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FOSSIL INSECTS

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ARANEIDAE RECORDED FROM MADAGASCAR

By V. D. Roth, D. Silva Davila, D. Ubick, and C. E. Griswold

(complete synonymies and references at Platnick, N. I. 2003.
The world spider catalog, version 3.5. American Museum of Natural History, online at http://research.amnh.org/entomology/spiders/catalog81-87/index.html)




ARANEIDAE Simon, 1895

Acantharachne Tullgren, 1910
Nomen nudum: A. tuleari Emerit, 1978.- Emerit, 2000: 157, fig. 19.
f giltayi Lessert, 1938....................Congo, Madagascar
  A. g. Lessert, 1938: 451, f. 38-40 (Df).
  A. g. Emerit, 2000: 155, f. 17-18 (f).

f madecassa Emerit, 2000....................Madagascar
  A. m. Emerit, 2000: 153, f. 12-13 (Df).

f milloti Emerit, 2000....................Madagascar
  A. m. Emerit, 2000: 154, f. 14-16 (Df).

Acrosomoides Simon, 1887
f acrosomoides (O. P.-Cambridge, 1879)....................Madagascar
  Gasteracantha a. O. P.-Cambridge, 1879c: 289, pl. 27, f. 19 (Df).
  A. a. Emerit, 1974: 61, f. 20-21, 22A-C, 26C (Tf from Gasteracantha).
  A. a. Emerit, 1982b: 154, f. 1.

Arachnura Vinson, 1863
f scorpionoides Vinson, 1863 *....................Congo, Ethiopia, Madagascar, Mauritius
  Arachnoura s. Vinson, 1863: 291, 310, pl. 13, f. 1 (Df).
  Hapalochrota caudata Keyserling, 1864: 82, pl. 3, f. 6-11 (Df).
  A. scorpionides Uyemura, 1976b: 26, f. 1.2, 4, 6, 2.1-2, 3.1, 4.1 (f).

Araneus Clerck, 1757
j isabella (Vinson, 1863)....................Madagascar
  Epeira i. Vinson, 1863: 157, 308, pl. 4, f. 2 (Dj).

f kraepelini (Lenz, 1891)....................Madagascar
  Epeira k. Lenz, 1891: 176, pl. 2, f. 16 (Df).

f lenzi (Roewer, 1942)....................Madagascar
  Epeira albomaculata Lenz, 1886: 388, pl. 10, f. 1 (Df; preoccupied by Lucas, 1846).
  A. l. Roewer, 1942a: 810 (replacement name).

mf madagascaricus (Strand, 1908)....................Madagascar
  Aranea rufipalpis madagascarica Strand, 1908e: 290, 293 (Dmf).
  Aranea m. Roewer, 1942a: 810.

f margitae (Strand, 1917)....................Madagascar
  Epeira cinerea Lenz, 1886: 390, pl. 10, f. 13 (Df; preoccupied).
  Aranea m. Strand, 1917c: 71 (replacement name).

jj nossibeus (Strand, 1907)....................Madagascar
  Aranea nossibea Strand, 1907d: 732 (Djj).
  Aranea nossibea Strand, 1907o: 51 (Djj).

f pallescens (Lenz, 1891)....................Madagascar
  Epeira p. Lenz, 1891: 174, pl. 2, f. 15 (Df).

f saccalava (Strand, 1907)....................Madagascar
  Aranea s. Strand, 1907d: 731 (Df).
  Aranea s. Strand, 1907o: 57 (Df).

f sambava (Strand, 1907)....................Madagascar
  Aranea s. Strand, 1907d: 731 (Df).
  Aranea s. Strand, 1907o: 61 (Df).

Argiope Audouin, 1826
f coquereli (Vinson, 1863)....................Zanzibar, Madagascar
  Epeira coquerelii Vinson, 1863: 201, 312, pl. 8, f. 1 (Df).
  A. c. Karsch, 1879a: 339.
  A. c. Bjørn, 1997: 217, f. 43-48 (f).

mf ranomafanensis Bjørn, 1997....................Madagascar
  A. r. Bjørn, 1997: 235, f. 93-98 (Dmf).

Augusta O. P.-Cambridge, 1877
mf glyphica (Guérin, 1839) *....................Madagascar
  Gasteracantha g. Guérin, 1839: 110 (Df).
  Augusta papilionacea O. P.-Cambridge, 1877c: 37, pl. 7, f. 6 (Df).
  A. g. Emerit, 1974: 33, f. 3A-B, 4A-F, 5A-C, 6B (Tf from Gasteracantha, Dm).
  A. g. Scharff & Coddington, 1997: 372, f. 4 (m).

