Institute for Biodiversity Science and Sustainability
Online Version, Updated 30 March 2017


Catalog of Fishes
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Catalog of Fishes - version of 30 March 2017 (Continuously updated since the early 1980s.)

Richard van der Laan is now a co-author of the Catalog of Fishes. This becomes Eschmeyer, W. N., R. Fricke, and R. van der Laan.

Other projects include Fricke (early references, new references, and corrections).

In this edition, we provide 123 new species in 2017.

Bill Eschmeyer, Florida Museum of Natural History, 1659 Museum Road, Gainesville, FL 32611 USA and California Academy of Sciences.

We are making additions to:

Van der Laan, R., W. N. Eschmeyer & R. Fricke (2014) (11 Nov.), Family-group names of Recent fishes. Zootaxa Monograph 3882 (1), 1-230. DOI 10.11646/zootaxa.3882.1.1

Download the addenda to the family-group list as a PDF.

See Family Group Names for future updates by R. van der Laan.

By default, search results will not include item 3 names -- unavailable names that detract from the main list of available names. These are mostly names mentioned in synonymy, names in lists, etc. They are still in the database, so if you want to see them, check the box labeled Include unavailable names before searching.

Citing the Catalog of Fishes

Eschmeyer, W. N. and R. Fricke, and R. van der Laan (eds).  CATALOG OF FISHES: GENERA, SPECIES, REFERENCES. ( Electronic version accessed dd mmm 2017. [This version was edited by Bill Eschmeyer.]

Fricke, R. (ed).  REFERENCES. ( Electronic version accessed dd mmm 2017. [Ron Fricke devoted much time to improving references and journals, many of which had not been examined in 15 to 25 years. Please remember that the title of a paper often differs in three places: the title on the paper itself, the contents listed by the journal, and in the PDF provided by the journal. Ron tends to follow the title on the pdf for recent papers and the title on the original for earlier publications. Many of our dates of publication are documented for priority purposes. Some current journals have a target date, but actual publication is often delayed. For nomenclatural date problems, we will help if we can.]

van der Laan, R., W. N. Eschmeyer and R. Fricke.  FAMILY-GROUP NAMES. ( Electronic version accessed dd mmm 2017.

Fricke, R. & Eschmeyer, W. N.  JOURNALS. ( Electronic version accessed dd mmm 2017. [Includes all journals appearing in the Catalog, including publication information and ISSN numbers.]

Fricke, R. & Eschmeyer, W. N.  GUIDE TO FISH COLLECTIONS. ( Electronic version accessed dd mmm 2017. [Arranged by museum abbreviation and by country, includes type catalogs and historical publications and www sites where available.]

Eschmeyer, W. N. & Fong, J. D.  SPECIES BY FAMILY/SUBFAMILY. ( Electronic version accessed dd mmm 2017. [Recalculated with each new version; based on current literature, this provides all available species names, valid species, and species described in the last 10 years by family/subfamily.]

Please remember that if you got to the Catalog of Fishes from Fishbase, you are NOT in Fishbase. Fishbase is a more comprehensive database for a different audience, but they use Catalog of Fishes information in their authority file.

Some other useful sites for fishes are (1) Fishbase [Fishbase team, Manila]; (2) -- A comprehensive database on Shorefishes of the Tropical Eastern Pacific (and of the eastern Atlantic) [Ross Robertson]; (3) -- A database on elasmobranchs, with over 13,000 references, information on taxa, type specimens, etc. [Jürgen Pollerspöck]; (4) -- a comprehensive relational database of more than 99,350 scientific species name combinations and over 34,300 well-identified (mostly marine) fish pictures [Dennis Polack]; (5) Freshwater Fish List, 19th edition October 2016; (6) ETYFish Project (derivations of Fish Names) by Christopher Scharpf; and (7) BioStor, for Biodiversity Heritage Library references. A very useful site is [not associated with Fishbase or Worldfish]. Maintained by Michal Mikšik and Erwin Schraml. They post new fish taxa and references as they are published.


A summary paper was published in 2010 from the Catalog of Fishes database that might be of interest to users of the Catalog. "Marine fish diversity: history of knowledge and discovery (Pisces)." By William N. Eschmeyer, Ronald Fricke, Jon D. Fong & Dennis A. Polack. Zootaxa, no. 2525:19-50 (2 July 2010, open access, click title to download).

As always, notifying us of missed original descriptions and errors in the database or providing PDFs of recent papers is most welcome.

Bill Eschmeyer edited type localities and standardized many place names. Our style is to use current place names found on an English map (or from Wikipedia and Google sites). Should you need original localities, they can be found in the original descriptions or from particular museum holdings (often online).

