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Species that contain: Salmo and trutta [ 81 ] records abanticus, Salmo trutta Tortonese [E.] 1954:19, Pl. 1 (fig. 3); Figs. 2, 3a-b [Publications Institute Hydrobiology University of Istanbul (Sér. B) v. 2 (no. 1); ref. 12668] Lake Abant, northern Anatolia, Asiatic Turkey. Holotype (unique): MSNM 1 [ex MSNM 5031]. •Listed -- (Kottelat 1997:144 [ref. 22952]). •Synonym of Salmo labrax Pallas 1814 -- (Fricke et al. 2007:55 [ref. 29533]). •Synonym of Salmo trutta Linnaeus 1758 -- (Kalayci et al. 2018:803 [ref. 36785]). •Valid as Salmo abanticus Tortonese 1954 -- (Turan et al. 2010:362 [ref. 30689], Turan et al. 2011:32 [ref. 31466], Turan et al. 2012:234 [ref. 32276], Turan et al. 2014:285 [ref. 33242], Turan et al. 2014:149 [ref. 33615], Çiçek et al. 2015:150 [ref. 35151], Turan et al. 2017:62 [ref. 35485], Çiçek et al. 2018:21 [ref. 36267], Çiçek et al. 2020:255 [ref. 37644], Schöffmann 2021:191 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Çiçek et al. 2023:421 [ref. 40625], Turan et al. 2024:88 [ref. 40793], Çiçek et al. 2024:285 [ref. 40789], Küçük et al. 2024:400 [ref. 40993]). Current status: Valid as Salmo abanticus Tortonese 1954. Salmonidae: Salmoninae. Distribution: Asia Minor: Lake Abant, Bolu Province, northwestern Anatolia (Turkey); introduced in Black Sea watersheds. Habitat: freshwater, brackish, marine. adriatica, Trutta Kolombatović [G.] 1890:166 [4] [Glasnik Hrvatskoga Naravoslovnoga Društva v. 5; ref. 20232] Vranjic, Croatia, Adriatic Sea, Mediterranean Sea. Holotype (unique): whereabouts unknown. On p. 4 of separate. •Questionably a synonym of Salmo marmoratus Cuvier 1829 -- (Kottelat 1997:138 [ref. 22952], Bianco 2013:3 [ref. 32897]). Current status: Uncertain as Salmo marmoratus Cuvier 1829. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. albus, Salmo Bonnaterre [J. P.] (ex Pennant) 1788:161 [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] Rivers and seas of Great Britain. No types known. Based on the 'Gwiniad' of Pennant 1769:267, Pl. 16 [ref. 18527]. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). •See Dorofeeva & Savvaitova 1998:40 [ref. 23574]. Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. albus, Salmo Walbaum [J. J.] 1792:76 [Petri Artedi sueci genera piscium Part 3; ref. 4572] Rivers and seas of Great Britain; Carolina, U.S.A. No types known. Based in part on the 'Gwiniad' of Pennant 1769:267, Pl. 16 [ref. 18527], and on Pennant 1785:Suppl. 140 [ref. 18530]. Objectively invalid; preoccupied by Salmo albus Bonnaterre 1788. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). 1°Homonym. Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. albus, Salmo salvelinus var. Fitzinger [L. J. F. J.] 1832:338 [Beiträge zur Landeskunde Österreichs unter der Enns, Wien v. 1; ref. 5019] No types known. Not available, name only; from Leytha, Fischa, Traisen (Danube system), Austria. •Synonym of Salmo trutta Linnaeus 1758. Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. aralensis, Salmo trutta Berg [L. S.] 1908:317 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 13 (no. 2); ref. 14303] Aral Sea near Amu-Darya River mouth, Kazakhstan. Holotype (unique): ZIN 14418. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies as described -- (Berg 1948:249 [ref. 12882], see Coad 1981:8 [ref. 22348]). •Listed -- (Kottelat 1997:144 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. argenteus, Fario Valenciennes [A.] in Cuvier & Valenciennes 1848:294, Pl. 616 [Histoire naturelle des poissons v. 21; ref. 1013] Many localities. Syntypes: MNHN. Kottelat 1997:142 [ref. 22952] "restricted" the type locality to the Atlantic drainage of France, but the Code does not provide for such restriction in that manner. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. argentinus, Salmo Linnaeus [C.] (ex Koelreuter) 1766:511 [Systema naturae sive regna tria naturae v. 1 (pt 1); ref. 2786] Brazil. No types known. Unneeded replacement name for Trutta dentata Koelreuter 1763. •Synonym of Piabucus dentatus (Koelreuter 1763) -- (Moreira in Reis et al. 2003:173 [ref. 27061], Moreira 2024:377 [ref. 40817]). Replacement Name. Current status: Synonym of Piabucus dentatus (Koelreuter 1763). Iguanodectidae. Habitat: freshwater. ausonii, Salar Valenciennes [A.] in Cuvier & Valenciennes 1848:319, Pl. 618 [Histoire naturelle des poissons v. 21; ref. 1013] Localities in France, Germany, and Italy. Syntypes: MNHN 0002 (1), 3647 (1), A-7590 (2), 1953 (2), 3638 to 3643 (8), 3650 and 3651 (2), 3648 (1), 3637 (1), 3652 to 3654 (3), A-8567-68 (1, 1), 3644 to 3646 (3), A-8573 (1), 3649 (1), 3634 (1), A-9702 (1), 3635 and 3636 and 4920 (3). Type catalog: Bertin 1940:303-304 [ref. 293]. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. bailloni, Salar Valenciennes [A.] in Cuvier & Valenciennes 1848:342, Pl. 619 [Histoire naturelle des poissons v. 21; ref. 1013] Somme River at Abbeville, France. Lectotype: MNHN 0000-6321. Paralectotypes: MNHN 0000-2909 (2) Norway. Type catalog: Bertin 1940:305 [ref. 293] with MNHN 6321 as holotype and 2909 as paratypes. Lectotype designated by Eschmeyer 1998:186 [ref. 23416]. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. balcanica, Trutta Karaman [S.] 1927:253, Fig. 1 [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 2 (nos 1-2); ref. 15859] Ohrid Lake, southern Yugoslavia/eastern Albania. No types known. •Valid as Salmo balcanicus (Karaman 1927) -- (Kottelat 1997:133 [ref. 22952], Kottelat & Freyhof 2007:420 [ref. 29996], Shumka et al. 2023:39 [ref. 40747]). Current status: Valid as Salmo balcanicus (Karaman 1927). Salmonidae: Salmoninae. Distribution: Southeastern Europe: Lake Ohrid (Albania and Macedonia). Habitat: freshwater. brachypoma, Salmo Günther [A.] 1866:87, Figs. [Catalogue of the fishes in the British Museum v. 6; ref. 1983] Firth of Forth, Tweed and Ouse rivers, England. Syntypes: (7) BMNH 1839.1.4.23 (1, stuffed) Firth of Forth, 1848.1.12.239 (1, LHS skin) Firth of Forth, 1865.9.25.1-2 (2) River Ouse. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. caecifer, Salmo Parnell [R.] 1838 [Memoirs of the Wernerian Natural History Society, Edinburgh v. 7 (art. 2) (for 1831-1837); ref. 18519] Loch Leven, Scotland. Syntypes: whereabouts unknown. Original not seen. