Species that contain: Nemacheilus and barbatulus [ 17 ] records

barbatula, Cobitis Linnaeus [C.] 1758:303 [Systema Naturae, Ed. X v. 1; ref. 2787] Fresh waters of Europe and Asia. Syntypes: NRM 68 (1 or 3). Probably 3 syntypes with no geographic origin, Norén et al. 2018:64 [ref. 35742]. •Valid as Cobitis barbatula Linnaeus 1758 -- (Herrera 1896:25 [ref. 39336]). •Valid as Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:868 [ref. 20661] as Nemachilus, Lelek 1987:256 [ref. 22962] as Noemacheilus, Chereshnev 1996:601 [ref. 24659] as Noemacheilus). •Valid as Noemacheilus barbatulus (Linnaeus 1758), subspecies barbatulus -- (Mitrofanov & Dukravec 1989:58 [ref. 37748]). •Valid as Orthrias barbatulus (Linnaeus 1758) -- (Bănărescu & Nalbant 1995:449 [ref. 23187], Prokofiev 2003:726 [ref. 27494], Soric 2000:61 [ref. 28770]). •Valid as Orthrias barbatulus (Linnaeus 1758) and subspecies barbatulus (Linnaeus 1758) -- (Prokofiev 2007:15 [ref. 30343]). •Valid as Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:94 [ref. 23591], Novikov et al. 2000:161 [ref. 40929], Bogutskaya et al. 2001:45 [ref. 26178], Hanel 2003:55 [ref. 27894], Bogutskaya & Naseka 2004:112 [ref. 28183], Economidis 2005:67 [ref. 28213], Vassilev & Pehlivanov 2005:172 [ref. 28369], Hanel & Lusk 2005:290 [ref. 28804], Fricke et al. 2007:50 [ref. 29533], Kottelat & Freyhof 2007:329 [ref. 29996], Kottelat 2012:77 [ref. 32367], Parin et al. 2014:102 [ref. 33547], Barbieri et al. 2015:75 [ref. 35029], Çiçek et al. 2015:148 [ref. 35151], Norén et al. 2018:59 [ref. 35742], Tutman et al. 2018:220 [ref. 35876], Ćaleta et al. 2019:149 [ref. 37026], Gauliard et al. 2019:175 [ref. 36787], Lorenzoni et al. 2019:251 [ref. 37108], Pozdeev et al. 2022:6 [ref. 39511], Dyldin et al. 2023:[7] [ref. 40328], Nagy et al. 2023:356 [ref. 40348]). Current status: Valid as Barbatula barbatula (Linnaeus 1758). Nemacheilidae. Distribution: Northern and eastern Europe and western Asia. Habitat: freshwater, brackish.

caucasicus, Nemachilus barbatulus var. Berg [L. S.] 1898:67 [26] [Izvestija Kavkazskago Muzeja = Mitteilungen des Kaukasischen Museums = Bulletin du Musée du Caucase v. 1 (no. 3); ref. 20653] Argun River, Terek River basin, Russia. Syntypes: (many). Original genus is correctly Nemacheilus. On p. 26 of Russian issue. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies as described -- (Berg 1949:872 [ref. 20661] as Nemachilus, Lelek 1987:256 [ref. 22962] as Noemacheilus). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies -- (Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:95 [ref. 23591], Gabrielyan 2001:28 [ref. 25867], Ninua & Japoshvili 2008:167 [ref. 31251], Kottelat 2012:77 [ref. 32367] with question, Parin et al. 2014:103 [ref. 33547] with question). Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae. Habitat: freshwater, brackish.

