 1854; replaced by Chondrostoma oligolepis Robalo, Doadrio, Almada & Kottelat 2005. •Treated as valid as Rutilus macrolepidotus (Steindachner 1866) -- (Kottelat 1997:79 [ref. 22952]). •See Ketmaier et al. 1998:47 [ref. 23809]. •Incorrectly treated as valid as Chondrostoma macrolepidotum (Steindachner 1866) -- (Doadrio & Carmona 2003:30 [ref. 28132], Coelho et al. 2005:202 [ref. 28430]). •Synonym of Achondrostoma oligolepis (Robalo, Doadrio, Almada & Kottelat 2005). 1°Homonym. Current status: Synonym of Achondrostoma oligolepis (Robalo, Doadrio, Almada & Kottelat 2005). Leuciscidae: Leuciscinae. Habitat: freshwater.

meidingeri, Leuciscus Heckel [J. J.] 1851:148 [Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften v. 6; ref. 23010] Atter Lake, Austria. Syntypes: NMW 49826 (1), 50451-52 (1, 1). Also appeared in Heckel 1851:290 [ref. 18258], and in more detail in Heckel 1852:88, Pl. 9 [ref. 18260]. •Synonym of Pararutilus frisii (Nordmann 1840), but a valid subspecies -- (Lelek 1987:233 [ref. 22962] as meidingeri). •Valid as Rutilus meidingeri (Heckel 1851) -- (Kottelat 1997:79 [ref. 22952], Bogutskaya & Iliadou 2006:294 [ref. 28494], Kottelat & Freyhof 2007:242 [ref. 29996], Marić 2010:157 [ref. 31515]). Current status: Valid as Rutilus meidingeri (Heckel 1851). Leuciscidae: Leuciscinae. Distribution: Austria, formerly Germany. Habitat: freshwater.

pagellus, Leuciscus De Filippi [F.] 1844:400 [14] [Pesci finora osservati in Lombardia v. 1; ref. 19961] Lombardy, northern Italy. Syntypes: whereabouts unknown. On p. 14 of separate. Also appeared in De Filippi 1845:95 [ref. 18079]. •Synonym of Rutilus rubilio (Bonaparte 1837) -- (Bianco & Taraborelli 1985:136 [ref. 21914]). •Synonym of Rutilus aula (Bonaparte 1841) -- (Kottelat 1997:77 [ref. 22952]). •Synonym of Leucos aula (Bonaparte 1841) Current status: Synonym of Leucos aula (Bonaparte 1841). Leuciscidae: Leuciscinae. Habitat: freshwater.

pallens, Leuciscus Blanchard [C. E.] 1866:386, Fig. 88 [Les poissons des eaux douces de la France; ref. 310] Small rivers in the vicinity of Annecy, France. Syntypes: ?MNHN. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

pauperum, Leuciscus De Filippi [F.] 1844:401 [15] [Pesci finora osservati in Lombardia v. 1; ref. 19961] Lombardy, northern Italy. Syntypes: whereabouts unknown. On p. 15 of separate. Also appeared in De Filippi 1845:97 [ref. 18079]. •Synonym of Rutilus rubilio (Bonaparte 1837) -- (Bianco & Taraborelli 1985:136 [ref. 21914]). •Synonym of Rutilus aula (Bonaparte 1841) -- (Kottelat 1997:77 [ref. 22952]). •Synonym of Leucos aula (Bonaparte 1841) Current status: Synonym of Leucos aula (Bonaparte 1841). Leuciscidae: Leuciscinae. Habitat: freshwater.