Caerostris Thorell, 1868
f cowani Butler, 1882....................Madagascar
  C. c. Butler, 1882a: 103, pl. 6, f. 4 (Df).
  C. excellens Butler, 1882a: 105, pl. 6, f. 5 (Df).
  C. c. Grasshoff, 1984: 755, f. 34-36 (f, S).

f ecclesiigera Butler, 1882....................Madagascar
  C. e. Butler, 1882a: 102, pl. 6, f. 1-2 (Df).
  C. e. Grasshoff, 1984: 754, f. 31-33 (f).

f extrusa Butler, 1882....................Madagascar
  C. e. Butler, 1882a: 100, pl. 6, f. 3 (Df).
  C. bankana Strand, 1915d: 225 (Df).
  C. e. Grasshoff, 1984: 752, f. 28-30 (f, S).

f hirsuta (Simon, 1895)....................Madagascar
  Trichocharis h. Simon, 1895a: 835 (Df).
  C. h. Grasshoff, 1984: 756, f. 4, 39 (f).

mf mitralis (Vinson, 1863) *....................Central Africa, Madagascar
  Epeira paradoxa Doleschall, 1859: 37, pl. 9, f. 11, pl. 10, f. 8 (Df; preoccupied by Lucas, 1846).
  Epeira m. Vinson, 1863: 230, 314, pl. 9, f. 2-4 (Df).
  C. m. Grasshoff, 1984: 743, f. 10, 13, 19-21, 45 (mf, S).

mf sexcuspidata (Fabricius, 1793)....................Africa, Madagascar, Comoro Is., Aldabra
  Aranea s. Fabricius 1793: 427 (Df).
  Epeira tuberculosa Vinson, 1863: 228, 314, pl. 14, f. 2 (Df).
  C. s. Grasshoff, 1984: 736, f. 8-9, 11, 14-18, 46-47 (mf, S).

Chorizopes O. P.-Cambridge, 1870
mf antongilensis Emerit, 1997....................Madagascar
  C. a. Emerit, 1997: 44, f. 1-3 (Dmf).

mf madagascariensis Emerit, 1997....................Madagascar
  C. m. Emerit, 1997: 47, f. 4-5 (Dmf).

Coelossia Simon, 1895
f trituberculata Simon, 1903....................Mauritius, Madagascar
  C. t. Simon, 1903b: 139 (Df).

Cyclosa Menge, 1866
m hova Strand, 1907....................Madagascar
  C. h. Strand, 1907d: 730 (Dm).
  C. h. Strand, 1907o: 47 (Dm).

mf sanctibenedicti (Vinson, 1863)....................Madagascar, Réunion
  Epeira sancti-benedicti Vinson, 1863: 204, 312, pl. 14, f. 3 (Df).
  C. s. Lopez, 1990: 18, f. 7 (Dm).

Cyphalonotus Simon, 1895
f columnifer Simon, 1903....................Madagascar
  C. c. Simon, 1903b: 140 (Df).

Cyrtarachne Thorell, 1868
mf ixoides (Simon, 1870)....................Mediterranean to Georgia, Madagascar
  Peltosoma i. Simon, 1870b: 315, pl. 3, f. 8 (Df).
  C. tuberculiferum Berland, 1927b: 10, f. 1 (mf).
  C. ixodoides Emerit, 2000: 148, f. 3-5 (f).

f madagascariensis Emerit, 2000....................Madagascar
  C. m. Emerit, 2000: 147, f. 1-2 (Df).

Cyrtophora Simon, 1864
mf citricola (Forskål, 1775) *....................Old World, Hispaniola, Colombia
  Aranea c. Forskål, 1775: 86 (D).
  Epeira opuntiae Vinson, 1863: 212, 313, pl. 9, f. 1 (mf, in part).
  Epeira flava Vinson, 1863: 222, 313, pl. 8, f. 3 (Df).
  Epeira purpurea Vinson, 1863: 225, 313, pl. 8. f. 4 (Df).
  C. c. Scharff & Coddington, 1997: 375, f. 21 (m).
  C. c. Levy, 1998a: 323, f. 47-56 (mf).

Exechocentrus Simon, 1889
f lancearius Simon, 1889 *....................Madagascar
  E. l. Simon, 1889b: 227 (Df).
  E. l. Simon, 1895a: 888, f. 951-953 (f).
  E. l. Emerit, 2000: 159, f. 20-21 (f).