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. cambricus, Salmo Donovan [E.] 1806:3 unnum. pp., Pl. 91 [The natural history of British fishes, including scientific and general descriptions of the most interesting species, and an extensive selection of accurately finished coloured plates.; ref. 17408] Glamorganshire and Carmarthenshire, Wales, U.K. No types known. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. caspius, Salmo Kessler [K. F.] 1877:62, Pl. 2 (fig. 15) [The Aralo-Caspian Expedition; ref. 2597] Kura River near Bozhii Promysel fishing grounds, Azerbaijan. Syntypes: (3) not at ZIN. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies -- (Berg 1948:242 [ref. 12882], Reshetnikov et al. 1997:729 [ref. 24702], Dorofeeva & Savvaitova 1998:36 [ref. 23574]). •Listed -- (Kottelat 1997:143 [ref. 22952]). •Synonym of Salmo trutta Linnaeus 1758 -- (Kalayci et al. 2018:803 [ref. 36785]). •Valid as Salmo caspius Kessler 1877 -- (Fricke et al. 2007:55 [ref. 29533], Ninua & Japoshvili 2008:168 [ref. 31251], Esmaeili et al. 2010:374 [ref. 33391], Turan et al. 2010:362 [ref. 30689], Turan et al. 2011:32 [ref. 31466], Turan et al. 2012:234 [ref. 32276], Turan et al. 2014:285 [ref. 33242], Turan et al. 2014:149 [ref. 33615], Jouladeh-Roudbar et al. 2015:891 [ref. 34062], Çiçek et al. 2015:150 [ref. 35151], Esmaeili et al. 2017:76 [ref. 35254], Turan et al. 2017:62 [ref. 35485], Esmaeili et al. 2018:57 [ref. 36089], Ninua et al. 2018:[12] [ref. 35786], Çiçek et al. 2020:255 [ref. 37644], Dyldin et al. 2020:83 [ref. 37569], Jouladeh-Roudbar et al. 2020:220 [ref. 37983], Kaya et al. 2020:340 [ref. 37582], Kuljanishvili et al. 2020:[6] [ref. 37535], Turan et al. 2020:1461 [ref. 37608], Schöffmann 2021:208 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Pozdeev et al. 2022:6 [ref. 39511], Turan et al. 2022:56 [ref. 39307], Çiçek et al. 2023:422 [ref. 40625], Çiçek et al. 2024:287 [ref. 40789]). Current status: Valid as Salmo caspius Kessler 1877. Salmonidae: Salmoninae. Distribution: Eurasia: Kura River drainage and southern and southwestern Caspian Sea watersheds. Habitat: freshwater, brackish, marine. ciscaucasicus, Salmo trutta Dorofeeva [E. A.] 1967:15 [Voprosy Ikhtiologii v. 7 (no. 1, art. 42); ref. 7957] Keyranchay River, Samur delta, eastern Ciscaucasia, Russia. Holotype: ZIN 26244. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies as described -- (Reshetnikov et al. 1997:729 [ref. 24702], Dorofeeva & Savvaitova 1998:36 [ref. 23574]). •Listed -- (Kottelat 1997:144 [ref. 22952]). •Valid as Salmo ciscaucasicus Dorofeeva 1967 -- (Kottelat & Freyhof 2007:427 [ref. 29996], Ninua et al. 2018:[12] [ref. 35786], Dyldin et al. 2020:84, 103 [ref. 37569], Kaya et al. 2020:340 [ref. 37582] as aff. ciscaucasicus, Kuljanishvili et al. 2020:[6] [ref. 37535], Schöffmann 2021:213 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Turan et al. 2022:57 [ref. 39307], Çiçek et al. 2023:423 [ref. 40625], Çiçek et al. 2024:287 [ref. 40789]). Current status: Valid as Salmo ciscaucasicus Dorofeeva 1967. Salmonidae: Salmoninae. Distribution: Eurasia: Terek River and rivers flowing into the Caspian Sea from the north and northwest. Habitat: freshwater, brackish, marine. cornubiensis, Salmo Walbaum [J. J.] 1792:65 [Petri Artedi sueci genera piscium Part 3; ref. 4572] Lake Lo, Cornwall, England. No types known. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. cumberland, Salmo Lacepède [B. G. E.] 1803:695, 696 [Histoire naturelle des poissons (Lacepède) v. 5; ref. 4930] Lakes of Cumbreland and Scotland, U.K. No types known. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. dentex, Salar Heckel [J. J.] 1851:371, Pl. 10 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 8; ref. 18259] Narenta [Nerteva River], Bosnia-Herzegovina. Lectotype: NMW 65895. Paralectotypes: NMW 16505 (1, dry), 65887 (9), 65888 (4, poor condition), 65895 (4), 92806 (1, dry), 95284 (1), 95285 (1); NRM 25000 (1). See Kottelat 1997:135 [ref. 22952] for localities. Lectotype selected by Delling 2011:339 [ref. 31191]. •Questionably valid as Salmo dentex (Heckel 1851) -- (Kottelat 1997:135 [ref. 22952], Delling 2002:299 [ref. 26660]). •Synonym of Salmo trutta Linnaeus 1758, subspecies dentex (Heckel 1851) -- (Ćaleta et al. 2015:85 [ref. 34877]). •Valid as Salmo dentex (Heckel 1851) -- (Mrakovcic et al. 1995:183 [ref. 23079], Zupančič 2008:62 [ref. 29926], Kottelat & Freyhof 2007:421 [ref. 29996], Snoj et al. 2010:[1] 199 [ref. 41208], Delling 2011:339 [ref. 31191], Bianco 2013:3 [ref. 32897], Ćaleta et al. 2019:177 [ref. 37026] with question). Current status: Valid as Salmo dentex (Heckel 1851). Salmonidae: Salmoninae. Distribution: Southeastern Europe: Adriatic basin. Habitat: freshwater. eriox, Salmo Linnaeus [C.] 1758:308 [Systema Naturae, Ed. X v. 1; ref. 2787] European seas and rivers. No types known. Neotype designated by Fricke 1999:29 [ref. 24101], but withdrawn in Fricke 2000:639 [ref. 24537]. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952], Scott & Crossman 1998:200 [ref. 24570], Fricke 1999:29 [ref. 24101]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. estuarius, Salmo Knox [R.] 1855:4662 [The Zoologist: a Popular Miscellany of Natural History v. 13; ref. 18428] England and Scotland. Syntypes: whereabouts unknown. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. ezenami, Salmo trutta Berg [L. S.] 1948:253 [267] [Freshwater fishes of the U.S.S.R. and adjacent countries No. 27; ref. 12882] Lake Ezenam, Daghestan, Caucasus, Russia, elevation 1882 meters. Syntypes: ZIN 28356 (11). On p. 267 of English translation. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies as described -- (Reshetnikov et al. 1997:729 [ref. 24702], Dorofeeva & Savvaitova 1998:36 [ref. 23574]). •Listed -- (Kottelat 1997:144 [ref. 22952]). •Valid as Salmo ezenami Berg 1948 -- (Bogutskaya et al. 2001:46 [ref. 26178], Bogutskaya & Naseka 2004:156 [ref. 28183], Kottelat & Freyhof 2007:428 [ref. 29996], Dyldin et al. 2020:88, 103, 113 [ref. 37569]). Current status: Valid as Salmo ezenami Berg 1948. Salmonidae: Salmoninae. Distribution: Eurasia: northern Caucasus region. Habitat: freshwater, brackish, marine. fario, Salmo Linnaeus [C.] 1758:309 [Systema Naturae, Ed. X v. 1; ref. 2787] Rivers of Sweden and Switzerland. No types known. •Valid as Salmo fario Linnaeus 1758 -- (Okada 1961:724 [ref. 6363]). •Synonym of Salmo trutta Linnaeus 1758, subspecies trutta but as a valid morpha -- (Lelek 1987:105 [ref. 22962], Hanel 2003:58 [ref. 27894]). •Synonym of Salmo trutta Linnaeus 1758 -- (Berg 1948:255 [ref. 12882], Cadwallader & Backhouse 1983:215 [ref. 25473], Smith 1986:231 [ref. 22186], Baxter & Stone 1995:191 [ref. 23374], La Rivers 1994:318 [ref. 23389], Kottelat 1997:141 [ref. 22952], Scott & Crossman 1998:200 [ref. 24570], Kalayci et al. 2018:803 [ref. 36785]). •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies fario Linnaeus 1758 -- (Svetovidov in Whitehead et al. 1984:383 [ref. 13675], Pequeño 1989:29 [ref. 14125], Talwar & Jhingran 1991:721 [ref. 20764], Wu & Wu 1992:128 [ref. 21205], Zhang et al. 1995:43 [ref. 22734], Zhu 1995:10 [ref. 25213], Dorofeeva 1998:442 [ref. 24680] as a valid morpha, Rafique 2000:323 [ref. 25220], Mirza 2003:3 [ref. 27265] as a valid subspecies, Gabrielyan 2001:24 [ref. 25867], Vassilev & Pehlivanov 2005:164 [ref. 28369], Shrestha 2008:202 [ref. 29923], Gurung et al. 2013:4884 [ref. 32969], Zhang et al. 2016:212 [ref. 34477]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. farioides, Salmo Karaman [S.] 1938:137, Fig. 3 [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 18; ref. 17197] Krka River in Croatia, near the town of Knin. Neotype: MCCI/P/501. Syntypes: whereabouts unknown. See Kottelat 1997:135 [ref. 22952] for localities. Neotype selected by Bianco 2014:65 [ref. 32897]. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies -- (Mrakovcic et al. 1995:183 [ref. 23079]). •Questionably valid as Salmo farioides Karaman 1938 -- (Kottelat 1997:135 [ref. 22952]). •Valid as Salmo farioides Karaman 1928 -- (Kottelat & Freyhof 2007:421 [ref. 29996], Zupančič 2008:63 [ref. 29926], Delling 2011:337 [ref. 31191], Bianco 2013:4 [ref. 32897], Barbieri et al. 2015:82 [ref. 35029], Ćaleta et al. 2019:177 [ref. 37026], Schöffmann 2021:142 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Shumka et al. 2023:39 [ref. 40747]). Current status: Valid as Salmo farioides Karaman 1938. Salmonidae: Salmoninae. Distribution: Southeastern Europe: eastern Adriatic slope (Croatia, Bosnia-Herzegovina, Montenegro, Albania, Macedonia and Greece). Habitat: freshwater. fluvialis, Trutta Dralet [É.-F.] 1821:18 [Considérations sur l'histoire naturelle des poissons, sur la pêche et les lois qui la régissent.; ref. 17418] No locality. No types known. Based on Rondelet. •Synonym of Salmo trutta Linnaeus 1758. Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater. forestensis, Salmo fario var. Bloch [M. E.] & Schneider [J. G.] 1801:400 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Europe (several localities); Siberia, Russia. No types known. Species epithet corrected from faris in corrigenda. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. gadoides, Salmo Lacepède [B. G. E.] 1803:157, 224 [Histoire naturelle des poissons (Lacepède) v. 5; ref. 4930] Trouville pond, near Rouen, France. No types known. Originally gadoïdes. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. gaimardi, Salar Valenciennes [A.] in Cuvier & Valenciennes 1848:341 [Histoire naturelle des poissons v. 21; ref. 1013] Iceland. Syntypes: MNHN 0000-3657 to 0000-3661 (5, poor condition). Type catalog: Bertin 1940:304 [ref. 293]. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. gallivensis, Salmo Günther [A.] 1866:88, Fig. [Catalogue of the fishes in the British Museum v. 6; ref. 1983] Galway and Ballinahuich Fishery, Ireland. Syntypes: (at least 4, but not the hybrids) BMNH 1865.8.30.1-2 (2), 1865.8.30.3 (1), 1865.8.30.8 (1)., ?1865.8.30.4-7 (4). •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. illanca, Salmo Wartmann [B.] 1783:55 [Schriften der Berlinischen Gesellschaft Naturforschender Freunde v. 4; ref. 17831] Germany. No types known. •Synonym of Salmo lacustris Linnaeus 1758 -- (Kottelat 1997:143 [ref. 22952]). •Synonym of Salmo trutta Linnaeus 1758. Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater. islayensis, Salmo Thomson [J.] 1873:86 [Hardwicke's Science -- Gossip: an illustrated medium of interchange and gossip. London v. 8 (no. 83) (for 1872); ref. 19841] Loch Na-Maorichen, Islay Island, U. K. No types known. Syntypes: whereabouts unknown. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. krkensis, Trutta obtusirostris Karaman [S.] 1927:259, Fig. 4 [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 2 (nos 1-2); ref. 15859] Krka River at Knin, Croatia. Syntypes: whereabouts unknown. •Synonym of Salmothymus obtusirostris (Heckel 1851), but a valid subspecies -- (Mrakovcic et al. 1995:184 [ref. 23079], Schöffmann 2021:154 [ref. 38997]). •Synonym of Salmothymus obtusirostris (Heckel 1851) -- (Kottelat 1997:144 [ref. 22952]). •Synonym of Salmo obtusirostris (Heckel 1851) but a doubtful subspecies -- (Snoj et al. 2002:399 [ref. 26461]). •Valid as Salmo krkensis (Karaman 1927) -- (Zupančič 2008:64 [ref. 29926]). Current status: Synonym of Salmo obtusirostris (Heckel 1851). Salmonidae: Salmoninae. Habitat: freshwater. labrax, Salmo Pallas [P. S.] 1814:346 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Sivastopol and Biyuk-ozen River; Chersones; and Ochakov, Crimea, Ukraine. Syntypes: whereabouts unknown. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies -- (Berg 1948:239 [ref. 12882], Svetovidov 1973:145 [ref. 7169], Svetovidov in Whitehead et al. 1984:383 [ref. 13675], Lelek 1987:109 [ref. 22962], Reshetnikov et al. 1997:729 [ref. 24702], Dorofeeva & Savvaitova 1998:36 [ref. 23574], Dorofeeva 1998:442 [ref. 24680], Vassilev & Pehlivanov 2005:165 [ref. 28369], Parin et al. 2014:138 [ref. 33547]). •Synonym of Salmo trutta Linnaeus 1758 -- (Kalayci et al. 2018:803 [ref. 36785]). •Valid as Salmo labrax Pallas 1814 -- (Kottelat 1997:136 [ref. 22952], Bilecenoğlu et al. 2002:40 [ref. 26753], Delling 2002:299 [ref. 26660], Fricke et al. 2007:55 [ref. 29533], Kottelat & Freyhof 2007:429 [ref. 29996], Ninua & Japoshvili 2008:168 [ref. 31251], Turan et al. 2010:362 [ref. 30689], Turan et al. 2014:149 [ref. 33615], Çiçek et al. 2015:150 [ref. 35151], Artüz & Fricke 2019:553 [ref. 36612], Ćaleta et al. 2019:178 [ref. 37026], Özuluğ & Saç 2019:32 [ref. 36599] as cf. labrax, Çiçek et al. 2020:255 [ref. 37644], Dyldin et al. 2020:84, 103 [ref. 37569], Kuljanishvili et al. 2020:[6] [ref. 37535], Abliazov et al. 2021:8 [ref. 38576], Schöffmann 2021:173 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Niță et al. 2022:129 [ref. 40471], Turan et al. 2022:62 [ref. 39307], Çiçek et al. 2023:425 [ref. 40625], Shumka et al. 2023:39 [ref. 40747], Çiçek et al. 2024:290 [ref. 40789]). Current status: Valid as Salmo labrax Pallas 1814. Salmonidae: Salmoninae. Distribution: Eurasia: Black Sea and adjacent watersheds; introduced in Tigris-Euphrates watersheds. Habitat: freshwater, brackish, marine. lacustris, Salmo Linnaeus [C.] 1758:309 [Systema Naturae, Ed. X v. 1; ref. 2787] Switzerland (several localities). No types known. Kottelat 1997:143 [ref. 22952] "restricted" the type locality to Lake Constance, Switzerland, but the Code does not provide for such restriction in that manner. •Synonym of Salmo trutta Linnaeus 1758 -- (Berg 1948:250 [ref. 12882]). •Synonym of Salmo trutta Linnaeus 1758 or valid as Salmo lacustris Linnaeus 1758 -- (Kottelat 1997:143 [ref. 22952]). •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies -- (Svetovidov in Whitehead et al. 1984:382 [ref. 13675], Lelek 1987:112 [ref. 22962]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. laevis, Salmo trutta var. Walbaum [J. J.] 1792:60 [Petri Artedi sueci genera piscium Part 3; ref. 4572] No locality stated. No types known. Unneeded replacement name or unjustified emendation of Salmo levis Mohr 1786. •Synonym of Salvelinus alpinus (Linnaeus 1758) -- (Kottelat 1997:152 [ref. 22952]). Current status: Synonym of Salvelinus alpinus (Linnaeus 1758). Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. lapasseti, Salmo Zill [M.] 1858:127 [Annales des Sciences Naturelles, Paris (Zoologie) (Sér. 4) v. 9; ref. 17709] El-abaïch, tributary of Oued-Z'hour, entering the Mediterranean about 30 kilometers west of Collo, Algeria. Syntypes: (several) whereabouts unknown. •Synonym of Salmo trutta Linnaeus 1758 -- (Behnke 1984:125 [ref. 6177]). •Questionably a synonym of Salmo cettii Rafinesque 1810 -- (Kottelat 1997:135 [ref. 22952]). •Synonym of Salmo macrostigma (Duméril 1858) -- (Delling & Doadrio 2005:50 [ref. 28211], Doadrio et al. 2015:2 [ref. 33950]). Current status: Synonym of Salmo macrostigma (Duméril 1858). Salmonidae: Salmoninae. Habitat: freshwater. lemanus, Salmo Cuvier [G.] 1829:303 [Le Règne Animal (Edition 2) v. 2; ref. 995] European lakes. Syntypes: MNHN A-8842 (1, dry) Lac Lémon, ?5467 (1) Lake Geneva. Type catalog: Bertin 1940:302 [ref. 293]. See also Valenciennes in Cuvier & Valenciennes 1848:302 [ref. 1013]. •Synonym of Salmo trutta Linnaeus 1758 or synonym of Salmo lacustris Linnaeus 1758 Great European Lakes stock -- (Kottelat 1997:143 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. letnica, Trutta Karaman [S.] 1924:20 [Pisces Macedoniae; ref. 15902] Lake Ohrid, Republic of Macedonia. Syntypes: (several) IEBMT. One specimen from Prespa Lakes was caught in 2011 [ref. 35029]. •Valid as Salmo letnica (Karaman 1924) -- (Baxter & Stone 1995:188 [ref. 23374], Kottelat 1997:137 [ref. 22952], Dorofeeva 1998:442 [ref. 24680], Fuller et al. 1999:256 [ref. 25838], Delling 2002:299 [ref. 26660], Delling & Doadrio 2005:55 [ref. 28211], Kottelat & Freyhof 2007:422 [ref. 29996], Barbieri et al. 2015:83 [ref. 35029], Schöffmann 2021:159 [ref. 38997], Shumka et al. 2023:39 [ref. 40747]). Current status: Valid as Salmo letnica (Karaman 1924). Salmonidae: Salmoninae. Distribution: Southeastern Europe: Lake Ohrid (Albania and Macedonia). Habitat: freshwater. levenensis, Salmo Yarrell [W.] (ex Walker) 1839:9, Fig. (pt. 2) [A history of British fishes, illustrated by nearly 400 wood-cuts. Supplement.; ref. 17748] Loch Levin, Scotland. Syntypes: whereabouts unknown. Appeared first as name only and not available in Neill 1808:541 [ref. 23048] and in Walker 1812:373 [ref. 23049]. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952], Scott & Crossman 1998:198 [ref. 24570]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. loensis, Salmo fario var. Walbaum [J. J.] 1792:61 [Petri Artedi sueci genera piscium Part 3; ref. 4572] Lake Lo, Cornwall, England. No types known. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. macedonica, Trutta fario var. Karaman [S.] 1924:16 [Pisces Macedoniae; ref. 15902] Mouth of Treska River, Vardar basin, Republic of Macedonia. Syntypes: (several) IEBMT. •Valid as Salmo macedonicus (Karaman 1924) -- (Kottelat 1997:138 [ref. 22952], Kottelat & Freyhof 2007:422 [ref. 29996], Simonović et al. 2007:368 [ref. 33985], Delling 2011:335 [ref. 31191], Barbieri et al. 2015:83 [ref. 35029], Schöffmann 2021:164 [ref. 38997]). Current status: Valid as Salmo macedonicus (Karaman 1924). Salmonidae: Salmoninae. Distribution: Southeastern Europe: Macedonia and Greece. Habitat: freshwater. macroptera, Trutta fario Chichkoff [G.] 1939:121 [French p. 177], Pl. 2 (fig. 4) [Godisnik na Sofijskija Universitet = Annuaire de l'Université de Sofia, Fiziko-Matematiceski Fakultet = Faculté des Sciences Physiques et Mathématiques, Sofija v. 35 (no. 3); ref. 17182] Gorno Ribno Esero [Ezero] and Dolno Ribno Esero [Exero], Struma River basin, Rila range, Bulgaria. Syntypes: whereabouts unknown. •Questionably a synonym of Salmo macedonicus (Karaman 1924) -- (Kottelat 1997:138 [ref. 22952]). •Synonym of Salmo trutta Linnaeus 1758, subspecies fario Linnaeus 1758 -- (Vassilev & Pehlivanov 2005:165 [ref. 28369]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish. macrostigma, Salar Duméril [A. H. A.] 1858:160 [Comptes Rendus Hebdomadaires des Séances de l'Académie des Sciences v. 47; ref. 23040] Oued-el-Abaïch, Kabylie, 40 kilometers west of Collo, Algeria. Syntypes: (2) MNHN A-7585 (1). Type catalog: Bertin 1940:305 [ref. 293] with a holotype. Also appeared in Duméril 1858:396-399, pl. 10 [ref. 23041] and in L'Institute, v. 26:256, and in Bull. Soc. Acclim. Paris, v. 5:444-446 -- with slightly different titles; earliest not researched. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies -- (Berg 1948:259 [ref. 12882], Behnke 1984:125 [ref. 6177], Lelek 1987:114 [ref. 22962], Poll & Gosse 1995:101 [ref. 24781]). •Questionably a synonym of Salmo cettii Rafinesque 1810 -- (Kottelat 1997:135 [ref. 22952], Bianco 2013:3 [ref. 32897]). •Status uncertain -- (Delling & Doadrio 2005:49 [ref. 28211]). •Synonym of Salmo trutta Linnaeus 1758 -- (Tougard et al. 2018:302 [ref. 36173]). •Valid as Salmo macrostigma (Duméril 1858) -- (Turan et al. 2011:32 [ref. 31466] as cf. macrostigma from Turkey, Bianco & Delmastro 2011:2 [ref. 31648], Bianco 2013:2 [ref. 32897], Doadrio et al. 2015:2 [ref. 33950]). •Synonym of Salmo cettii Rafinesque 1810 -- (Splendiani et al. 2019:438 [ref. 37214]). Current status: Valid as Salmo macrostigma (Duméril 1858). Salmonidae: Salmoninae. Distribution: Algeria and Mediterranean Morocco. Habitat: freshwater, brackish, marine. marina, Trutta Moreau [É.] 1881:537 [Histoire naturelle des poissons (Moreau); ref. 3040] Meuse, Seine and Loire rivers, France. Syntypes: MNHN. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. marmorata, Trutta fario var. Siebold [C. T. E. von] 1863:187 [Verhandlungen der Schweizerischen Naturforschenden Gesellschaft zu Samaden 1863 (for 24-26 Aug.); ref. 37429] Adriatic Sea drainages, Ober-Engadin, Switzerland. Syntypes: whereabouts unknown. Subjectively invalid; secondarily preoccupied by Salmo marmoratus Cuvier 1829. •Questionably a synonym of Salmo marmoratus Cuvier 1829 -- (Kottelat 1997:138 [ref. 22952]). 2°Homonym. Current status: Uncertain as Salmo marmoratus Cuvier 1829. Salmonidae: Salmoninae. Habitat: freshwater. marmoratus, Salmo Cuvier [G.] 1829:304 [Le Règne Animal (Edition 2) v. 2; ref. 995] Lakes of Lombardia, Italy. Syntypes: whereabouts unknown. •Synonym of Salmo trutta Linnaeus 1758, subspecies marmoratus (Cuvier 1829) -- (Ćaleta et al. 2015:85 [ref. 34877]). •Valid as Salmo marmoratus Cuvier 1829 -- (Bianco 1988:148 [ref. 22383], Mrakovcic et al. 1995:183 [ref. 23079], Kottelat 1997:138 [ref. 22952], Dorofeeva 1998:438 [ref. 24680], Delling 2002:283 [ref. 26660], Delling & Doadrio 2005:55 [ref. 28211], Schöffmann 2005:20 [ref. 33986], Kottelat & Freyhof 2007:416 [ref. 29996], Simonović et al. 2007:368 [ref. 33985], Zupančič 2008:65 [ref. 29926], Delling 2011:340 [ref. 31191], Bianco 2013:3 [ref. 32897], Ćaleta et al. 2019:178 [ref. 37026], Lorenzoni et al. 2019:253 [ref. 37108], Splendiani et al. 2019:437 [ref. 37214], Kovačić et al. 2020:13 [ref. 37519], Schöffmann 2021:134 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Shumka et al. 2023:40 [ref. 40747]). Current status: Valid as Salmo marmoratus Cuvier 1829. Salmonidae: Salmoninae. Distribution: Southern Europe: Po River basin, lakes Maggiore, Lugano and Poschivo (northern Italy). Introduced elsewhere. Habitat: freshwater. microps, Salmo Hardin [S.] 1862:383 [Öfversigt af Kongliga Vetenskaps-Akademiens förhandlingar, Kungliga Svenska Vetenskapsakademien. v. 18; ref. 18245] Lake Vänern, Sweden. Syntypes: whereabouts unknown. •Synonym of Salmo trutta Linnaeus 1758 or Salmo lacustris Linnaeus 1758 or Salmo truttula Nilsson 1832 -- (Kottelat 1997:143 [ref. 22952]). •Synonym of Salmo trutta Linnaeus 1758. Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. mistops, Salmo Günther [A.] 1866:105, Fig. [Catalogue of the fishes in the British Museum v. 6; ref. 1983] Eidfjord River, Norway. Syntypes: (at least 4) BMNH 1865.8.23.2-4 (3), 1865.8.23.8 (1). •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. montenigrina, Trutta Karaman [S.] 1933:104 [1] [Ribarski List, Polusluzbeni Organ Ministarstva Poljoprivrede, Sarajevo v. 8 (nos 9-10); ref. 18372] Moraca [Morace] River at Titograd, Montenegro. Syntypes: whereabouts unknown. On p. 1 of separate. •Synonym of Salmothymus obtusirostris (Heckel 1851) -- (Kottelat 1997:144 [ref. 22952]). •Synonym of Salmo montenigrina (Heckel 1851). •Valid as Salmo montenigrinus (Karaman 1933) -- (Kottelat & Freyhof 2007:423 [ref. 29996]). Current status: Valid as Salmo montenigrinus (Karaman 1933). Salmonidae: Salmoninae. Distribution: Bosnia-Herzegovina and Montenegro. Habitat: freshwater. orcadensis, Salmo Günther [A.] 1866:91 [Catalogue of the fishes in the British Museum v. 6; ref. 1983] Loch Stennis, Orkney Islands, Scotland. Syntypes: BMNH 1856.9.19.765-766 (2). •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. oxianus, Salmo Kessler [K. F.] 1874:35, Pl. 5 (fig. 21) [Izvestiia Imperatorskago Obschchestva Liubitelei Estestvozaniia, Antropologii i Etnografii v. 11; ref. 2596] Darmut stream, tributary of Kizil-Su River, Upper Amu Darya River basin, Afghanistan. Lectotype: ZMMU P-833. Paralectotypes: BMNH uncat. (1, stuffed), BMNH uncat (skin in bottle); ZMMU P-833a (1), P-1871 (1); none at ZIN. Type catalog: Svetovidova 1978:256 [ref. 9011], Vasil'eva et al. 2001:38-39 [ref. 25714]. Lectotype selected by Salmanov & Dorofeyeva 1990:818 [ref. 20193]. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies oxianus Kessler 1874 -- (Berg 1948:258 [ref. 12882], Dorofeeva 1998:442 [ref. 24680], Salmanov & Dorofeyeva 1990:817 [ref. 20193]). •Listed -- (Kottelat 1997:143 [ref. 22952]). •Valid as Salmo oxianus Kessler 1874 -- (Schöffmann 2021:226 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Çiçek et al. 2023:43 [ref. 40326]). Current status: Valid as Salmo oxianus Kessler 1874. Salmonidae: Salmoninae. Distribution: North Asia: Amu Darya River basin (Kyrgyzstan, Tajikistan and Afghanistan). Habitat: freshwater, brackish, marine. oxyrhynchus, Salmo (Trutta) obtusirostris var. Steindachner [F.] 1882:75, Pl. 4 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 86 (1. Abth.); ref. 4233] Rivers Narenta [Neretva] and Grupa near Metcovich, Croatia. Syntypes: (several) NWM 82297 (2). Species not listed in earlier abstract [ref. 20485]. •Synonym of Salmothymus obtusirostris (Heckel 1851), but a valid subspecies -- (Mrakovcic et al. 1995:184 [ref. 23079], Schöffmann 2021:150 [ref. 38997]). •Synonym of Salmothymus obtusirostris (Heckel 1851) -- (Kottelat 1997:144 [ref. 22952]). •Synonym of Salmo obtusirostris (Heckel 1851) but a doubtful subspecies -- (Snoj et al. 2002:399 [ref. 26461]). Current status: Synonym of Salmo obtusirostris (Heckel 1851). Salmonidae: Salmoninae. Habitat: freshwater. pallaryi, Salmo Pellegrin [J.] 1924:971 [Comptes Rendus Hebdomadaires des Séances de l'Académie des Sciences v. 178; ref. 23042] Lake Aguelman de Sidi Ali or Mohamed, 30 kilometers north-west of Itzer, Middle Atlas, Morocco. Syntypes: (2) MNHN 1923-0066 (1), 1924-0146 (1). Type catalog: Bertin 1940:305 [ref. 293] with a holotype, 1923-66. Also appeared in Pellegrin 1924:182, Fig. [ref. 15462]. Fowler's use of Salmo eriox pallayeri Hoyer & Michalski 1915 was apparently a misspelling as pallayeri of Salmo pallaryi Pellegrin 1924 and with wrong author and date (see Kottelat 1997:142 [ref. 22952]). Salmo fario is the only trout name that appeared in Hoyer & Michalski 1915:212 [ref. 22001]. •Possible synonym of Salmo trutta Linnaeus 1758, subspecies macrostigmus (Duméril 1855) -- (Behnke 1984:125 [ref. 6177]). •Questionably a synonym of Salmo cettii Rafinesque 1810 -- (Kottelat 1997:135 [ref. 22952]). •Valid as Salmo pallaryi Pellegrin 1924 -- (Delling & Doadrio 2005:49, 52 [ref. 28211], Doadrio et al. 2015:2 [ref. 33950], Delling et al. 2023:[6] [ref. 39858]). Current status: Valid as Salmo pallaryi Pellegrin 1924. Salmonidae: Salmoninae. Distribution: North Africa: Lake Aguelman de Sidi Ali (Morocco) [extinct]. Habitat: freshwater. pallasii, Salmo Günther [A.] 1866:112 [Catalogue of the fishes in the British Museum v. 6; ref. 1983] Soldaja, southern shore of Crimea, Ukraine. No types known. Appeared as Salmo trutta (pallasii) on p. 112 and pallasii alone in index; on p. 112 it is stated, "As we have no specimens to decide this point, we are not entitled to propose a change in the specific denomination." Regarded as available. •Synonym of Salmo labrax Pallas 1814 -- (Kottelat 1997:137 [ref. 22952]). Current status: Synonym of Salmo labrax Pallas 1814. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. pellegrini, Salmo trutta var. Werner [F.] 1931:262, Pl. 1 (fig. 1) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftlichen Classe v. 140 (Abt. 1) (nos 3-4); ref. 16344] Ourika stream near Timishi, Grand Atlas, Morocco. Syntypes: MCZ 32705 (1) Oued Ouri, NMW 19546 (1). •Synonym of Salmo trutta Linnaeus 1758 -- (Behnke 1984:125 [ref. 6177]). •Questionably a synonym of Salmo cettii Rafinesque 1810 -- (Kottelat 1997:135 [ref. 22952]). •Status uncertain -- (Delling & Doadrio 2005:61 [ref. 28211]) •Valid as Salmo pellegrini Werner 1931 -- (Doadrio et al. 2015:2 [ref. 33950], Segherloo et al. 2021:10 [ref. 39584]). Current status: Valid as Salmo pellegrini Werner 1931. Salmonidae: Salmoninae. Distribution: North Africa: Morocco. Habitat: freshwater. phinoc, Salmo Shaw [G.] 1804:54 [General zoology or systematic natural history; ref. 4015] Esk River in Cumberland, Great Britain. No types known. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. platycephalus, Salmo (Platysalmo) Behnke [R. J.] 1968:2, Figs. 1-2 [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 66; ref. 250] Tributary of Seyhan River basin, about 30 kilometers south of Pinarbasi, Turkey. Holotype: ZMH H4089. Paratypes: Colorado Coop. Fish. Unit (2). Type catalog: Wilkens 1977:155 [ref. 22013]. •Synonym of Salmo trutta Linnaeus 1758 -- (SuŜnik et al. 2004:947 [ref. 33786]). •Valid as Salmo platycephalus Behnke 1968 -- (Delling & Doadrio 2005:55 [ref. 28211], Fricke et al. 2007:56 [ref. 29533], Turan et al. 2010:362 [ref. 30689], Turan et al. 2011:32 [ref. 31466], Turan et al. 2012:221 [ref. 32276], Turan et al. 2014:284 [ref. 33242], Turan et al. 2014:147 [ref. 33615], Çiçek et al. 2015:150 [ref. 35151], Turan et al. 2017:63 [ref. 35485], Çiçek et al. 2018:22 [ref. 36267], Bayçelebi 2020:760 [ref. 37964], Çiçek et al. 2020:255 [ref. 37644], Schöffmann 2021:198 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Çiçek et al. 2023:426 [ref. 40625], Çiçek et al. 2024:293 [ref. 40789], Turan et al. 2024:90 [ref. 40793], Küçük et al. 2024:400 [ref. 40993]). Current status: Valid as Salmo platycephalus Behnke 1968. Salmonidae: Salmoninae. Distribution: Asia Minor: upper Seyhan River basin, Mediterranean tributary (Turkey). Habitat: freshwater, brackish. polyosteus, Salmo Günther [A.] 1866:111 [Catalogue of the fishes in the British Museum v. 6; ref. 1983] Lapland. Syntypes: BMNH 1862.10.24.15 (1, male), 1862.10.24.16 (1, female), 1862.10.24.18-23 (6). •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. rappii, Salmo Günther [A.] 1866:82 [Catalogue of the fishes in the British Museum v. 6; ref. 1983] Lake Constance, Europe. Syntypes: (3) BMNH 1866.1.10.2-3 (2), ZMB 3718 (1, disintegrated). Based on literature sources and specimens. •Synonym of Salmo trutta Linnaeus 1758 or synonym of Salmo lacustris Linnaeus 1758 -- (Kottelat 1997:143 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. relicta, Trutta salar Malmgren [A. J.] 1863:59 [333] [Kritisk öfversigt af Finlands fisk-fauna; ref. 13933] Lake Ladoga at Keksgolm [Kexholm], Finland [now in Russia]. Syntypes: whereabouts unknown. Original not seen. On p. 333 of German translation. •Synonym of Salmo salar Linnaeus 1758 -- (Berg 1948:229 [ref. 12882], Kottelat 1997:139 [ref. 22952], Dymond 1963:498 [ref. 24571], Denys et al. 2022:392 [ref. 40085]). Current status: Synonym of Salmo salar Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. rhenana, Salmo lacustris var. Fatio [V.] 1890:347 [Faune des Vertébrés de la Suisse. Vol. V; ref. 14495] Rhine River basin, downriver from the falls [of Schaffhausen], Switzerland. Syntypes: MHNG 806.96 (1). Type catalog: Weber 1998:11 [ref. 32707]. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. rhodanensis, Salmo eriox Fowler [H. W.] 1974:62 [Quarterly Journal of the Taiwan Museum (Taipei) v. 27 (nos 1/2); ref. 7180] Rhône River, France. Syntypes: MNHN 1922-0164 (1) discarded in 1947. Based on infrasubspecific Salmo trutta forma major faces rhodanensis Roule 1923. •Valid as Salmo rhodanensis Fowler 1974 -- (Kottelat 1997:138 [ref. 22952], Kottelat & Freyhof 2007:417 [ref. 29996], Schöffmann 2021:102 [ref. 38997]). Current status: Valid as Salmo rhodanensis Fowler 1974. Salmonidae: Salmoninae. Distribution: Western Europe: France, Switzerland, France and Italy. Habitat: freshwater. romanovi, Salmo lacustris var. Kavraisky [F. F.] 1896:43, Pls. 8-9 [Die Lachse der Kaukasusländer und ihrer angrenzenden Meere; ref. 14634] Lake Tabistskhuri, Georgia, Eurasia. Syntypes: (20) ?BMNH 1897.2.9.21-22 (2). •Synonym of Salmo trutta Linnaeus 1758 -- (Berg 1948:252 [ref. 12882]). •Listed -- (Kottelat 1997:144 [ref. 22952]). •Possible synonym of Salmo caspius Kessler 1877 -- (Turan et al. 2022:57 [ref. 39307]). Current status: Synonym of Salmo caspius Kessler 1877. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. salmanata, Trutta Strøm [H.] 1784:123 [Physisk-oeconomisk beskrivelse over Eger-praestgiaeld i Aggerhuus-stift i Norge; ref. 22961] Eger, Buskerud District, Norway. No types known. •Questionably a synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Uncertain as Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. salonitana, Trutta obtusirostris Karaman [S.] 1927:256 [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 2 (nos 1-2); ref. 15859] Jadro River at Split and Krka River at Knin, Croatia. Syntypes: whereabouts unknown. •Synonym of Salmothymus obtusirostris (Heckel 1851), but a valid subspecies -- (Mrakovcic et al. 1995:184 [ref. 23079] as solitana). •Synonym of Salmothymus obtusirostris (Heckel 1851) -- (Kottelat 1997:144 [ref. 22952]). •Synonym of Salmo obtusirostris (Heckel 1851) but a doubtful subspecies -- (Snoj et al. 2002:399 [ref. 26461]). •Valid as Salmo salonitana (Karaman 1927) -- (Zupančič 2008:67 [ref. 29926]) Current status: Synonym of Salmo obtusirostris (Heckel 1851). Salmonidae: Salmoninae. Habitat: freshwater. saxatilis, Salmo Schrank [F. von P.] 1798:320 [Fauna Boica. Durchgedachte Geschichte der in Baieren einheimischen und zahmen Thiere. v. 1; ref. 6444] Bavaria and other localities. No types known. Syntypes: ZSM [old collection] destroyed in WWII. Type catalog: Neumann 2006:279 [ref. 28925]. Based on Bloch, p. 157, Pl. 23 [ref. 17254]. Kottelat 1997:137, 141 [ref. 22952] "restricted" the type locality to Germany, but the Code does not provide for such restriction in that manner. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. septentrionalis, Salmo lacustris var. Fatio [V.] 1890:354 [Faune des Vertébrés de la Suisse. Vol. V; ref. 14495] Original not seen. Unneeded substitute name for Salmo lacustris var. rhenana Fatio 1890. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. spectabilis, Salar Valenciennes [A.] in Cuvier & Valenciennes 1848:340 [Histoire naturelle des poissons v. 21; ref. 1013] Neva River at St. Petersburg, Russia. Syntypes: MNHN 0000-3655-0000-3656 and A-7593 (3). Type catalog: Bertin 1940:304 [ref. 293]. •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:142 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. spurius, Salmo Pallas [P. S.] 1814:343 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Gulf of Finland, Baltic Sea [originally Russian rivers]. Neotype: ZIN 40749. Neotype designated by Bogutskaya & Dorofeeva 2007:136 [ref. 36499]. •In part a synonym of Salmo trutta Linnaeus 1758 -- (Berg 1948:242 [ref. 12882]). •Listed -- (Kottelat 1997:143 [ref. 22952]). •Synonym of Salmo trutta Linnaeus 1758, subspecies trutta Linnaeus 1758 -- (Bogutskaya & Dorofeeva 2007:135 [ref. 36499]). Current status: Uncertain as Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. stroemii, Salmo Gmelin [J. F.] 1789:1374 [Caroli a Linné ... Systema Naturae per regna tria naturae v. 1 (pt 3); ref. 18139] Denmark [now Sondmor, Norway]. See Kottelat 1997:141 [ref. 22952] for locality information. •Questionably a synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Uncertain as Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. sylvaticus, Salmo Gmelin [J. F.] 1789:1367 [Caroli a Linné ... Systema Naturae per regna tria naturae v. 1 (pt 3); ref. 18139] Norway; Berlin, Germany; Danzig [Gdansk], Poland. Syntypes: ZMB (lost). •Synonym of Salmo trutta Linnaeus 1758 -- (Kottelat 1997:141 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. taleri, Trutta Karaman [S.] 1933:2 [Ribarski List, Polusluzbeni Organ Ministarstva Poljoprivrede, Sarajevo v. 7 (nos. 9-10); ref. 18371] Upper Zeta River at Niksic, Lake Skadar basin, Montenegro. Syntypes: whereabouts unknown. •Tentatively valid as Salmo taleri (Karaman 1933) -- (Kottelat 1997:140 [ref. 22952]). •Valid as Salmo taleri (Karaman 1933) -- (Kottelat & Freyhof 2007:426 [ref. 29996], Simonović et al. 2007:368 [ref. 33985], Schöffmann 2021:176 [ref. 38997]). Current status: Valid as Salmo taleri (Karaman 1933). Salmonidae: Salmoninae. Distribution: Southeastern Europe: Montenegro. Habitat: freshwater. trutta, Salmo Linnaeus [C.] 1758:308 [Systema Naturae, Ed. X v. 1; ref. 2787] European rivers. No types known. Neotype designated by Fricke 1999:24 [ref. 24101], but withdrawn in Fricke 2000:639 [ref. 24537]. •See discussion in Kottelat 1997:141 [ref. 22952]. •Valid as Salmo trutta Linnaeus 1758 -- (Berg 1948:235 [ref. 12882], Ovchynnyk 1967:26 [ref. 37559], Svetovidov 1973:146 [ref. 7169], Lee et al. 1980:109 [ref. 22416], Cadwallader & Backhouse 1983:151 [ref. 25473], Behnke 1984:125 [ref. 6177], Svetovidov in Whitehead et al. 1984:382 [ref. 13675], Araga in Masuda et al. 1984:38 [ref. 6441], Smith 1986:230 [ref. 22186], Robins & Ray 1986:76 [ref. 23100], Lelek 1987:101 [ref. 22962], Robison & Buchanan 1988:126 [ref. 23393], Scott & Scott 1988:134 [ref. 25518], Paxton et al. 1989:165 [ref. 12442], Pequeño 1989:29 [ref. 14125], Paulin et al. 1989:91 [ref. 24556], Kawanabe & Mizuno 1989:148 [ref. 25598], McAllister 1990:85 [ref. 14674], Tomelleri & Eberle 1990:52 [ref. 24089], Page & Burr 1991:51 [ref. 18983], Coad 1991:20 [ref. 15702], Boschung 1992:68 [ref. 23239], Etnier & Starnes 1993:348 [ref. 22809], Skelton 1993:261 [ref. 24071], Stearley & Smith 1993:21 [ref. 26683], Gomon et al. 1994:231 [ref. 22532], La Rivers 1994:318 [ref. 23389], Jenkins & Burkhead 1994:580 [ref. 21581], Baxter & Stone 1995:190 [ref. 23374], Coad 1995:25 [ref. 23608], Stauffer et al. 1995:237 [ref. 23376], Cross & Collins 1995:170 [ref. 23394], Economidis 1995:207 [ref. 23025], Apostolidis et al. 1996:551 [ref. 24743], Apostolidis et al. 1996:608 [ref. 24724], Davies & McDowall in McDowall 1996:81 [ref. 22909], Chereshnev 1996:604 [ref. 24659], Andersen & Deacon 1996:126 [ref. 24717], Coad 1996:71 [ref. 25297], Coad 1996:85 [ref. 25304], Chilcott & Humphries 1996:145 [ref. 25528], Mettee et al. 1996:425 [ref. 25836], Sigler & Sigler 1996:204 [ref. 25943], Bianco et al. 1996:55 [ref. 26724], Kottelat 1997:141 [ref. 22952], Murdy et al. 1997:100 [ref. 23144], Reshetnikov et al. 1997:729 [ref. 24702], Thurow et al. 1997:1106 [ref. 25523], Bianco 1997:612 [ref. 26725], Dorofeeva & Savvaitova 1998:36 [ref. 23574], Coad 1998:103 [ref. 23869], Morgan et al. 1998:39 [ref. 24611], Scott & Crossman 1998:197 [ref. 24570], Dorofeeva 1998:442 [ref. 24680], Fricke 1999:28 [ref. 24101], Fuller et al. 1999:258 [ref. 25838], Dyer 2000:89 [ref. 26678], Lyons et al. 2000:51 [ref. 26594], Nakabo 2000:299 [ref. 25086], Novikov et al. 2000:161 [ref. 40929], Hutchins 2001:18 [ref. 25847], Gabrielyan 2001:25 [ref. 25867] as species and subspecies, Bogutskaya et al. 2001:46 [ref. 26178], Bianco & Ketmaier 2001:194 [ref. 26728], Skelton 2001:261 [ref. 27406], Allen et al. 2002:358 [ref. 25930], Nakabo 2002:298 [ref. 26001], Moyle 2002:293 [ref. 26449], Delling 2002:283 [ref. 26660], López et al. 2003:67 [ref. 27366], Wydoski & Whitney 2003:92 [ref. 27816], Hanel 2003:57 [ref. 27894], Nelson et al. 2004:86 [ref. 27807], Bailey et al. 2004:156 [ref. 27882], Miller & Robison 2004:258 [ref. 27931], Boschung & Mayden 2004:354 [ref. 27995], Seegers et al. 2003:40 [ref. 28087], Hammer & Walker 2004:91 [ref. 28129], Menni 2004:85 [ref. 28131], Bogutskaya & Naseka 2004:157 [ref. 28183], Delling & Doadrio 2005:55 [ref. 28211], Hanel & Lusk 2005:305 [ref. 28804], Duhamel et al. 2005:110 [ref. 32059], Schöffmann 2005:20 [ref. 33986], Paxton et al. 2006:415 [ref. 28994], Scharpf 2006:33 [ref. 30386], Fricke 2007:26 [ref. 30577], Fricke et al. 2007:56 [ref. 29533], Kottelat & Freyhof 2007:408 [ref. 29996], Vasil'eva 2007:52 [ref. 30517], McPhail 2007:311 [ref. 31199], Simonović et al. 2007:368 [ref. 33985], Raadik 2008:213 [ref. 30624], Ninua & Japoshvili 2008:168 [ref. 31251], Minckley & Marsh 2009:97 [ref. 31114], Esmaeili et al. 2010:374 [ref. 33391], Lipej & Dulčić 2010:21 [ref. 36649] subspecies trutta, Page & Burr 2011:400 [ref. 31215], Marshall 2011:188 [ref. 31755], Hammer et al. 2012:66 [ref. 32389], Page et al. 2013:87 [ref. 32708], Parin et al. 2014:137 [ref. 33547] suspecies trutta, Sarmiento et al. 2014:135, 189 [ref. 35004], Jouladeh-Roudbar et al. 2015:891 [ref. 34062], McDowall & Roberts 2015:415 [ref. 34244], Jiménez-Prado et al. 2015:293 [ref. 34894], Barbieri et al. 2015:84 [ref. 35029], Mirande & Koerber 2015:49 [ref. 35117], Çiçek et al. 2015:150 [ref. 35151], Reis et al. 2016:18 [ref. 34496], Esmaeili et al. 2017:77 [ref. 35254], Esmaeili et al. 2018:57 [ref. 36089], Fricke et al. 2018:64 [ref. 35805], Kuiter 2018:108 [ref. 36369], Martins & Carneiro 2018:31 [ref. 36550], Tougard et al. 2018:307 [ref. 36173], Ćaleta et al. 2019:179 [ref. 37026], Carneiro et al. 2019:86 [ref. 37250], Lorenzoni et al. 2019:253 [ref. 37108], Çiçek et al. 2020:255 [ref. 37644], Dyldin et al. 2020:84, 103 [ref. 37569], Jouladeh-Roudbar et al. 2020:221 [ref. 37983], Kovačić et al. 2020:12 [ref. 37519], Bañón & Maño 2021:53 [ref. 38883], Iglésias et al. 2021:171 [ref. 38818], Schöffmann 2021:35 [ref. 38997], Segherloo et al. 2021:10 [ref. 39584], Bagley et al. 2023:316 [ref. 41241], Çiçek et al. 2023:43 [ref. 40326], Çiçek et al. 2023:427 [ref. 40625], Çiçek et al. 2023:152 [ref. 40728], Nagy et al. 2023:357 [ref. 40348], Page et al. 2023:83 [ref. 40505], Çiçek et al. 2024:294 [ref. 40789]). Current status: Valid as Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Distribution: Northeastern Atlantic: Norway south to Iberian Peninsula; Baltic Sea; North Sea; Mediterranean Sea; Europe: northeastern Atlantic watersheds and upper Danube basin, south to Sicily; introduced widely elsewhere. Habitat: freshwater, brackish, marine. truttula, Salmo Nilsson [S.] 1832:5 [Prodromus ichthyologiae Scandinavicae; ref. 3204] Sea around Göteborg and Lake Vättern, Sweden. •Synonym of Salmo trutta Linnaeus 1758 or synonym of Salmo lacustris Linnaeus 1758 or valid -- (Kottelat 1997:143 [ref. 22952]). Current status: Uncertain as Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. typicus, Salmo letnica Stefanovic [D.] 1948:181, 4 figs. [Posebna izdanja / Srpska akademija nauka i umetnosti. Beograd 139; ref. 23038] Original not seen. Unneeded replacement name for Trutta letnica Karaman 1924 according to Kottelat 1997:137 [ref. 22952]. •Synonym of Salmo letnica (Karaman 1924) -- (Kottelat 1997:137 [ref. 22952]). Replacement Name. Current status: Synonym of Salmo letnica (Karaman 1924). Salmonidae: Salmoninae. Habitat: freshwater. ungeri, Trutta Vásárhelyi [S.] 1940:188, Fig. 2 [Zoologischer Anzeiger v. 132 (no. 7/8); ref. 23044] Várvölgy creek in Bükk Range, Kom. Borsod, Hungary. Syntypes: (3) whereabouts unknown. •Tentatively a synonym of Salmo labrax Pallas 1814 -- (Kottelat 1997:137 [ref. 22952]). Current status: Synonym of Salmo labrax Pallas 1814. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. variabilis, Trutta Lunel [G.] 1874:146, Fig.; Pls. 16-18 [Histoire naturelle des poissons du bassin du Léman; ref. 18460] Lake Leman, Switzerland. Syntypes: MHNG 807.35 (1), 816.06 (1), 816.07 (1). Type catalog: Weber 1998:11 [ref. 32707]. •Synonym of Salmo trutta Linnaeus 1758 or synonym of Salmo lacustris Linnaeus 1758 [Great European Lakes stock] -- (Kottelat 1997:143 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. vario, Salmo Nau [B. S.] 1787:22 [Oekonomische Naturgeschichte der Fische in der Gegend um Mainz; ref. 18473] Lohr River near Lohr am Main, Bayern, Germany. No types known. Unneeded new name for Salmo fario Linnaeus 1758. •Objective synonym of Salmo fario Linnaeus 1758. •Synonym of Salmo trutta Linnaeus 1758. Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater. venernensis, Salmo Günther [A.] 1866:110 [Catalogue of the fishes in the British Museum v. 6; ref. 1983] Lake Vänern, Sweden. Syntypes: (at least 7) BMNH 1829.11.8.6 (1), 1853.3.16.37 (1, missing), 1862.3.16.2 (1, stuffed and mounted), 1862.3.16.3 (1, stuffed and mounted). Based on literature and specimens. •Synonym of Salmo trutta Linnaeus 1758 -- (Berg 1948:250 [ref. 12882]). •Synonym of Salmo trutta Linnaeus 1758 or Salmo lacustris Linnaeus 1758 or Salmo truttula Nilsson 1832 -- (Kottelat 1997:143 [ref. 22952]). Current status: Synonym of Salmo trutta Linnaeus 1758. Salmonidae: Salmoninae. Habitat: freshwater, brackish, marine. visovacensis, Salmo Taler [Z.] 1950:118, Fig. [Glasnik Hrvatskoga Prirodoslovniga drustva (2B) v. 2 (no. 3); ref. 12923] Visovac Lake, Dalmatia. Syntypes: whereabouts unknown. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies visovacensis Karaman 1938 -- (Mrakovcic et al. 1995:183 [ref. 23079], Ćaleta et al. 2015:85 [ref. 34877]). •Questionably valid as Salmo visovacensis Taler 1950 -- (Kottelat 1997:143 [ref. 22952]). •Synonym of Salmo farioides Karaman 1938 -- (Bianco 2013:4 [2014:65] [ref. 32897]). •Valid as Salmo visovacensis Taler 1950 -- (Schöffmann 2021:146 [ref. 38997]). Current status: Valid as Salmo visovacensis Taler 1950. Salmonidae: Salmoninae. Distribution: Southern Europe: Dalmatia. Habitat: freshwater. zrmanjaensis, Salmo farioides Karaman [S.] 1938:138 [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 18; ref. 17197] Zrmanja River near Obrovac and Zegar, former Jugoslavian Republic of Macedonia. Syntypes: whereabouts unknown. •Synonym of Salmo trutta Linnaeus 1758, but a valid subspecies zrmanjaensis Karaman 1938 -- (Mrakovcic et al. 1995:183 [ref. 23079], Ćaleta et al. 2015:85 [ref. 34877]). •Questionably valid as Salmo zrmanjaensis Karaman 1938 -- (Kottelat 1997:143 [ref. 22952]). •Synonym of Salmo farioides Karaman 1938 -- (Bianco 2013:4 [ref. 32897]). •Valid as Salmo zrmanjaensis Karaman 1938 -- (Schöffmann 2021:145 [ref. 38997]). Current status: Valid as Salmo zrmanjaensis Karaman 1938. Salmonidae: Salmoninae. Distribution: Southeastern Europe: Macedonia. Habitat: freshwater. |