Institute for Biodiversity Science and Sustainability
Online Version, Updated 2 March 2018


Catalog of Fishes
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Catalog of Fishes - version of 2 March 2018 (Continuously updated since the early 1980s, around the turn of the month.)

Richard van der Laan is now a co-author of the Catalog of Fishes. This becomes Eschmeyer, W. N., R. Fricke, and R. van der Laan.

Other projects include Fricke (early references, new references, and corrections).

In this edition, we provide 77 new species in 2018. In 2017 there were 463 new species described.

Bill Eschmeyer, Florida Museum of Natural History, 1659 Museum Road, Gainesville, FL 32611 USA and California Academy of Sciences.

We are making additions to:

Van der Laan, R., W. N. Eschmeyer & R. Fricke (2014) (11 Nov.), Family-group names of Recent fishes. Zootaxa Monograph 3882 (1), 1-230. DOI 10.11646/zootaxa.3882.1.1

Download the addenda to the family-group list as a PDF.

See Family Group Names for future updates by R. van der Laan.

By default, search results will not include item 3 names -- unavailable names that detract from the main list of available names. These are mostly names mentioned in synonymy, names in lists, etc. They are still in the database, so if you want to see them, check the box labeled Include unavailable names before searching.

Citing the Catalog of Fishes

Eschmeyer, W. N. and R. Fricke, and R. van der Laan (eds).  CATALOG OF FISHES: GENERA, SPECIES, REFERENCES. ( Electronic version accessed dd mmm 2017. [This version was edited by Bill Eschmeyer.]

Fricke, R. (ed).  REFERENCES. ( Electronic version accessed dd mmm 2017. [Ron Fricke devoted much time to improving references and journals, many of which had not been examined in 15 to 25 years. Please remember that the title of a paper often differs in three places: the title on the paper itself, the contents listed by the journal, and in the PDF provided by the journal. Ron tends to follow the title on the pdf for recent papers and the title on the original for earlier publications. Many of our dates of publication are documented for priority purposes. Some current journals have a target date, but actual publication is often delayed. For nomenclatural date problems, we will help if we can.]

van der Laan, R., W. N. Eschmeyer and R. Fricke.  FAMILY-GROUP NAMES. ( Electronic version accessed dd mmm 2017.

Eschmeyer, W. N. and R. Fricke, and R. van der Laan (eds).  CATALOG OF FISHES: CLASSIFICATION. ( Electronic version accessed dd mmm 2017.

Fricke, R. & Eschmeyer, W. N.  JOURNALS. ( Electronic version accessed dd mmm 2017. [Includes all journals appearing in the Catalog, including publication information and ISSN numbers.]

Fricke, R. & Eschmeyer, W. N.  GUIDE TO FISH COLLECTIONS. ( Electronic version accessed dd mmm 2017. [Arranged by museum abbreviation and by country, includes type catalogs and historical publications and www sites where available.]

Eschmeyer, W. N. & Fong, J. D.  SPECIES BY FAMILY/SUBFAMILY. ( Electronic version accessed dd mmm 2017. [Recalculated with each new version; based on current literature, this provides all available species names, valid species, and species described in the last 10 years by family/subfamily.]

Please remember that if you got to the Catalog of Fishes from Fishbase, you are NOT in Fishbase. Fishbase is a more comprehensive database for a different audience, but they use Catalog of Fishes information in their authority file.

Some other useful sites for fishes are (1) Fishbase [Fishbase team, Manila]; (2) -- A comprehensive database on Shorefishes of the Tropical Eastern Pacific (and of the eastern Atlantic) [Ross Robertson]; (3) -- A database on elasmobranchs, with over 13,000 references, information on taxa, type specimens, etc. [Jürgen Pollerspöck]; (4) -- a comprehensive relational database of more than 99,350 scientific species name combinations and over 34,300 well-identified (mostly marine) fish pictures [Dennis Polack]; (5) Freshwater Fish List, 23rd edition December 2017; (6) ETYFish Project (derivations of Fish Names) by Christopher Scharpf; and (7) BioStor, for Biodiversity Heritage Library references. A very useful site is [not associated with Fishbase or Worldfish]. Maintained by Michal Mikšik and Erwin Schraml. They post new fish taxa and references as they are published.