compressirostris, Nemachilus Warpachowski [N. A.] 1897:270, Pl. 11 (figs. 1-3) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 2; ref. 4515] Lakes in northwestern Mongolia [possibly an error, probably a lake in the vicinity of Kosh-Agach]. Lectotype: ZIN 11298 (158 mm). Paralectotypes: ZIN 11298 (1). Original genus is correctly Nemacheilus. For remarks on systematics see Kottelat 2006:51 [ref. 28831]. Lectotype selected by Prokofiev 2016:392 [ref. 34693]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Barbatula toni (Dybowski 1869) -- (Vasil'eva 1998:95 [ref. 23591], Sideleva 2001:54 [ref. 26181], Sideleva 2003:11 [ref. 27815], Prokofiev 2016:392 [ref. 34693] with doubt). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Valid as Barbatula compressirostris (Warpachowski 1897) -- (Kottelat 2006:51 [ref. 28831], Cao et al. 2012:236 [ref. 32335], Kottelat 2012:78 [ref. 32367] with question, Prokofiev 2015:460 [ref. 34065] with question, Prokofiev 2018:273 [ref. 35878], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula compressirostris (Warpachowski 1897). Nemacheilidae. Distribution: East Asia: northwestern Mongolia and Tuva (Russia). Habitat: freshwater.

cuneicephalus, Barbatula Shaw [T.-H.] & Tchang [T.-L.] 1931:81, Fig. 10 [Bulletin of the Fan Memorial Institute of Biology, Peiping (Zoological Series) v. 2 (no. 5); ref. 16334] San-kai-tien, near Beijing [Peiping], China. Holotype (unique): ZMFMIB (now ASIZB) 6237. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:871 [ref. 20661] as Nemachilus). •Valid as Triplophysa cuneicephala (Shaw & Tchang 1931) -- (Zhu 1989:77 [ref. 17744], Zhu 1995:113 [ref. 25213], Wang et al. 2001:171 [ref. 26566], Kottelat 2012:124 [ref. 32367], Zhang et al. 2016:137 [ref. 34477]). Current status: Valid as Triplophysa cuneicephala (Shaw & Tchang 1931). Nemacheilidae. Distribution: China. Habitat: freshwater.

fowleri, Barbatula toni Nichols [J. T.] 1925:3 [American Museum Novitates No. 171; ref. 3178] Eastern Tombs, Chihli, Hopeh [Hebei] Province, China. Now: Zunhua, 40°11'N, 117°58'E, China. Holotype: AMNH 8409. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Barbatula barbatula (Linnaeus 1758), subspecies nuda (Bleeker 1868) -- (Zhu 1989:30 [ref. 17744]). •Synonym of Barbatula toni (Dybowski 1869), but a valid subspecies as described -- (Vasil'eva 1998:96 [ref. 23591]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula nuda (Bleeker 1869) with question -- (Kottelat 2012:78 [ref. 32367]). •Synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831], Prokofiev 2015:461 [ref. 34065]). •Synonym of Barbatula pechiliensis (Fowler 1899) -- (Chen et al. 2019:15 [ref. 36569]). Current status: Synonym of Barbatula pechiliensis (Fowler 1899). Nemacheilidae. Habitat: freshwater.

hispanica, Nemacheilus barbatulus Lelek [A.] 1987:256 [The freshwater fishes of Europe. Threatened fishes of Europe v. 9; ref. 22962] Nervion River at Durango and Tajo River, Spain. Syntypes: MGAB (11). Available by reference to infrasubspecific hispanica Bǎcescu-Meşter 1967, with Bǎcescu-Meşter specimens becoming the types. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies as described -- (Lelek 1987:256 [ref. 22962] as Noemacheilus). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Barbatula quignardi (Băcescu-Meşter1967) -- (Gauliard et al. 2019:176 [ref. 36787]). •Valid as Barbatula hispanica (Lelek 1987) -- (Denys et al. 2021:5 [ref. 38347]). Current status: Valid as Barbatula hispanica (Lelek 1987). Nemacheilidae. Distribution: Southwestern Europe: Ebro drainage and Cantabria (Spain); Adour drainage (southwestern France). Habitat: freshwater.