Gasteracantha
Sundevall, 1833
mf rhomboidea madagascariensis Vinson, 1863....................Madagascar
  G. m. Vinson, 1863: 242, 315, pl. 9, f. 6 (Df).
  G. r. m. Emerit, 1974: 91, f. 27C, 28C, 36, 37A-E (f, Dm).

f rufithorax Simon, 1881....................Madagascar
  G. r. Simon, 1881e: 2 (Df).
  G. r. Emerit, 1974: 97, f. 38B, 41B (f, removed from S).

f sanguinolenta andrefanae Emerit, 1974....................Madagascar
  G. s. a. Emerit, 1974: 83, f. 33A-C (Df).

mf sanguinolenta bigoti Emerit, 1974....................Madagascar
  G. s. b. Emerit, 1974: 84, f. 28E, 34, 35A-C (Dmf).

mf sanguinolenta mangrovae Emerit, 1974....................Madagascar
  G. s. m. Emerit, 1974: 87, f. 28D, 32A (Dmf).

mf thorelli Keyserling, 1864....................Madagascar
  G. thorellii Keyserling, 1864: 67, pl. 1, f. 4 (Df).
  G. t. Emerit, 1974: 111, f. 28B, 48C, 50, 51A-C (f, Dm).

f versicolor avaratrae Emerit, 1974....................Madagascar
  G. v. a. Emerit, 1974: 109, f. 48A, 49A-C (Df).

mf versicolor formosa Vinson, 1863....................Madagascar
  G. f. Vinson, 1863: 244, 315, pl. 9, f. 7 (Df).
  G. f. O. P.-Cambridge, 1879c: 285, pl. 26, f. 11 (f).
  G. v. f. Emerit, 1974: 109, f. 2A-C, 27A, 28A, 29, 45A-E, 48B (f, placed as subspecies, Dm).

Isoxya Simon, 1885
mf cowani (Butler, 1882)....................Madagascar
  Gasteracantha c. Butler, 1882b: 766, pl. 57, f. 5 (Dm).
  Gasteracantha c. Dahl, 1914: 255 (Df).
  I. c. Emerit, 1974: 49, f. 7A, 11, 12A-C, 16A (Tmf from Gasteracantha).

mf mahafalensis Emerit, 1974....................Madagascar
  I. m. Emerit, 1974: 43, f. 7C, 8, 9A-C, 10A-C, 16B (Dmf).

f milloti Emerit, 1974....................Madagascar
  I. m. Emerit, 1974: 55, f. 15, 16C (Df).

f reuteri (Lenz, 1886)....................Madagascar
  Gasteracantha r. Lenz, 1886: 382, pl. 10, f. 3 (Df).
  I. r. Emerit, 1974: 51, f. 7B, 13, 14A-C, 16D (f).

Larinia Simon, 1874
(=Drexelia McCook, 1892 by Harrod, Levi & Leibensperger, 1991: 243, after Levi, 1975b: 102)
f tamatave (Grasshoff, 1971)....................Madagascar
  Drexelia t. Grasshoff, 1971a: 91, f. 42a-d (Df).

Madacantha Emerit, 1970
mf nossibeana (Strand, 1916) *....................Madagascar
  Gasteracantha n. Strand, 1916b: 64 (Df).
  Madacantha n. Emerit, 1974: 66, f. 23, 24A-C, 25A-B, 26B (mf).

Nemoscolus
Simon, 1895
f waterloti Berland, 1920....................Madagascar
  N. w. Berland, 1920a: 385, f. 1-5 (Df).

Neoscona
Simon, 1864
mf angulatula (Schenkel, 1937)....................Madagascar, Aldabra, Kenya
  Araneus angulatulus Schenkel, 1937: 383, f. 4 (Df).
  N. dripaca Roberts, 1983: 275, f. 199-201, 211-212 (Dm).
  N. a. Grasshoff, 1986: 106, f. 153-159 (Tf from Araneus, Sm).

mf cereolella (Strand, 1907)....................Congo, East Africa, Madagascar
  Aranea c. Strand, 1907d: 732 (Dmf).
  Afraranea c. Archer, 1951a: 22 (Tmf from Araneus).

mf moreli (Vinson, 1863)....................Cuba to Argentina, Africa to Seychelles
  Epeira morelii Vinson, 1863: 166, 309, pl. 4, f. 4 (Dm).
  N. m. Grasshoff, 1986: 56, f. 71-78 (mf, S).

