Species that contain: Noemacheilus and toni [ 2 ] records markakulensis, Nemacheilus barbatulus Menshikov [M. I.] 1939:141, Fig. [Ucenye zapiski / Permskij Ordena Trudovogo Krasnogo Znameni Gosudarstvennyj Universitet Imeni A. M. Gor'kogo, Ministerstvo Vyssego i Srednego Special'nogo Obrazovanija RSFSR. v. 3 (no. 2); ref. 21170] Lake Marka-kul', Irtysh basin, Kazakhstan. Syntypes: ZIN 26864 (6). Author also seen as Men'shikov. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:872 [ref. 20661] as Nemachilus). •Synonym of Noemacheilus barbatulus (Linnaeus 1758), but a valid subspecies markakulensis Manshikov 1939 -- (Mitrofanov & Dukravec 1989:60 [ref. 37748]). •Synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (authors). •Valid as Barbatula markakulensis (Menshikov 1939) -- (Kottelat 2012:78 [ref. 32367] with question, Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:396 [ref. 34693] as makakulensis, Prokofiev 2018:273 [ref. 35878]). Current status: Valid as Barbatula markakulensis (Menshikov 1939). Nemacheilidae. Distribution: North Asia: Markakul Lake (Kazakhstan). Habitat: freshwater.
toni, Cobitis Dybowski [B. N.] 1869:957, Pl. 18 (fig. 10) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 19; ref. 1169] Onon and Ingoda rivers, Amur River basin, Russia. Syntypes and/or Dybowski specimens: MNHN B-0738 (8), B-0739 (4); NMW 50473 (2). Type catalog: Bertin & Estève 1948:98 [ref. 19574]. For remarks on systematics see Kottelat 2006:53 [ref. 28831]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies -- (Berg 1949:869 [ref. 20661] as Nemachilus, Kawanabe & Mizuno 1989:398 [ref. 25598] as Neomacheilus, Watanabe 1998:263 [ref. 23491]). •Synonym of Noemacheilus barbatulus (Linnaeus 1758), but a valid subspecies toni -- (Chereshnev 1996:601 [ref. 24659], Mitrofanov & Dukravec 1989:58 [ref. 37748], Nakabo 2000:276 [ref. 25086], Nakabo 2002:276 [ref. 26001]). •Synonym of Barbatula barbatula, subspecies nuda (Bleeker 1865) -- (Zhu 1989:29 [ref. 17744]). •Valid as Nemacheilus toni (Dybowski 1869) -- (Wang 1984:82 [ref. 22951], Sawada in Masuda et al. 1984:59 [ref. 6441] as Noemacheilus, Bogutskaya & Naseka 1996:47 [ref. 22798], Matsuura et al. 2000:176 [ref. 26000], Choi et al. 2003:178, 517 [ref. 26218]). •Valid as Orthrias toni (Dybowski 1869) -- (Bănărescu & Nalbant 1995:449 [ref. 23187], Kim et al. 2000:224 [ref. 25434], Kim & Park 2002:193 [ref. 25913], Jang et al. 2003:119 [ref. 27681]). •Synonym of Orthrias nudus (Bleeker 1865) -- (Kim 1997:555 [ref. 24032]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies toni (Dybowski 1869) -- (Prokofiev 2003:736 [ref. 27494], Prokofiev 2007:17 [ref. 30343]). •Valid as Barbatula toni (Dybowski 1869) -- (Okada 1961:541 [ref. 6363], Nikiforov et al. 1997:330 [ref. 24694], Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:95 [ref. 23591], Sheiko & Fedorov 2000:17 [ref. 25650], Chereshnev et al. 2001:37 [ref. 26736], Tokranov 2001:268 [ref. 25407], Zemnukhov et al. 2001:420 [ref. 25412], Kottelat 2001:49 [ref. 25482], Pietsch et al. 2001:140 [ref. 25688], Bogutskaya et al. 2001:45 [ref. 26178], Sideleva 2001:54 [ref. 26181], Fedorov et al. 2003:25 [ref. 27579], Alekseyev et al. 2004:54 [ref. 27625], Sideleva 2003:11 [ref. 27815], Bogutskaya & Naseka 2004:113 [ref. 28183], Kottelat 2006:53 [ref. 28831], Ocock et al. 2006:28 [ref. 28933], Kottelat & Freyhof 2007:330 [ref. 29996], Bogutskaya et al. 2008:339 [ref. 30085], Cao et al. 2012:241 [ref. 32335], Kottelat 2012:80 [ref. 32367], Parin et al. 2014:103 [ref. 33547], Prokofiev 2015:460 [ref. 34065], Prokofiev 2016:399 [ref. 34693], Dyldin & Orlov 2016:669 [ref. 34853] with question, Prokofiev 2018:274 [ref. 35878], Chen et al. 2019:8 [ref. 36569], Dyldin et al. 2020:87 [ref. 37569], An et al. 2021:121 [ref. 38850], Dyldin et al. 2021:19 [ref. 38804] with question, Dyldin & Orlov 2021:63 [ref. 38158] with question, Dyldin et al. 2023:[8] [ref. 40328] with question). Current status: Valid as Barbatula toni (Dybowski 1869). Nemacheilidae. Distribution: East Asia: Amur (Heilongjiang) and Yin River drainages (Russia, Mongolia, northern China and northern Korea). Habitat: freshwater.
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