Species that contain: Pungitius and platygaster [ 7 ] records

aralensis, Gasterosteus platygaster var. Kessler [K. F.] 1877:4 [The Aralo-Caspian Expedition; ref. 2597] Aral Sea and mouth of Amu Darya River, Uzbekistan. Syntypes: ZIN 3712 (3). •Synonym of Pungitius platygaster (Kessler 1859) -- (Monod 1973:286 [ref. 7193], Keivany & Nelson 2000:117 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856]). •Synonym of Pungitius platygaster (Kessler 1859), but a valid subspecies aralensis Kessler 1877 -- (Berg 1949:970 [ref. 20662], Mitrofanov & Dukravec 1989:114 [ref. 37748], Reshetnikov et al. 1997:747 [ref. 24702], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Jouladeh-Roudbar et al. 2020:286 [ref. 37983], Çiçek et al. 2024:376 [ref. 40789]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish.

caucasicus, Gasterosteus platygaster var. Kessler [K. F.] 1877:3 [The Aralo-Caspian Expedition; ref. 2597] Transcaucasia. Syntypes: ZIN 2401 (12+). •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Kottelat 1997:166 [ref. 22952], Keivany & Nelson 2000:116 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983], Çiçek et al. 2024:376 [ref. 40789]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

danubica, Gasterosteus platygaster var. Steindachner [F.] 1899:542 [4] [Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften v. 108 (pt. 1) (no. 7) (for July 1899); ref. 10017] Danube swamps near Negotin, Srebia; Save [Sava] River at Belgrad, Serbia. Syntypes: NMW 6823 (6). On p. 4 of separate. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Kottelat 1997:166 [ref. 22952], Keivany & Nelson 2000:117 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

kessleri, Gasterosteus pungitius var. Yakovlev [V. E.] 1870:110 [10] [On the new and little-known species of fish found in the mouths of the Volga; ref. 19610] Lakes near Astrakhan, Volga River Delta, Russia. Syntypes: whereabouts unknown. On p. 10 of separate. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Coad 1995:27 [ref. 23608], Kottelat 1997:166 [ref. 22952], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Keivany & Nelson 2000:116 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

niger, Gasterosteus pungitius var. Yakovlev [V. E.] 1870:110 [10] [On the new and little-known species of fish found in the mouths of the Volga; ref. 19610] Lakes near Astrakhan, Volga River Delta, Russia. Syntypes: whereabouts unknown. On p. 10 of separate. Objectively invalid; preoccupied by Gasterosteus niger Cuvier 1829. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Coad 1995:27 [ref. 23608], Kottelat 1997:166 [ref. 22952], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Keivany & Nelson 2000:116 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). 1°Homonym. Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

nuda, Pygosteus platygaster Berg [L. S.] (ex Nikolskii) 1905:218 [Izviestii Turkestanskago otdiela Russkago geograficheskago obschchestva v. 4; ref. 15338] Lake Charkal, Ural River basin, Russia. Syntypes: ZIN 10613. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Kottelat 1997:167 [ref. 22952], Keivany & Nelson 2000:117 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

platygaster, Gasterosteus Kessler [K. F.] 1859:202 [Bulletin de la Société Impériale des Naturalistes de Moscou v. 32 (no. 3); ref. 18391] Odessa and side arm of Dnieper River in Aleschki, Ukraine. Syntypes: BMNH 1897.7.5.2 [ex ZIN] (2), ZIN 2350-51 (6, 6+). Additional material: (14) ZIN 2350-51 (6, 6+) Dnieper River near Aleschki, Ukraine. Also appeared in Kessler 1860:17, Pl. 1 (fig. 1) [ref. 18393]. •May be a subspecies of Pungitius pungitius (Linnaeus 1758) -- (Coad 1995:27 [ref. 23608]). •Valid as Pungitius platygaster (Kessler 1859), subspecies platygaster Kessler 1859 -- (Mitrofanov & Dukravec 1989:112 [ref. 37748]). •Valid as Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Coad 1981:16 [ref. 22348], Banister in Whitehead et al. 1986:642 [ref. 13676], Coad 1995:26 [ref. 23608], Economidis 1995:208 [ref. 23025], Keivany et al. 1997:558 [ref. 22904], Kottelat 1997:166 [ref. 22952], Reshetnikov et al. 1997:747 [ref. 24702], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Coad 1998:103 [ref. 23869], Keivany & Nelson 1998:458 [ref. 24681], Keivany & Nelson 2000:116 [ref. 24720], McLennan & Mattern 2001:11 [ref. 25222], Bogutskaya et al. 2001:46 [ref. 26178], Bilecenoğlu et al. 2002:64 [ref. 26753], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Bogutskaya & Naseka 2004:181 [ref. 28183], Vassilev & Pehlivanov 2005:175 [ref. 28369], Fricke et al. 2007:73 [ref. 29533], Kottelat & Freyhof 2007:498 [ref. 29996], Vasil'eva 2007:71 [ref. 30517], Esmaeili et al. 2010:375 [ref. 33391], Parin et al. 2014:205 [ref. 33547] as subspecies platycephalus, Jouladeh-Roudbar et al. 2015:895 [ref. 34062], Barbieri et al. 2015:97 [ref. 35029], Çiçek et al. 2015:151 [ref. 35151], Takahashi et al. 2016:49 [ref. 34337], Esmaeili et al. 2017:95 [ref. 35254], Esmaeili et al. 2018:77 [ref. 36089], Artüz & Fricke 2019:555 [ref. 36612], Jouladeh-Roudbar et al. 2020:286 [ref. 37983], Kuljanishvili et al. 2020:[5] [ref. 37535], Matsumoto et al. 2021:9 [ref. 38508], Wang et al. 2021:[3] [ref. 38800], Çiçek et al. 2023:55 [ref. 40326], Çiçek et al. 2024:376 [ref. 40789]). Current status: Valid as Pungitius platygaster (Kessler 1859). Gasterosteidae. Distribution: Eurasia: Black, Caspian and Aral Sea basins. Habitat: freshwater, brackish.