pausingeri, Leuciscus Heckel [J. J.] 1843:1039 [49] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Egelsee, Austria. Syntypes: NMW 49884-86 (1, 1, 1). On p. 49 of separate [ref. 2067]. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

pleurobipunctatus, Rutilus Stephanidis [A.] 1939:19, Pl. 2 [The freshwater fishes of northern Greece and the island of Corfu.; ref. 21156] Thiamos, Lourou, Acheron and Evinos streams; Lake Trichonis, area of Kalamictisa; Corfu, Greece. Syntypes: whereabouts unknown. •Valid as Leuciscus pleurobipunctatus (Stephanidis 1939) -- (Bianco 1988:4 [ref. 22381], Kottelat 1997:69 [ref. 22952], Bianco et al. 1996:50 [ref. 26724]). •Valid as Phoxinellus pleurobipunctatus (Stephanidis 1939) -- (Economidis 1995:208 [ref. 23025]). •Valid as Telestes pleurobipunctatus (Stephanidis 1939) -- (Ketmaier et al. 1998:47 [ref. 23809], Bogutskaya et al. 2006:336 [ref. 29055], Kottelat & Freyhof 2007:286 [ref. 29996], Perea et al. 2010:15 [ref. 31802], Barbieri et al. 2015:68 [ref. 35029], Buj et al. 2019:7 [ref. 36636], Shumka et al. 2023:40 [ref. 40747]). Current status: Valid as Telestes pleurobipunctatus (Stephanidis 1939). Leuciscidae: Leuciscinae. Distribution: Southern Europe: coastal rivers in southern Albania to the Alfios River, southwestern Greece. Habitat: freshwater.

prasinus, Leuciscus Agassiz [L.] 1835:46, Pl. 2 [not Pl. 7] [Mémoires de la Société Neuchateloise des Sciences Naturelles v. 1; ref. 22] Lake Neuchâtel, Switzerland. Syntypes: MHNN 994 (2), 994B (2), 995 (1). Type catalog: Kottelat 1984:148 [ref. 21784]. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:80 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

roseus, Leuciscus Bonaparte [C. L.] 1839:fasc. 24, punt. 125, Pl. 114 (fig. 1) [Iconografia della fauna italica; ref. 4895] Piémonte, Italy. Holotype (unique): ANSP 6525. Bonaparte specimens: MNHN 0000-3916 (1) Rome. Type catalog: Bertin & Estève 1948:57 [ref. 19574], Almaça 1969:1118 [ref. 20712], Böhlke 1984:89 [ref. 13621]. Puntata written as 126 on first page, corrected to 125 in index. •Synonym of Rutilus pigus (Lacepède 1803) -- (Bianco & Taraborelli 1985:152 [ref. 21914], Kottelat 1997:79 [ref. 22952]). Current status: Synonym of Rutilus pigus (Lacepède 1803). Leuciscidae: Leuciscinae. Habitat: freshwater.

rubella, Leuciscus Bonaparte [C. L.] 1837:fasc. 20, punt. 103, Pl. 112 (fig. 1) [Iconografia della fauna italica; ref. 4893] Viterbo; Marche; Naples; Perugino; Temi; Reiti, Italy. Syntypes: ANSP 6487-6508 (22), MNHN 0000-3924 (1) Rome, NMW 50772 (1). Type catalog: Bertin & Estève 1948:57 [ref. 19574], Almaça 1969:1117 [ref. 20712], Böhlke 1984:89 [ref. 13621]. •Synonym of Rutilus rubilio (Bonaparte 1837) -- (Bianco & Taraborelli 1985:136 [ref. 21914], Kottelat 1997:80 [ref. 22952]). •Synonym of Sarmarutilus rubilio (Bonaparte 1837). Current status: Synonym of Sarmarutilus rubilio (Bonaparte 1837). Leuciscidae: Leuciscinae. Habitat: freshwater.