kirinensis, Barbatula toni Tchang [T.-L.] 1932:115, Fig. 2 [Bulletin of the Fan Memorial Institute of Biology, Peiping (Zoological Series) v. 3 (no. 8); ref. 16401] Kirin, Jilin Province, northeastern China. Holotype: ZMFMIB (now ASIZB) 7931. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:870 [ref. 20661] as Nemachilus). •Synonym of Barbatula barbatula (Linnaeus 1758), subspecies nuda (Bleeker 1868) -- (Zhu 1989:30 [ref. 17744]). •Synonym of Barbatula toni (Dybowski 1869), but a valid subspecies as described -- (Vasil'eva 1998:96 [ref. 23591]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula nuda (Bleeker 1964) -- (Kottelat 2012:79 [ref. 32367] with question). •Synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831], Cao et al. 2012:241 [ref. 32335], Prokofiev 2015:461 [ref. 34065]). •Valid as Barbatula kirinensis Tchang 1932 -- (Chen et al. 2019:12 [ref. 36569], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula kirinensis Tchang 1932. Nemacheilidae. Distribution: East Asia: Jingpo Lake, Tumen River and Zhu'Erduo River (northern China). Habitat: freshwater.

markakulensis, Nemacheilus barbatulus Menshikov [M. I.] 1939:141, Fig. [Ucenye zapiski / Permskij Ordena Trudovogo Krasnogo Znameni Gosudarstvennyj Universitet Imeni A. M. Gor'kogo, Ministerstvo Vyssego i Srednego Special'nogo Obrazovanija RSFSR. v. 3 (no. 2); ref. 21170] Lake Marka-kul', Irtysh basin, Kazakhstan. Syntypes: ZIN 26864 (6). Author also seen as Men'shikov. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:872 [ref. 20661] as Nemachilus). •Synonym of Noemacheilus barbatulus (Linnaeus 1758), but a valid subspecies markakulensis Manshikov 1939 -- (Mitrofanov & Dukravec 1989:60 [ref. 37748]). •Synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (authors). •Valid as Barbatula markakulensis (Menshikov 1939) -- (Kottelat 2012:78 [ref. 32367] with question, Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:396 [ref. 34693] as makakulensis, Prokofiev 2018:273 [ref. 35878]). Current status: Valid as Barbatula markakulensis (Menshikov 1939). Nemacheilidae. Distribution: North Asia: Markakul Lake (Kazakhstan). Habitat: freshwater.

oreas, Orthrias Jordan [D. S.] & Fowler [H. W.] 1903:769, Fig. 2 [Proceedings of the United States National Museum v. 26 (no. 1332); ref. 2461] Chitose in Iburi, Hokkaido, Japan. Holotype (unique): Sapporo Mus. Non-types: BMNH 1905.2.4.134 [from Jordan] (1). •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Barbatula barbatula (Linnaeus 1758), subspecies nuda (Bleeker 1868) -- (Zhu 1989:29 [ref. 17744]). •Synonym of Barbatula toni (Dybowski 1869) -- (Okada 1961:541 [ref. 6363], Vasil'eva 1998:95 [ref. 23591], Sideleva 2001:54 [ref. 26181], Sideleva 2003:11 [ref. 27815], Kottelat 2006:53 [ref. 28831]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies oreas (Jordan & Fowler 1903) -- (Prokofiev 2007:15 [ref. 30347]). •Valid as Barbatula oreas (Jordan & Fowler 1903) -- (Kottelat 2012:79 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula oreas (Jordan & Fowler 1903). Nemacheilidae. Distribution: East Asia: Hokkaido (Japan) and South Korea. Habitat: freshwater.