A summary paper was published in 2010 from the Catalog of Fishes database that might be of interest to users of the Catalog. "Marine fish diversity: history of knowledge and discovery (Pisces)." By William N. Eschmeyer, Ronald Fricke, Jon D. Fong & Dennis A. Polack. Zootaxa, no. 2525:19-50 (2 July 2010, open access, click title to download).

As always, notifying us of missed original descriptions and errors in the database or providing PDFs of recent papers is most welcome.

Bill Eschmeyer edited type localities and standardized many place names. Our style is to use current place names found on an English map (or from Wikipedia and Google sites). Should you need original localities, they can be found in the original descriptions or from particular museum holdings (often online).

Many scientific names of fish species are based on personal names; some were found to be incorrect. For example, Jordan and co-workers spelled Japanese names ending in "a" with a terminal "e" as in matsubarae; that is acceptable, with the name Latinized first. So those names do not end in "i". Remarks are provided by Eschmeyer in this version for most species based on personal names. For a review of this very technical subject, see Dubois 2007: Zootaxa No. 1550:49-68.

New treatment of synonyms: Often an author revises a genus and treats a species in detail but does not mention the species synonyms (if any). If he/she does not mention the synonyms (rather common in recent treatments) and moves the valid “parent” species to a different genus, do we move the synonyms automatically, although the revising author may not have studied or assessed the synonyms? In the past, we did not move the synonyms. Typically, this often resulted in the synonyms being in a different current genus than the valid parent species. We now move the synonyms to the genus of the valid parent species, but no “status reference” showing the new generic placement is found with the synonym. Some synonyms will be moved incorrectly, but it is considered more desirable to have the potential synonyms in the same current genus as the valid parent species.

Some conventions used in the Catalog

Author plus date and a species code: The code of Zoological Nomenclature does not require a comma between the author and date, only a suggestion. In fact, we consider that confusing as that is traditionally used to show a cited reference in journal publications and not a species authorship and date. Genus+species+author+date (with parentheses as needed) is a (nearly) unique code or formula to define every species [2 known exceptions]. So you will find no comma between the author and date. Some journals are now treating species in this way.

Figures: Under a new species heading, some journals list (a) only the major figure, some list (b) all figures (including maps, habitats and graphs), and (c) some give figures showing the full view and any illustrated parts of the species. We give all figures that show a full view and any anatomical part of the species, so our listing of figures often differs from the treatment by a journal in the original description (as listed in the species heading).

Type localities: We give a current place name that one will find on a Google search or in an ‘English’ atlas. For some species described many years ago, one may find both the original locality and in parentheses a modern name for that locality. Many collections now have their holdings on-line, and it is there one can usually find the actual original locality if needed. Current localities are often augmented by additional information, such as coordinates.

Authors: We have added initials in brackets to distinguish all individual authors of genera and species.

Other sections

Species of Fishes by Family/Subfamily -- Total valid taxa reflect new taxa added minus taxa moved into synonymy plus taxa newly moved to valid. The classification is as used in the Catalog of Fishes. Prepared by Bill Eschmeyer and Jon Fong, with modifications by Richard van der Laan.

Family-Group Names -- Priority applies to Family-group names. It did not in early years. As classifications are refined with use of more subfamilies and tribes, it is required to use the oldest available family-group name for these new taxa.

Ichthyological Collections -- Presented are those specimen collections mentioned in the main Catalog of Fishes database, especially as repositories of type specimens. Besides museum names and abbreviations, this includes WWW sites, cross-referencing of abbreviations, and selection of museums by country, along with type catalogs, historical articles, name changes, transfers, and other items for individual collections. Prepared by Ron Fricke and Bill Eschmeyer.