Many scientific names of fish species are based on personal names; some were found to be incorrect. For example, Jordan and co-workers spelled Japanese names ending in "a" with a terminal "e" as in matsubarae; that is acceptable, with the name Latinized first. So those names do not end in "i". Remarks are provided by Eschmeyer in this version for most species based on personal names. For a review of this very technical subject, see Dubois 2007: Zootaxa No. 1550:49-68.

New treatment of synonyms: Often an author revises a genus and treats a species in detail but does not mention the species synonyms (if any). If he/she does not mention the synonyms (rather common in recent treatments) and moves the valid “parent” species to a different genus, do we move the synonyms automatically, although the revising author may not have studied or assessed the synonyms? In the past, we did not move the synonyms. Typically, this often resulted in the synonyms being in a different current genus than the valid parent species. We now move the synonyms to the genus of the valid parent species, but no “status reference” showing the new generic placement is found with the synonym. Some synonyms will be moved incorrectly, but it is considered more desirable to have the potential synonyms in the same current genus as the valid parent species.

Some conventions used in the Catalog

Author plus date and a species code: The code of Zoological Nomenclature does not require a comma between the author and date, only a suggestion. In fact, we consider that confusing as that is traditionally used to show a cited reference in journal publications and not a species authorship and date. Genus+species+author+date (with parentheses as needed) is a (nearly) unique code or formula to define every species [2 known exceptions]. So you will find no comma between the author and date. Some journals are now treating species in this way.

Figures: Under a new species heading, some journals list (a) only the major figure, some list (b) all figures (including maps, habitats and graphs), and (c) some give figures showing the full view and any illustrated parts of the species. We give all figures that show a full view and any anatomical part of the species, so our listing of figures often differs from the treatment by a journal in the original description (as listed in the species heading).

Type localities: We give a current place name that one will find on a Google search or in an ‘English’ atlas. For some species described many years ago, one may find both the original locality and in parentheses a modern name for that locality. Many collections now have their holdings on-line, and it is there one can usually find the actual original locality if needed. Current localities are often augmented by additional information, such as coordinates.

Authors: We have added initials in brackets to distinguish all individual authors of genera and species.

Other sections

Species of Fishes by Family/Subfamily -- Total valid taxa reflect new taxa added minus taxa moved into synonymy plus taxa newly moved to valid. The classification is as used in the Catalog of Fishes. Prepared by Bill Eschmeyer and Jon Fong, with modifications by Richard van der Laan.

Family-Group Names -- Priority applies to Family-group names. It did not in early years. As classifications are refined with use of more subfamilies and tribes, it is required to use the oldest available family-group name for these new taxa.

Ichthyological Collections -- Presented are those specimen collections mentioned in the main Catalog of Fishes database, especially as repositories of type specimens. Besides museum names and abbreviations, this includes WWW sites, cross-referencing of abbreviations, and selection of museums by country, along with type catalogs, historical articles, name changes, transfers, and other items for individual collections. Prepared by Ron Fricke and Bill Eschmeyer.

Ichthyological Journals -- There are nearly 2600 individual journals or monograph series in the Catalog of Fishes database. We provide full citations, and other information such as changes in titles, published duration, ISSN numbers, WWW sites, etc. Prepared by Ron Fricke and Bill Eschmeyer.


I am fortunate to have Sherry Bostick as a technician who enters new species and other information under my direction; she is an excellent typist, find errors, and I look forward to her visits.