pechiliensis, Nemachilus Fowler [H. W.] 1899:181 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 51; ref. 17572] Tan lan Ho River, tributary of Shan lan Ho River, about 30 miles northeast of Lama-miau or Dolon-nor, Pechili Province, Inner Mongolia. Holotype (unique): ANSP 16394 (missing). Type catalog: Böhlke 1984:63 [ref. 13621]. Original genus is correctly Nemacheilus. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Barbatula toni (Dybowski 1869) -- (Okada 1961:541 [ref. 6363] as pichiliensis, Kottelat 2006:53 [ref. 28831]). •Synonym of Barbatula barbatula (Linnaeus 1758), subspecies nuda (Bleeker 1868) -- (Zhu 1989:29 [ref. 17744]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula toni fowleri Fowler 1925 -- (Cao et al. 2012:241 [ref. 35335]). •Synonym of Barbatula nuda (Bleeker 1964) -- (Kottelat 2012:78 [ref. 32367] with question). •Possible senior synonym of Barbatula potaninorum (Prokofiev 2007) -- (Prokofiev 2015:461 [ref. 34065]). •Valid as Barbatula pechiliensis (Fowler 1899) -- (Chen et al. 2019:15 [ref. 36569]). Current status: Valid as Barbatula pechiliensis (Fowler 1899). Nemacheilidae. Distribution: Luan, Hai, Yin and Nu'er River drainages and Dali-Nur Lake, northern China. Habitat: freshwater.

quignardi, Noemacheilus barbatulus Băcescu-Meşter [L.] 1967:359, Figs. 1 (A, B, C), 5 (C, D) [Travaux du Muséum d'Histoire Naturelle "Grigore Antipa" v. 7; ref. 7940] Le Lez River, near Montpellier, France. Holotype: MGAB 77. Paratypes: MGAB 78 (1), MGAB (16). Genus is correctly Nemacheilus. •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies as described -- (Vasil'eva 1998:95 [ref. 23591]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies quignardi (Băcescu-Meşter 1967) -- (Prokofiev 2007:15 [ref. 30343]). •Valid as Barbatula quignardi (Băcescu-Meşter 1967) -- (Kottelat & Freyhof 2007:330 [ref. 29996], Kottelat 2012:79 [ref. 32367], Gauliard et al. 2019:176 [ref. 36787]). Current status: Valid as Barbatula quignardi (Băcescu-Meşter 1967). Nemacheilidae. Distribution: Southern France and northwestern Spain. Habitat: freshwater.

sibiricus, Nemacheilus Gratzianov [V. I.] 1907:164, 168 [Trudy Otdela Ichtiologii Imperatorskago Russkago Obshchestva Akklimatizacii Zivotnych' i Rastenii. v. 4; ref. 1871] Biya River near Biysk [Biysk], Altai, Russia. Lectotype: ZUMM 2531 (140 mm). Paralectotypes: ZMMU P-2532 (now 2). Type catalog: Vasil'eva et al. 2001:25 [ref. 25714]. Lectotype selected by Prokofiev 2016:394 [ref. 34693]. Former syntypes ZMMGU 2532 (3) = Barbatula compressirostris, see Prokofiev 2018:273 [ref. 35878]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Nemacheilus toni (Dybowski 1869) -- (Bogutskaya & Naseka 1996:47 [ref. 22798]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (based on status of Barbatula). •Questionably the same as Nemacheilus tominus Ruzssky 1920 -- (Kottelat 2012:80 [ref. 32367]). •Synonym of Barbatula toni (Dybowski 1869) -- (Vasil'eva 1998:95 [ref. 23591], Sideleva 2001:54 [ref. 26181], Sideleva 2003:11 [ref. 27815], Kottelat 2006:53 [ref. 28831], Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:391 [ref. 34693] with question). Current status: Synonym of Barbatula toni (Dybowski 1869). Nemacheilidae. Habitat: freshwater.