Ichthyological Journals -- There are nearly 2600 individual journals or monograph series in the Catalog of Fishes database. We provide full citations, and other information such as changes in titles, published duration, ISSN numbers, WWW sites, etc. Prepared by Ron Fricke and Bill Eschmeyer.


I am fortunate to have Sherry Bostick as a technician who enters new species and other information under my direction; she is an excellent typist, find errors, and I look forward to her visits.

Acknowledgments for Bill Eschmeyer (2018): Nicolas Alvarez-Pliego, Jonathan Armbruster, Nicolas Bailly-coff, José Milton Barbosa, David A. Boyd, Ralf Britz, Dave Catania, Patrick Ciccotto, Kevin Conway, Matthew Craig, Carlos Donascimiento, Benjamin Frable, Uiblein Franz, Tom Fraser, Martin Gomon, P. Alexander Hulley, Howard Jelks, Thomas Litz, Alejandro Londoño-Burbano, Mark McGrouther, Paulo Lucinda, John Lundberg, Michal Mikšík, Hiroyuki Motomura, Makoto Okamoto, Michael Oliver, Felipe Ottoni, Larry M. Page, Arturo Acero Pizarro, Dennis Polack, Stuart Poss, Roberto E. Reis, Ross Robertson, Rob Robins, Christopher Scharpf, David G. Smith, Maria Laura de Souza Delapieve, John Sullivan, William White, Richard Winterbottom and Dan Zimberlin.

Acknowledgments for Bill Eschmeyer (2017): K. V. Akhilesh, Gerry Allen, Mark Allen, Armando J. Almeida, Nisreen Alwan, William D. Anderson Jr, Carlos A. Ardila Rodriguez, Nicolas Bailly, Ramiro E. Barriga Salazar, Tim Berra, Roger Bills, Gert Boden, Philippe Borsa, David Boyd, Matt Broadhead, George Burgess, Bárbara Calegari, Miguel Caneiro, Fernando Carvalho, Dave Catania, Albert Chakona, Patrick Ciccotto, Kevin Conway, Vahe Demirjian, Rafael Bañón Díaz, Fabio Di Dario, Carlos Donascimiento, Jean-Dominique Durand, Yury Dyldin, Dave Ebert, Marco Endruweit, Andre Esguicero, Carl Ferraris, Adania Flemming, Ralph Foster, Benjamin Frable, Libert Francois, Tom Fraser, Isabel Gandarilla, Hugo Gante, Michel Donato Gianeti, Carter Gilbert, Martin F. Gomon, David Greenfield, Renny Kurnia Hadiaty, Karsten Hartel, Elaine Heemstra, Hans Ho, Doug Hoese, Gabriela Hogue, Erling Holm, Hisashi Imamura, Tomio Iwamoto, Laith Jawad, Howard Jelks, Dave Johnson, Max Kasparek, Seishi Kimura, Stefan Koerber, Keita Koeda, Muddu Krishna, Richard van der Laan, Kuldeep K. Lal, Helen Larson, Peter Last, Sébastien Lavoué, Jeff Leis, Thomas Litz, Hugo López, Zilda Margarete Seixas de Lucena, John Lundberg, Nathan Lujan, James Maclaine, Javier Maldonado, Manoela Marinho, Douglas Markle, Eddie Matheson, George Mendes T. Mattox, Mizuki Matsunuma, Catherine W. Mecklenburg, Filipe Augusto Gonçalves de Melo, Marcelo Roberto Souto de Melo, Michael Mincarone, Marcos Mirande, Hiroyuki Motomura, Bruce Mundy, Nozomu Muto, Naohide Nakayama, Gavin Naylor, Andre Luiz Netto-Ferreira, Dirk Neumann, Leo Nico, Jørgen Nielsen, Huu Duc Nguyen, Makoto Okamoto, Michael Oliver, Larry Page, Paolo Parenti, Keith Phillip, John Pogonoski, Dennis Polack, Riley Pollom, Stuart Poss, Artem Prokofiev, Graham Proudlove, John Randall, Lucia Rapp Py-Daniel, Roberto E. Reis, Tyson Roberts, Rob Robins, Uwe Römer, Juan Jose Rosso, Mark Sabaj Perez, Norma Salcedo, Frank Schäfer, Christopher Scharpf, Ulrich Schliewen, Erwin Schraml, Werner Schwarzhans, Randal Singer, David G. Smith, Bill Smith-Vaniz, Matthias Stehmann, Nicolas Straube, John Sullivan, Milton Tan, Don Taphorn, Sandrine Tercerie, Alfred Thomson, Luke Tornabene, Tom Trnski, Denis Tweddle, Franz Uiblein, James Anyelo Vanegas-Rios, Benjamin Victor, Chelsea Walery, Simon Weigmann, David De Weirdt, William White, Jeff Williams, Richard Winterbottom, Jouke van der Zee, Mikhail Zhukov and Dan Zimberlin. Plus any others we may have missed.

