Species that contain: Pygosteus and pungitius [ 4 ] records

kaibarae, Pygosteus Tanaka [S.] 1915:565 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 27 (no. 325); ref. 4324] Kichisho-in, southwest of Kyoto, Japan. Lectotype: ZUMT 8197. Paralectotypes: ZUMT 59848-54 (7). Lectotype established by Tanaka 1918:491 [ref. 14911], as ZUMT 8197 (identity of specimen not researched). •Synonym of Pungitius sinensis (Guichenot 1869), but a valid subspecies -- (Ida in Masuda et al. 1984:84 [ref. 6441]). •See Kim 1997:392 [ref. 24032]. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Okada 1961:622 [ref. 6363], Keivany & Nelson 2000:118 [ref. 24720]). •Valid as Pungitius kaibarae (Tanaka 1915) -- (Nakabo 2000:514 [ref. 25086], Kim & Park 2002:314 [ref. 25913], Nakabo 2002:514 [ref. 26001], Choi et al. 2003:231, 553 [ref. 26218], Jang et al. 2003:120 [ref. 27681], Bogutskaya et al. 2008:349 [ref. 30085], Parin et al. 2014:204 [ref. 33547], Takahashi et al. 2016:49 [ref. 34337], Matsumoto et al. 2021:16 [ref. 38508], Wang et al. 2021:[3] [ref. 38800]). Current status: Valid as Pungitius kaibarae (Tanaka 1915). Gasterosteidae. Distribution: Northwestern Pacific: Sea of Japan (Hyogo, Kyoto and Osaka prefectures, Japan and eastern side of Korean Peninsulae). IUCN (2021): Not assessed. Habitat: freshwater, brackish.

nuda, Pygosteus platygaster Berg [L. S.] (ex Nikolskii) 1905:218 [Izviestii Turkestanskago otdiela Russkago geograficheskago obschchestva v. 4; ref. 15338] Lake Charkal, Ural River basin, Russia. Syntypes: ZIN 10613. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Kottelat 1997:167 [ref. 22952], Keivany & Nelson 2000:117 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

steindachneri, Pygosteus Jordan [D. S.] & Snyder [J. O.] 1901:747 [Proceedings of the United States National Museum v. 23 (no. 1235); ref. 2504] Replacement name for Gasterosteus japonicus Steindachner 1881, preoccupied by Gasterosteus japonicus Houttuyn 1782. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Okada 1961:621 [ref. 6363], Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius sinensis (Guichenot 1869) -- (Bogutskaya et al. 2008:350 [ref. 30085]). Replacement Name. Current status: Synonym of Pungitius sinensis (Guichenot 1869). Gasterosteidae. Habitat: freshwater, brackish, marine.

undecimalis, Pygosteus Jordan [D. S.] & Starks [E. C.] 1902:62, Fig. 1 [Proceedings of the United States National Museum v. 26 (no. 1308); ref. 2525] Chitose, Hokkaido, Japan. Holotype: CAS-SU 7119. Paratypes: CAS-SU 7153 (5). Type catalog: Böhlke 1953:59 [ref. 12291]. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies tymensis (Nikolskii 1889) -- (Okada 1961:619 [ref. 6363], Monod 1973:285 [ref. 7193], Keivany & Nelson 2000:119 [ref. 24720]). •Synonym of Pungitius tymensis (Nikolskii 1889) -- (Berg 1949:968 [ref. 20662], Dyldin & Orlov 2017:58 [ref. 35138]). Current status: Synonym of Pungitius tymensis (Nikolskii 1889). Gasterosteidae. Habitat: freshwater.