sturanyi, Nemachilus Steindachner [F.] 1892:378 [22] [131], Pl. 2 (fig. 3) [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 59 (1. Abth.); ref. 14517] Lake Ohrid [Ochrida] at Pestani, between Ohrid City and Naum Monastery, Yugoslavia/Albania. Holotype (unique): NMW 48440. Original genus is correctly Nemacheilus. On p. 22 of separate; on p. 131 of abstract. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies -- (Lelek 1987:256 [ref. 22962] as Noemacheilus). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies -- (Vasil'eva 1998:95 [ref. 23591], Economidis 2005:67 [ref. 28213]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies sturanyi (Steindachner 1892) -- (Soric 2000:64 [ref. 28770], Prokofiev 2007:15 [ref. 30343]). •Valid as Barbatula sturanyi (Steindachner 1892) -- (Kottelat 2006:89 [ref. 28831], Kottelat & Freyhof 2007:331 [ref. 29996], Kottelat 2012:80 [ref. 32367], Shumka et al. 2023:36 [ref. 40747]). Current status: Valid as Barbatula sturanyi (Steindachner 1892). Nemacheilidae. Distribution: Southeastern Europe: Lake Ohrid basin and upper Drin River (Macedonia and Albania). Habitat: freshwater.

taurica, Nemachilus barbatulus var. Kessler [K. F.] 1877:171 [The Aralo-Caspian Expedition; ref. 2597] Selgir River, Crimea, Ukraine. Syntypes: ZIN 3029 (3), 3032 (3). Original genus is correctly Nemacheilus. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:868 [ref. 20661] as Nemachilus). •Synonym of Barbatula barbatula (Linnaeus 1758). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies barbatulus (Linnaeus 1758) -- (Prokofiev 2007:54 [ref. 30343]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (based on status of Barbatula) Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae. Habitat: freshwater, brackish.

tomianus, Nemacheilus barbatulus Ruzsky [M. D.] 1920:36, Figs. 1-2 [Trudy estestvenno-isthrichsk. Otdel., Tomsk No. 1; ref. 21169] Basins of the rivers Tom, Ob and Yenisei, Russia. Syntypes: not researched. See Reshetnikov et al. 1997:743 [ref. 24702]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:871 [ref. 20661] as Nemachilus). •Synonym of Barbatula toni (Dybowski 1869) -- (Vasil'eva 1998:96 [ref. 23591], Sideleva 2001:54 [ref. 26181], Sideleva 2003:11 [ref. 27815], Kottelat 2006:53 [ref. 28831]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Valid as Barbatula tomiana (Ruzsky 1920) -- (Kottelat 2012:80 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:399 [ref. 34693], Romanov et al. 2017:823 [ref. 35687], Prokofiev 2018:274 [ref. 35878], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula tomiana (Ruzsky 1920). Nemacheilidae. Distribution: East Asia: Altai Republic (Russia) and Mongolia. Habitat: freshwater.