Acknowledgments for Ron Fricke (2018): Sylvain Adnet, Gastón Aguilera, Gerry Allen, Dmitry Astakhov, Arcady Balushkin, Esra Bayςelebi, Ronaldo Borges Barthem, David Bellwood, Nickolai Bochkarev, Guamuchirai Chakona, Natalia Chernova, Todd Clardy, Fenton Cotterill, Laura Donin, Guy Duhamel, Guilherme Dutra, Yury Dyldin, Anthony Echelle, Marco Endruweit, Deniz Ergüden, Brit Finucci, Nelson Ferreira Fontoura, Ofer Gon, Renny Hadiaty, Elisabeth Henschel, Douglass F. Hoese, A. K. Jaiswar, Dušan Jelić, Misel Jelić, Hichem Kara, Subodha Kumar Karna, Axel Katz, Cüneyt Kaya, Philippe Keith, Keita Koeda, Yoshinori Kumazawa, Boyd Kynard, Kosygin Laishram, Kuldeep Lal, Lalramliana, Pablo Lehmann, Jeff Leis, Li Chenhong, Alejandro Londoño-Burbano, Luiz Malabarba, Fatih Mangιt, Alexandre Marceniuk, Rene Martin, Michael Mincarone, Meera Mohandas, Vindhya Mohindra, Timo Moritz, Takahiko Mukai, Béla Nagy, Heok Hee Ng, Larry Page, Antoine Pariselle, Lubomír Piálek, Artem Prokofiev, Francisco Provenzano, Roberto Reis, R. K. Renjith, Baruch Rinkevich, Lukas Rüber, Takashi Satoh, Ivan Sazima, Christopher Scharpf, Nalani Schnell, Hiroshi Senou, Esmaile Shakman, Valentina Sideleva, Raimundo da Silva, Thiago Silva Loboda, Paul Skelton, Donald Taphorn, Luke Tornabene, Katerina Vasil'eva, Waikhom Vishwanath, Emmanuel Vrewen, Mark Wilson, Peter Wirtz.