mf subfusca (C. L. Koch, 1837)....................Old World
  Atea s. C. L. Koch, 1837b: 4 (D).
  Araneus hoffmanni Schenkel, 1937: 384, f. 5 (Df).
  N. s. Grasshoff, 1986: 15, f. 2, 4, 11-24 (mf, S).

mf triangula (Keyserling, 1864)....................Cape Verde to India
  Epeira nocturna Vinson, 1863: 161, 309, pl. 4, f. 3 (Df; preoccupied by Linnaeus, 1758).
  Epeira triangula Keyserling, 1864: 98, pl. 5, f. 12-14 (Dmf).
  N. t. Grasshoff, 1986: 84, f. 124-135 (mf, S).

m triangula mensamontella (Strand, 1907)....................Madagascar
  Aranea annulata m. Strand, 1907a: 534 (Dm).
  Aranea annulella m. Strand, 1907n: 623.

Paraplectana
Brito Capello, 1867
f walleri (Blackwall, 1865)....................West, Central Africa, Madagascar
  Eurysoma w. Blackwall, 1865b: 349 (Df).
  P. w. Simon, 1895a: 876, f. 941 (f).
  P. w. Emerit, 2000: 152

Pararaneus Caporiacco, 1940
f uncivulva (Strand, 1907)....................Madagascar
  Aranea u. Strand, 1907a: 535 (Df).
  P. u. Grasshoff, 1968: 94, f. 66a-d, 74-82 (Tf from Araneus).

Parmatergus Emerit, 1994
mf coccinelloides Emerit, 1994 *....................Madagascar
  P. c. Emerit, 1994: 159, f. 1-2, 3A-C, 4A-C, pl. IA-G, IIA-D (Dmf).

mf coccinelloides ambrae Emerit, 1994....................Madagascar
  P. c. a. Emerit, 1994: 163, f. 5 (Dmf).
  mf lens Emerit, 1994....................Madagascar
  P. l. Emerit, 1994: 164, f. 6A-C, 7A-B, pl. IIE (Dmf).

Pasilobus Simon, 1895
f antongilensis Emerit, 2000....................Madagascar
  P. a. Emerit, 2000: 152, f. 10-11 (Df).

f capuroni Emerit, 2000....................Madagascar
  P. c. Emerit, 2000: 151, f. 8-9 (Df).

Poltys C. L. Koch, 1843
f kochi Keyserling, 1864....................Mauritius, Madagascar
  P. k. Keyserling, 1864: 84, pl. 3, f. 1-5 (Df).
  P. madagascariensis Lenz, 1886: 387 (Df).
  P. k. Simon, 1889b: 225.

f reuteri Lenz, 1886....................Madagascar
  P. r. Lenz, 1886: 386 (Df).

j vesicularis Simon, 1889....................Madagascar
  P. v. Simon, 1889b: 225 (Dj).

Prasonica Simon, 1895
mf albolimbata Simon, 1895 *....................Congo, Madagascar
  P. a. Simon, 1895a: 794 (Df).
  P. a. Grasshoff, 1971b: 299, f. 52c, 53a-f (Simon's f is juvenile, Dmf).

f seriata Simon, 1895....................Africa, Madagascar
  P. s. Simon, 1895a: 794 (Df).
  P. s. Grasshoff, 1971b: 301, f. 49a-b, 50d-f, 51b-c, 52d, 56a-b, 57a-f (f, Dm, S)

Prasonicella Grasshoff, 1971
m cavipalpis Grasshoff, 1971 *....................Madagascar
  P. c. Grasshoff, 1971b: 307, f. 62a-c (Dm).

Pronous Keyserling, 1881
f tetralobus Simon, 1895....................Madagascar
  P. t. Simon, 1895a: 863 (Df).
  P. t. Emerit, 1974: 116

Pycnacantha
Blackwall, 1865
m fuscosa Simon, 1903....................Madagascar
  P. f. Simon, 1903b: 139 (Df).

Thelacantha Hasselt, 1882
mf brevispina (Doleschall, 1857) *....................Madagascar, India to Philippines, Australia
  Plectana b. Doleschall, 1857: 423 (Df).
  Gasteracantha borbonica Vinson, 1863: 236, 315, pl. 9, f. 5 (Df).
  T. b. Emerit, 1974: 57, f. 17, 18A-B, 19A-C, 26A, pl. 5, f. E (mf).


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