Acknowledgments for Bill Eschmeyer (2016): K. V. Akhilesh, Gerry Allen, Allen Allison, William Anderson, Marcelo Andrade, Carlos A. Ardila Rodriguez, Carole Baldwin, Philippe Béarez, Vinicius Bertaco, Roger Bills, Heiko Bleher, Philippe Borsa, Sherry Bostick, David Boyd, Bárbara Calegar, Emily Capuli, Fernando Carvalho, Tiago Carvalho, Dave Catania, Roman Causse, Paula Chainho, Kevin Conway, Raphael Covain, William Crampton, Erwin Delrieu-Trottin, Gabriel Deprá, Rafael Bañón Díaz, Carlos DoNascimiento, Yury Dyldin, Anthony Echelle, Lee Finley, Jon Fong, Tom Fraser, Matthias Geiger, Dani Golani, Martin Gomon, Karsten Hartel, Iraj Hashemzade, Mysi Hoang, Byron Hosking, M. Shahadat Hossain, Butch (Alexander) Hulley, Hisashi Imamura, Adrian Indemaur, Tomio Iwamoto, Juan Jacobo Schmitter-Soto, Laith Jawad, Howard Jelks, David Johnson, K. K. Joshi, Max Kasparek, Jinkoo Kim, Peter Konstantinidis, Sven Kullander, Richard van der Laan, Kuldeep K. Lal, Peter Last, Sebastien Lavoue, Boris Levin, François Libert, Thomas Litz, Hugo López, Nathan Lujan, John Lundberg, Hairong Luo, James Maclaine, Luiz Malabarba, Larry Marcelo, John McCosker, John McEachran, Catherine Mecklenburg, Michael Mincarone, Marcos Mirande, Hiroyuki Motomura, Hamed Mousavi-Sabet, Thomas Munroe, Diego Nadalin , Gavin Naylor, A. L. Netto-Ferreira, Dirk Neumann, Jørgen Nielsen, Makoto Okamoto, Larry Page, Mats Petersson, Ana Petry, Paulo Petry, Fritz Pfeil, Ted Pietsch, John Pogonoski, Stuart Poss, Peter Psomadakis, Brad Pusey, John Randall, Roberto E. dos Reis, Alexandre Ribeiro, Adela Y. Roa-Varon, Ross Robertson, Rob Robins, Uwe Römer, Juan Jose Rosso, Ronald de Ruiter, Barry Russell, Mark Sabaj Pérez, Artem Prokofiev, Christopher Scharpf, Ray Schmidt, Erwin Schraml, Werner Schwarzhans, Richard Sexton, M. Stehmann, Carla Stout, Camm Swift, Armando Jorge Telese Almeida, Ralf Thiel, Luke Tornabene, Davut Turan, James Van Tassell, James A. Vanegas-Rios, Katerina Vasil'eva, Benjamin Victor, Gregory Watkins-Colwell, Simon Weigmann, William White, Jeff Williams, Rick Winterbottom, Hermann. J. Wischmann, Uusuke Yamanoue, Lei Yang, Gus Yearsley, Henry Dario Agudelo Zamora, Angela Zanata, Cláudio Henrique Zawadzki and Mikhail Zhukov. Plus any others we may have missed.

Acknowledgments for Bill Eschmeyer (2017): K. V. Akhilesh, Gerry Allen, William D. Anderson Jr, Carlos A. Ardila Rodriguez, Tim Berra, David Boyd, Bárbara Calegari, Miguel Caneiro, Dave Catania, David De Weirdt, Rafael Banon Diaz, Fabio Di Dario, Carlos Donascimiento, Yury Dyldin, Dave Ebert, Carl Ferraris, Ralph Foster, Benjamin Frable, Libert Francois, Tom Fraser, Hugo Gante, Renny Kurnia Hadiaty, Karsten Hartel, Elaine Heemstra, Hans Ho, Erling Holm, Hisashi Imamura, Laith Jawad, Dave Johnson, Max Kasparek, Seishi Kimura, Stefan Koerber, Muddu Krishna, Richard van der Laan, Helen Larson, Peter Last, Thomas Litz, John Lundberg, Nathan Lujan, James Maclaine, Douglas F. Markle, Eddie Matheson, Mizuki Matsunuma, Marcelo Roberto Souto de Melo, Marcos Mirande, Hiroyuki Motomura, Dirk Neumann, Leo Nico, Jørgen Nielsen, Huu Duc Nguyen, Larry Page, Keith Phillip, Dennis Polack, Artem Prokofiev, John Randall, Lucia Rapp Py-Daniel, Roberto E. Reis, Rob Robins, Juan Jose Rosso, Frank Schäfer, Christopher Scharpf, Ulrich Schliewen, Erwin Schraml, David G. Smith, Bill Smith-Vaniz, Matthias Stehmann, John Sullivan, Milton Tan, Luke Tornabene, Denis Tweddle, James Anyelo Vanegas-Rios, Benjamin Victor, Chelsea Walery, Jouke van der Zee, Mikhail Zhukov and Dan Zimberlin. Plus any others we may have missed.