rubilio, Leuciscus Bonaparte [C. L.] 1837:fasc. 19, punt. 96, Pl. 111 (fig. 2) [Iconografia della fauna italica; ref. 4893] Lake Nemi, Italy. Lectotype: ANSP 6509 (93 mm SL). Paralectotypes: ANSP 6510-19 (10); ?NMW 49860 (1), 50772 (1). Type catalog: Böhlke 1984:89 [ref. 13621]. Lectotype selected by Bianco & Ketmaier 2014:394 [ref. 33417]. •Valid as Rutilus rubilio (Bonaparte 1837) -- (Bianco & Taraborelli 1985:136 [ref. 21914], Mrakovcic et al. 1995:184 [ref. 23079], Lelek 1987:230 [ref. 22962], Kottelat 1997:80 [ref. 22952], Bianco 1997:614 [ref. 26725], Bianco & Ketmaier 2001:194 [ref. 26728], Bogutskaya & Iliadou 2006:294 [ref. 28494], Kottelat & Freyhof 2007:245 [ref. 29996], Marić & Radujković 2009:145 [ref. 31438], Marić 2010:153 [ref. 31525], Perea 2010:15 [ref. 31802]). •Valid as Sarmarutilus rubilio (Bonaparte 1837) -- (Bianco & Ketmaier 2014:393 [ref. 33417], Lorenzoni et al. 2019:252 [ref. 37108], Petrosino et al. 2022:2 [ref. 39352]). Current status: Valid as Sarmarutilus rubilio (Bonaparte 1837). Leuciscidae: Leuciscinae. Distribution: Southern Europe: Italy endemic. Habitat: freshwater.

rutiloides, Leuciscus Selys-Longchamps [E. de] 1842:212, Pl. 7 (fig. 1) [Faune belge. Première partie; ref. 18717] Meuse River, Liège, Belgium. Holotype (unique): IRSNB 108. Type catalog: Walschaerts 1987:13 [ref. 20755]. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952], Bianco & Ketmaier 2014:382 [ref. 33417]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

rutilus, Cyprinus Linnaeus [C.] 1758:324 [Systema Naturae, Ed. X v. 1; ref. 2787] European lakes. Possible syntypes: LS 44 (1, right half-skin). Type catalog: Wheeler 1985:32 [ref. 20705]. For remarks on systematics see Kottelat 2006:49 [ref. 28831]. •Valid as Leuciscus rutilus (Linnaeus 1758) -- (Herrera 1896:20 [ref. 39336]). •Valid as Rutilus rutilus (Linnaeus 1758) -- (Berg 1949:494 [ref. 20661], Coad 1981:11 [ref. 22348], Cadwallader & Backhouse 1983:168 [ref. 25473], Mitrofanov et al. 1987:9 [ref. 37731], Paxton et al. 1989:217 [ref. 12442], Brumley in McDowall 1996:105 [ref. 22909], Chereshnev 1996:601 [ref. 24659], Kottelat 1997:80 [ref. 22952], Bogutskaya 1997:178 [ref. 23566], Reshetnikov et al. 1997:741 [ref. 24702], Bogutskaya 1998:70 [ref. 23589], Briolay et al. 1998:101 [ref. 25586], Coad 1998:102 [ref. 23869], Gabrielyan 2001:27 [ref. 25867], Bogutskaya et al. 2001:43 [ref. 26178], Bianco & Ketmaier 2001:200 [ref. 26728], Allen et al. 2002:362 [ref. 25930], Hanel 2003:48 [ref. 27894], Bogutskaya & Naseka 2004:87 [ref. 28183], Vassilev & Pehlivanov 2005:171 [ref. 28369], Hanel & Lusk 2005:188 [ref. 28804], Bogutskaya & Iliadou 2006:294 [ref. 28494], Kottelat 2006:49 [ref. 28831], Ocock et al. 2006:45 [ref. 28933], Allen et al. 2006:347 [ref. 29004], Fricke 2007:28 [ref. 30577], Fricke et al. 2007:46 [ref. 29533], Kottelat & Freyhof 2007:246 [ref. 29996], Ninua & Japoshvili 2008:167 [ref. 31251], Marić 2010:157 [ref. 31515], Perea et al. 2010:15 [ref. 31802], Esmaeili et al. 2010:371 [ref. 33391], Bianco & Ketmaier 2014:395 [ref. 33417], Parin et al. 2014:91 [ref. 33547], Jouladeh-Roudbar et al. 2015:879 [ref. 34062], Barbieri et al. 2015:62 [ref. 35029], Çiçek et al. 2015:147 [ref. 35151], Levin et al. 2017:39 [ref. 35061], Romanov et al. 2017:822 [ref. 35687], Kuiter 2018:97 [ref. 36369], Ćaleta et al. 2019:164 [ref. 37026], Dolgov et al. 2018:46 [ref. 41055], Lorenzoni et al. 2019:252 [ref. 37108], Özuluğ & Saç 2019:31 [ref. 36599], Çiçek et al. 2020:252 [ref. 37644], Jouladeh-Roudbar et al. 2020:145 [ref. 37983], Baikeche et al. 2021:959 [ref. 39080], Pozdeev et al. 2022:5 [ref. 39511], Çiçek et al. 2023:36 [ref. 40326], Çiçek et al. 2023:408 [ref. 40625], Dyldin et al. 2023:[34] [ref. 40328] with question, Nagy et al. 2023:357 [ref. 40348], Çiçek et al. 2024:252 [ref. 40789]). Current status: Valid as Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Distribution: Europe and western Asia. Widely introduced elsewhere. Habitat: freshwater, brackish.