Acknowledgments for Ron Fricke (2017): Hugo Aguirre, Windsor Aguirre, K. V. Akhilesh, Malek Ali, Armando Almeida, Felipe Alonso, William Anderson Jr., Levent Artüz, Muthukumarasamy Arunachalam, Carole Baldwin, Michel Bariche, Mathias Barrett, Sudesh Batuwita, Philippe Béarez, Tim Berra, Ricardo Betancur-R, Gökçen Bilge, Flávio Bockmann, Sergey Bogorodsky, Philippe Borsa, Pedro Bragança, Jack Briggs, Gregor Cailliet, Bárbara Calegari, Henri Cappetta, Fernando Carvalho, Alfredo Carvalho-Filho, Marie-Noëlle de Casamajor, Wei-Jen Chen, Xiaojong Chen, Natalia Chernova, Cem Dalyan, Arieli Cherobim, Patrick Ciccotto, Kevin Conway, Diushi Corona-Santiago, Wilson Costa, Giovanni Delmastro, Gaël Denys, David de Weirdt, Fabio Di Dario, Andrey Dolgov, Jakov Dulčić, Yury Dyldin, Soheil Eagderi, Marco Endruweit, Semih Engin, Deniz Ergüden, Hamid Esmaeili, Jörg Freyhof, Daniel Golani, Vladimir Golovanov, Weferson Júnio da Graςa, Dave Greenfield, İskender Gülle, Lubomir Hanel, Ian Harrison, Elaine Heemstra, Muhammad Iqbal, Robert Irving, Isaäc Isbrücker, Fernando Jerep, Dave Johnson, Pepijn Kamminga, Karuppiah Kannan, Unmesh Katwate, Toshio Kawai, Cüneyt Kaya, Stefan Koerber, Alexander Kotlyar, Maurice Kottelat, Kuldeep Lal, Sébastien Lavoué, Ezequiel Mabragaña, Alexandre Marceniuk, Mojtaba Masoudi, Anil Mohapatra, Muddula Krishna Naranji, Thomas Litz, Carlos de Lucena, Paulo Lucinda, Leonid Manilo, Tatiane Mariguela, Mizuki Matsunuma, Bruno Melo, Rico Morgenstern, Bruce Mundy, Nozomu Muto, Mikhail Nazarkin, Thomas Near, Kongbrailatpam Nebeshwar, Heok Hee Ng, Jørgen Nielsen, Willian Ohara, Makoto Okamoto, Larry Page, Carla Pavanelli, Edson Pereira, Khalid Pervaiz, Mathews Plamoottil, John Pogonoski, Andrea Polanco, Gianluca Polgar, Jan Poulsen, Artem Prokofiev, Richard Pyle, Jack Randall, Bettina Reichenbacher, Diana Duarte Rodrigues, Uwe Römer, Barry Russell, Etsuro Sawai, Christopher Scharpf, Ingo Schindler, Juan Schmitter-Soto, Erwin Schraml, Sally Schramm, Werner Schwarzhans, Ying Giat Seah, Iraj Segherloo, Hiroshi Senou, Malcolm Smale, William Smith-Vaniz, Melanie Stiassny, John Sullivan, K. S. Sumod, Ana Swarça, Luke Tornabene, Armi Torres, Davut Turan, Anyelo Vanegas Rios, Ekaterina Vasil'eva, Ernesto Velázquez-Velázquez, Sarah Viana, Francisco Villa-Navarro, Waikhom Vishwanath, Matheus Volcan, Elena Voronina, Olga Voskoboinikova, Emmanuel Vreven, Stephen Walsh, Wolmar Wosiacki, Yusuke Yamanoue, Akane Yoshikawa, Hyo Jae Yu, Claúdio Zawadzki.

Acknowledgments for Richard van der Laan (2018): Gastón Aguilera, Nicolás Álvarez-Pliego, Carlos Ardila Rodríguez, David Boyd, Patrick Ciccotto, André Esguícero, Jean Huber, Axel Katz, Stefan Koerber, Paul van der Laan, Thomas Litz, Michal Mikšík, Larry Page, Chris Scharpf, Erwin Schraml, John Sullivan and Elsbeth Zwart.

Acknowledgments for Richard van der Laan (2017): Nisreen Alwan, Carlos Ardila Rodríguez, Nicolas Bailly, Simmy Bezeng, Gert Boden, Fernando Carvalho, Patrick Ciccotto, Carlos DoNascimiento, Wolfgang Eberl, André Esguícero, Jean Huber, Daemin Kim, Stefan Koerber, Paul van der Laan, Kuldeep Lal, Anton Lamboj, Thomas Litz, Maisnam Manorama, Juan Marcos Mirande, Timo Moritz, Larry Page, Murillo Pastana, Roberto Reis, Chris Scharpf, Ingo Schindler, Erwin Schraml, Jos Snoeks, Dan Zimberlin and Elsbeth Zwart.

Bill Eschmeyer
California Academy of Sciences

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