Institute for Biodiversity Science and Sustainability
Online Version, Updated 3 September 2019

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Eschmeyer's Catalog of Fishes
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Román-Valencia, C., R. I. Ruiz C. and R. E. Barriga Salazar  2005 [ref. 28433]
Una nueva especie ecuatoriana del género de peces andinos Grundulus (Characiformes: Characidae). Revista de Biología Tropical v. 53 (no. 3-4): 537-544.