ryzela, Leuciscus Valenciennes [A.] in Cuvier & Valenciennes 1844:199 [Histoire naturelle des poissons v. 17; ref. 1010] Turin, Italy. Syntypes: MNHN 0000-3935 (2). Type catalog: Bertin & Estève 1948:58 [ref. 19574]. Not the same as Chondrostoma rysela on p. 395 in same work. •Synonym of Rutilus pigus (Lacepède 1803) -- (Kottelat 1997:79 [ref. 22952]). Current status: Synonym of Rutilus pigus (Lacepède 1803). Leuciscidae: Leuciscinae. Habitat: freshwater.

sardella, Leuciscus Valenciennes [A.] in Cuvier & Valenciennes 1844:243 [Histoire naturelle des poissons v. 17; ref. 1010] Sarno River at confluence with Garigliano, Italy. No types known. •Synonym of Rutilus rubilio (Bonaparte 1837) -- (Kottelat 1997:80 [ref. 22952]). •Synonym of Sarmarutilus rubilio (Bonaparte 1837). Current status: Synonym of Sarmarutilus rubilio (Bonaparte 1837). Leuciscidae: Leuciscinae. Habitat: freshwater.

scardinus, Leuciscus De Filippi [F.] 1844:400 [14] [Pesci finora osservati in Lombardia v. 1; ref. 19961] Lombardy, northern Italy. Syntypes: whereabouts unknown. On p. 14 of separate. Also appeared in De Filippi 1845:96 [ref. 18079]. •Synonym of Rutilus rubilio (Bonaparte 1837) -- (Bianco & Taraborelli 1985:136 [ref. 21914]). •Synonym of Rutilus aula (Bonaparte 1841) -- (Kottelat 1997:77 [ref. 22952]). •Synonym of Leucos aula (Bonaparte 1841) Current status: Synonym of Leucos aula (Bonaparte 1841). Leuciscidae: Leuciscinae. Habitat: freshwater.

selysii, Leuciscus Selys-Longchamps [E. de] (ex Heckel) 1842:210, Pl. 6 (fig. 1) [Faune belge. Première partie; ref. 18717] Longchamps-sur-Geer, Belgium. Syntypes: IRSBN, ?MHNN 996 (1). Type catalog: Kottelat 1984:148 [ref. 21784]. Predates Leuciscus selysii Heckel 1843 (see Kottelat 1984:148 [ref. 21784]). •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952], Bianco & Ketmaier 2014:382 [ref. 33417]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