Acknowledgments for Ron Fricke (2016): László Antal, William D. Anderson Jr., Juan Carlos Arronte, Muthukumarasamy Arunachalam, Andrey Balanov, Vinicius Bertaco, K. K. Bineesh, Sergey Bogorodsky, Jörg Bohlen, David Boseto, Pedro Bragança, Fernando Carvalho, Wei-Jen Chen, Xiaoyong Chen, Erdoğan Çiçek, Kendall Clements, Oliver Crimmen, David Darom, Giuliano Doria, Du Li-Na, Jean-Dominique Durand, Yuri Dyldin, Marco Endruweit, Hamid Reza Esmaeili, Thomas Fraser, Stephen Gaimari, Matthias Geiger, Daniel Golani, Menachem Goren, Lubomir Hanel, Adeljean Ho, Peter Hundt, Isaac Isbrücker, Laith Jawad, Oliver Jewell, K. Kannan, E. S. Karmovskaya, Murat Kaya, Seishi Kimura, Marcus Knight, Alexander Kotlyar, Friedhelm Krupp, Aleksandr Kucheryavyy, Andrey Kukhlevskiy, Kuldeep Lal, Samuel Lalronunga, Edouard Lavergne, Michael Littmann, Thomas Litz, Adriana López-Arbarello, Sara Lourie, Carlos de Lucena, James Maclaine, Christophe Mailliet, Enrique Marschoff, Jesús Matallanas, Mizuki Matsunuma, Marcelo Melo, Juan Marcos Mirande, Hiroyuki Motomura, Thomas Near, Kongbrailatpam Nebeshwar, Heok Hee Ng, Nguyen Huu Duc, Dalton Nielsen, Willian Ohara, Makoto Okamoto, Alla Oleinik, Francesc Ordines, Thomas Orrell, Rafaela Ota, Müfit Özulug, Larry Page, Jorge Palma, Éric Parmentier, Didier Paugy, Mathews Plamoottil, Valentina Prirodina, Jürgen Pollerspöck, Artem Prokofiev, Peter Psomadakis, Brad Pusey, Olga Radchenko, Sihem Rafrafi-Nouira, Jack Randall, Bertrand Richer de Forges, Pedro Pereira Rizzato, Tyson Roberts, Cesar Román-Valencia, Uwe Römer, Barry Russell, Ivan Sazima, Christopher Scharpf, Ray Schmidt, Susana Schönhuth, Erwin Schraml, Bernard Séret, Bungdon Shangningam, Sergei Shedko, Bikramjit Sinha, Paul Skelton, Vic Springer, Nir Stern, Melanie Stiassny, Tan Heok Hui, Davut Turan, Peter Unmack, Stefano Valdesalici, Katerina Vasil'eva, Anatoly Velikanov, Kirill Vinnikov, Jacob Vinu, Waikhom Vishwanath, Olga Voskoboinikova, Emmanuel Vreven, William White, Alan Whitfield, Peter Wirtz, Uwe Zajonz, Cláudio Zawadzki.

Acknowledgments for Ron Fricke (2017): Windsor Aguirre, K. V. Akhilesh, Felipe Alonso, Muthukumarasamy Arunachalam, Carole Baldwin, Michel Bariche, Mathias Barrett, Sudesh Batuwita, Philippe Béarez, Tim Berra, Ricardo Betancur-R, Philippe Borsa, Pedro Bragança, Bárbara Calegari, Marie-Noëlle de Casamajor, Natalia Chernova, Arieli Cherobim, Wilson Costa, David de Weirdt, Jakov Dulčić, Soheil Eagderi, Marco Endruweit, Semih Engin, Hamid Esmaeili, Daniel Golani, Vladimir Golovanov, İskender Gülle, Lubomir Hanel, Ian Harrison, Elaine Heemstra, Muhammad Iqbal, Fernando Jerep, Dave Johnson, Unmesh Katwate, Toshio Kawai, Stefan Koerber, Alexander Kotlyar, Thomas Litz, Paulo Lucinda, Leonid Manilo, Tatiane Mariguela, Mizuki Matsunuma, Bruno Melo, Rico Morgenstern, Bruce Mundy, Jørgen Nielsen, Willian Ohara, Makoto Okamoto, Larry Page, Mathews Plamoottil, John Pogonoski, Gianluca Polgar, Artem Prokofiev, Richard Pyle, Jack Randall, Christopher Scharpf, Ingo Schindler, Erwin Schraml, Sally Schramm, Ying Giat Seah, Iraj Segherloo, Hiroshi Senou, Malcolm Smale, Melanie Stiassny, Ana Swarça, Armi Torres, Davut Turan, Ekaterina Vasil'eva, Francisco Villa-Navarro, Waikhom Vishwanath, Elena Voronina, Olga Voskoboinikova, Emmanuel Vreven, Wolmar Wosiacki, Yusuke Yamanoue, Akane Yoshikawa, Hyo Jae Yu.

Acknowledgments for Richard van der Laan (2016): Carlos Ardila, Alexandre Bannikov, Ralf Britz, Rudolf Caers (†), Paul van der Laan, Peter Last, Anton Lamboj, Luiz Malabarba, Herman Meeus, Bruno Melo, Marcos Mirande, Peter Rask Møller, Larry Page, Carla Pavanelli, Richard Pyle, Chris Scharpf, Ingo Schindler, Erwin Schraml, Bill Smith-Vaniz, Henk Stam, Stefano Valdesalici, Emmanuel Vreven, Will White, Axel Zarske and Elsbeth Zwart.

Acknowledgments for Richard van der Laan (2017): Carlos Ardila Rodríguez, Carlos DoNascimiento, Stefan Koerber, Paul van der Laan, Anton Lamboj, Timo Moritz, Larry Page, Chris Scharpf, Erwin Schraml and Elsbeth Zwart.

Bill Eschmeyer
California Academy of Sciences

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