Institute for Biodiversity Science and Sustainability
Online Version, Updated 2 March 2020

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Eschmeyer's Catalog of Fishes
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areolata, Perca Forsskål [P. S.] 1775:xi, 42 [Descriptiones animalium (Forsskål); ref. 1351] Jeddah, Saudi Arabia, Red Sea. Possible holotype: ZMUC P43570 (dry skin). Type catalog: Nielsen 1974:61 [ref. 9588] not definitely referred ZMUC P43570 as holotype due to confusion of some labels in Forsskål's collection. As Perca summana var. areolata on p. xi, and as "[PERCA summana] (c) Perca areolata" on p. 42. Possible type figured in Klausewitz & Nielsen 1965: Pl. 10 [ref. 20847]. •Valid as Epinephelus areolatus (Forsskål 1775) -- (Kyushin et al. 1977:208 [ref. 19756], Kyushin et al. 1982:175 [ref. 19754], Dor 1984:98 [ref. 29757], Katayama in Masuda et al. 1984:128 [ref. 6441], Heemstra & Randall 1986:523 [ref. 5667], Allen & Swainston 1988:56 [ref. 25477], Paxton et al. 1989:493 [ref. 12442], Lee 1990:36 [ref. 19830], Randall et al. 1990:105 [ref. 15987], Randall & Heemstra 1991:99 [ref. 19601], Baranes & Golani 1993:306 [ref. 22372], Heemstra & Randall 1993:112 [ref. 21042], Goren & Dor 1994:25 [ref. 25356], Randall 1995:128 [ref. 22896], Evseenko 1996:727 [ref. 24666], Mohsin & Ambak 1996:233 [ref. 27969], Allen 1997:84 [ref. 23977], Carpenter et al. 1997:143 [ref. 23922], Chen et al. 1997:45 [ref. 26476], Kuiter 1997:110 [ref. 25488], Randall et al. 1997:105 [ref. 25919], Heemstra & Randall 1999:2487 [ref. 24815], Nakabo 2000:715 [ref. 25086], Randall in Randall & Lim 2000:609 [ref. 25122], Laboute & Grandperrin 2000:181 [ref. 25191], Sadovy & Cornish 2000:80 [ref. 25480], Allen 2000:98 [ref. 25868], Hutchins 2001:30 [ref. 25847], Nakabo 2002:715 [ref. 26001], Allen & Adrim 2003:31 [ref. 26830], Manilo & Bogorodsky 2003:S104 [ref. 27377], Kimura et al. in Kimura & Matsuura 2003:51 [ref. 27510], Randall et al. 2004:11 [ref. 27624], Adrim et al. 2004:119 [ref. 27715], Randall 2005:154 [ref. 28239], Hoese et al. 2006:1008 [ref. 28999], Satapoomin in Motomura & Satapoomin 2009:86 [ref. 30492], Motomura et al. 2010:96 [ref. 31256], Fricke et al. 2011:384 [ref. 31242], Craig et al. 2011:85 [ref. 31786], Allen & Erdmann 2012:267 [ref. 31980], Randall et al. 2013:524 [ref. 32685], Fricke et al. 2013:256 [ref. 32706], Larson et al. 2013:99 [ref. 32988], Yoshida et al. 2013:99 [ref. 34464], Psomadakis et al. 2015:196 [ref. 34104], Rothman et al. 2016:479 [ref. 34171], Motomura et al. 2017:84 [ref. 35490], Ali et al. 2018:318 [ref. 36183], Bogorodsky & Randall 2018:[20] 258 [ref. 36492], Fricke et al. 2018:147 [ref. 35805], Jawad et al. 2018:103 [ref. 36262], Golani & Fricke 2018:62 [ref. 36273], Ma & Craig 2018:455 [ref. 36100], Satapoomin in Kimura et al. 2018:100 [ref. 36460], Eagderi et al. 2019:106 [ref. 37020], Fricke et al. 2019:116 [ref. 36673], Zajonz et al. 2019:70 [ref. 36871], Psomadakis et al. 2020:387 [ref. 37272], Ray & Mohapatra 2020:110 [ref. 37300]). Current status: Valid as Epinephelus areolatus (Forsskål 1775). Serranidae: Epinephelinae. Distribution: Red Sea, Indo-West Pacific: East Africa, Socotra, Persian Gulf and Madagascar east to Fiji, Tonga and Samoa, north to southern Japan, south to Western Australia and Queensland (Australia) and New Caledonia. Habitat: marine.