toni, Cobitis Dybowski [B. N.] 1869:957, Pl. 18 (fig. 10) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 19; ref. 1169] Onon and Ingoda rivers, Amur River basin, Russia. Syntypes and/or Dybowski specimens: MNHN B-0738 (8), B-0739 (4); NMW 50473 (2). Type catalog: Bertin & Estève 1948:98 [ref. 19574]. For remarks on systematics see Kottelat 2006:53 [ref. 28831]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies -- (Berg 1949:869 [ref. 20661] as Nemachilus, Kawanabe & Mizuno 1989:398 [ref. 25598] as Neomacheilus, Watanabe 1998:263 [ref. 23491]). •Synonym of Noemacheilus barbatulus (Linnaeus 1758), but a valid subspecies toni -- (Chereshnev 1996:601 [ref. 24659], Mitrofanov & Dukravec 1989:58 [ref. 37748], Nakabo 2000:276 [ref. 25086], Nakabo 2002:276 [ref. 26001]). •Synonym of Barbatula barbatula, subspecies nuda (Bleeker 1865) -- (Zhu 1989:29 [ref. 17744]). •Valid as Nemacheilus toni (Dybowski 1869) -- (Wang 1984:82 [ref. 22951], Sawada in Masuda et al. 1984:59 [ref. 6441] as Noemacheilus, Bogutskaya & Naseka 1996:47 [ref. 22798], Matsuura et al. 2000:176 [ref. 26000], Choi et al. 2003:178, 517 [ref. 26218]). •Valid as Orthrias toni (Dybowski 1869) -- (Bănărescu & Nalbant 1995:449 [ref. 23187], Kim et al. 2000:224 [ref. 25434], Kim & Park 2002:193 [ref. 25913], Jang et al. 2003:119 [ref. 27681]). •Synonym of Orthrias nudus (Bleeker 1865) -- (Kim 1997:555 [ref. 24032]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies toni (Dybowski 1869) -- (Prokofiev 2003:736 [ref. 27494], Prokofiev 2007:17 [ref. 30343]). •Valid as Barbatula toni (Dybowski 1869) -- (Okada 1961:541 [ref. 6363], Nikiforov et al. 1997:330 [ref. 24694], Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:95 [ref. 23591], Sheiko & Fedorov 2000:17 [ref. 25650], Chereshnev et al. 2001:37 [ref. 26736], Tokranov 2001:268 [ref. 25407], Zemnukhov et al. 2001:420 [ref. 25412], Kottelat 2001:49 [ref. 25482], Pietsch et al. 2001:140 [ref. 25688], Bogutskaya et al. 2001:45 [ref. 26178], Sideleva 2001:54 [ref. 26181], Fedorov et al. 2003:25 [ref. 27579], Alekseyev et al. 2004:54 [ref. 27625], Sideleva 2003:11 [ref. 27815], Bogutskaya & Naseka 2004:113 [ref. 28183], Kottelat 2006:53 [ref. 28831], Ocock et al. 2006:28 [ref. 28933], Kottelat & Freyhof 2007:330 [ref. 29996], Bogutskaya et al. 2008:339 [ref. 30085], Cao et al. 2012:241 [ref. 32335], Kottelat 2012:80 [ref. 32367], Parin et al. 2014:103 [ref. 33547], Prokofiev 2015:460 [ref. 34065], Prokofiev 2016:399 [ref. 34693], Dyldin & Orlov 2016:669 [ref. 34853] with question, Prokofiev 2018:274 [ref. 35878], Chen et al. 2019:8 [ref. 36569], Dyldin et al. 2020:87 [ref. 37569], An et al. 2021:121 [ref. 38850], Dyldin et al. 2021:19 [ref. 38804] with question, Dyldin & Orlov 2021:63 [ref. 38158] with question, Dyldin et al. 2023:[8] [ref. 40328] with question). Current status: Valid as Barbatula toni (Dybowski 1869). Nemacheilidae. Distribution: East Asia: Amur (Heilongjiang) and Yin River drainages (Russia, Mongolia, northern China and northern Korea). Habitat: freshwater.

vardarensis, Nemachilus barbatulus Karaman [S.] 1928:164 [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 6; ref. 15903] Vardar River basin, former Yugoslavian republic of Macedonia. Original genus is correctly Nemacheilus. Also in Karaman 1929:173 [ref. 15903]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies as described -- (Lelek 1987:256 [ref. 22962] as Noemacheilus). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies vardarensis (Karaman 1928) -- (Soric 2000:62 [ref. 28770]). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies vardarensis (Karaman 1928) -- (Vasil'eva 1998:95 [ref. 23591], Economidis 2005:67 [ref. 28213]). •Synonym of Barbatula barbatula (linnaeus 1758) -- (Kottelat 2012:78 [ref. 32367]). •Valid as Barbatula vardarensis (Karaman 1928) -- (Geiger et al. 2014:Table S2 [ref. 37408]). Current status: Valid as Barbatula vardarensis (Karaman 1928). Nemacheilidae. Distribution: Vardar / Axios River basin, Macedonia and Greece. Habitat: freshwater.