storerianus, Leuciscus Kirtland [J. P.] 1845:199 [Proceedings of the Boston Society of Natural History v. 1 (1841-1844); ref. 18409] Lake Erie, North America. No types known. Type catalog: Gilbert 1998:152 [ref. 23395]. Appeared first in Kirtland 1842:71 [ref. 18407] as "The true Rutilus plagurus of Raf., Dr. K. had been able finally to determine. The species formerly supposed by him [Kirtland] to be such, he now proposes to call R. Storerianus." No earlier treatment can be found; a full description under Leuciscus storerianus occurs as above, and the species apparently dates to that treatment. •Valid as Hybopsis storeriana (Kirtland 1845) -- (Lee et al. 1980:194 [ref. 22416], Smith 1986:139 [ref. 22186], Robison & Buchanan 1988:188 [ref. 23393], McAllister 1990:59 [ref. 14674], Scott & Crossman 1998:420 [ref. 24570]). •Valid as Macrhybopsis storeriana (Kirtland 1845) -- (Page & Burr 1991:104 [ref. 18983], Boschung 1992:52 [ref. 23239], Etnier & Starnes 1993:195 [ref. 22809], Warren et al. 1994:132 [ref. 25557], Stauffer et al. 1995:105 [ref. 23376], Cross & Collins 1995:79 [ref. 23394], Mettee et al. 1996:221 [ref. 25836], Gilbert 1998:152 [ref. 23395], Lyons et al. 2000:24 [ref. 26594], Ross et al. 2001:180 [ref. 25978], Nelson et al. 2004:73 [ref. 27807] dating to 1845, Bailey et al. 2004:81 [ref. 27882], Miller & Robison 2004:130 [ref. 27931], Boschung & Mayden 2004:210 [ref. 27995], Scharpf 2005:24 [ref. 28940], Page & Burr 2011:210 [ref. 31215], Page et al. 2013:73 [ref. 32708], Echelle et al. 2018:547 [ref. 35765], Page et al. 2023:72 [ref. 40505]). Current status: Valid as Macrhybopsis storeriana (Kirtland 1845). Leuciscidae: Pogonichthyinae. Distribution: North America: U.S.A. and Canada. Habitat: freshwater.

terekensis, Leuciscus rutilus var. Rossikov [K. M.] 1895:No. 54 [Kavkazskoe Sel'skoe Khozyajstvo 54-56; ref. 23013] Terek River, Mosdok, Russia. No types known. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.

trasimenicus, Leuciscus Bonaparte [C. L.] 1837:fasc. 20, punt. 103, Pl. 112 (fig. 4) [Iconografia della fauna italica; ref. 4893] Perugia, Natale, Lake Trasimeno, Italy. Syntypes: ANSP 5039-58 (20), ?NMW 50789 (1). Type catalog: Böhlke 1984:93 [ref. 13621]. •Synonym of Rutilus rubilio (Bonaparte 1837) -- (Bianco & Taraborelli 1985:136 [ref. 21914], Kottelat 1997:80 [ref. 22952]). •Synonym of Sarmarutilus rubilio (Bonaparte 1837). Current status: Synonym of Sarmarutilus rubilio (Bonaparte 1837). Leuciscidae: Leuciscinae. Habitat: freshwater.

virgo, Leuciscus Heckel [J. J.] 1852:69, Pls. 6-7 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 9; ref. 18260] Danube River near Vienna, Austria. Syntypes: NMW 22373 (1), 50626 (1), 94733 (4 pairs of pharyngeal bones). Type catalog: Palandačić et al. 2024:297 [ref. 41191]. •Synonym of Rutilus pigus (Lacepède 1803), but a valid subspecies -- (Lelek 1987:226, 227 [ref. 22962]). •Synonym of Rutilus pigus (Lacepède 1803) -- (Kottelat 1997:79 [ref. 22952]). •Valid as Rutilus virgo (Heckel 1852) -- (Bogutskaya & Iliadou 2006:294 [ref. 28494], Kottelat & Freyhof 2007:247 [ref. 29996], Bianco 2013:9 [ref. 32897], Ćaleta et al. 2015:37 [ref. 34877], Ćaleta et al. 2019:164 [ref. 37026], Nagy et al. 2023:357 [ref. 40348], Palandačić et al. 2024:298 [ref. 41191]). Current status: Valid as Rutilus virgo (Heckel 1852). Leuciscidae: Leuciscinae. Distribution: Europe. Habitat: freshwater.

vobla, Leuciscus rutilus var. Dikson [B. I.] 1909:31 [Rybolovstvo v basseine Volgi Vyche Saratova No. 8; ref. 23014] Saratow, Russia. No types known. Original not seen, from Kottelat 1997:81 [ref. 22952]. •Synonym of Rutilus rutilus (Linnaeus 1758) -- (Kottelat 1997:81 [ref. 22952]). Current status: Synonym of Rutilus rutilus (Linnaeus 1758). Leuciscidae: Leuciscinae. Habitat: freshwater, brackish.