Institute for Biodiversity Science and Sustainability
Online Version, Updated 6 November 2023

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Catalog of Fishes

Species that contain: Upeneus and tragula [ 6 ] records

alessandrini, Megalepis Bianconi [G. G.] 1857:100 [Rendiconto delle Sessioni dell'Accademia delle Scienze dell'Instituto dei Bologna 1856-1857; ref. 17950] Mozambique. No types known. Also appeared as new in Bianconi 1854:247, Piscium pl. 4 [ref. 295] and Bianconi 1857:453 [ref. 26453] with figure; earliest not researched. •Synonym of Upeneus tragula Richardson 1846 -- (Kottelat 2013:362 [ref. 32989]). Current status: Synonym of Upeneus tragula Richardson 1846. Mullidae. Habitat: brackish, marine.

caudalis, Upeneus sundaicus var. Popta [C. M. L.] 1921:206 [Zoologische Mededeelingen (Leiden) v. 6 (art. 15); ref. 15325] Tiworo, Muna, Sunda Islands, Indonesia. Holotype (unique): SMF 6601. •Synonym of Upeneus tragula Richardson 1846 -- (Kottelat 2013:362 [ref. 32989], Miesen et al. 2016:90 [ref. 34492]). Current status: Synonym of Upeneus tragula Richardson 1846. Mullidae. Habitat: brackish, marine.

niebuhri, Upeneus Guézé [P.] 1976:596 [Revue des Travaux de l'Institut des Pêches Maritimes v. 40 (no. 3-4); ref. 7337] Gulf of Suez, 28°54'29"N, 32°29' to 32°45'E, Red Sea, depth 62 meters. Holotype (unique): MNHN 1977-0174. Type catalog: Bauchot et al. 1985:8 [ref. 20735]. •Synonym of Upeneus tragula Richardson 1846 -- (Ben-Tuvia & Kissil 1988:13 [ref. 12819]). •Valid as Upeneus niebuhri (Guézé 1976) -- (Kottelat 2013:362 [ref. 32989], Uiblein & Gouws 2014:668 [ref. 33216], Bogorodsky & Randall 2018:[5] 243 [ref. 36492], Golani & Fricke 2018:104 [ref. 36273], Uiblein et al. 2020:550 [ref. 37717], Echreshavi et al. 2022:20 [ref. 39234], Uiblein & Randall 2022:371 [ref. 39855]). Current status: Valid as Upeneus niebuhri Guézé 1976. Mullidae. Distribution: Northern Red Sea endemic. Habitat: marine.

oligospilus, Upeneus Lachner [E. A.] 1954:525, Pl. 14 (fig. D) [Proceedings of the United States National Museum v. 103 (no. 3330); ref. 10031] Tarut Bay, Persian Gulf. Holotype: USNM 153988. Paratypes: USNM 147995 (11). •Synonym of Upeneus tragula Richardson 1846 -- (Randall 1995:243 [ref. 22896] or at best a subspecies, Carpenter et al. 1997:193 [ref. 23922], Randall 2001:3199 [ref. 26109], Kim & Nakaya 2002:130 [ref. 26150], Randall & Kulbicki 2006:305 [ref. 28802], Kottelat 2013:362 [ref. 32989]). •Valid as Upeneus oligospilus Lachner 1954 -- (Uiblein & Heemstra 2010:48 [ref. 30763], Uiblein & Heemstra 2011:587 [ref. 31346], Uiblein & Heemstra 2011:639 [ref. 31501], Uiblein & McGrouther 2012:63 [ref. 32282], Uiblein & Gouws 2014:670 [ref. 33216], Jawad et al. 2018:116 [ref. 36262], Eagderi et al. 2019:88 [ref. 37020], Uiblein et al. 2020:550 [ref. 37717], Echreshavi et al. 2022:20 [ref. 39234], Uiblein & Randall 2022:371 [ref. 39855]). Current status: Valid as Upeneus oligospilus Lachner 1954. Mullidae. Distribution: Northwestern Indian Ocean: Persian Gulf endemic. Habitat: brackish, marine.

tragula, Upeneus Richardson [J.] 1846:220 [Report of the British Association for the Advancement of Science 15th meeting [1845]; ref. 3742] Canton [Guangzhou], China. Syntypes: (3) BMNH 1968.3.11.12 (1, left half skin). Type information: Whitehead 1970:214 [ref. 11606]. The latin word tragula (= spear, javelin) is a noun and does not decline; the spelling is correct as tragula, regardless of generic assignment. •Valid as Upeneus tragula Richardson 1846 -- (Lachner 1954:522 [ref. 10031], Lachner 1960:4 [ref. 23973], Yamakawa in Masuda et al. 1984:163 [ref. 6441], Ben-Tuvia 1986:613 [ref. 5641], Ben-Tuvia & Kissil 1988:13 [ref. 12819], Allen & Swainston 1988:90 [ref. 25477], Winterbottom et al. 1989:41 [ref. 13251] as tragulus, Randall et al. 1990:212 [ref. 15987], Suzuki & Uno 1992:3 [ref. 24139], Kuiter 1993:202 [ref. 23929], Goren & Dor 1994:45 [ref. 25356], Randall 1995:243 [ref. 22896], Mohsin & Ambak 1996:387 [ref. 27969], Larson & Williams 1997:361 [ref. 23967], Allen 1997:142 [ref. 23977], Carpenter et al. 1997:193 [ref. 23922], Chen et al. 1997:109 [ref. 26476], Kuiter 1997:194 [ref. 25488], Randall et al. 1997:212 [ref. 25919], Myers 1999:161 [ref. 23965], Mishra et al. 1999:87 [ref. 25315], Johnson 1999:738 [ref. 25471], Allen 2000:105 [ref. 25868], Randall in Randall & Lim 2000:622 [ref. 25122], Nakabo 2000:872 [ref. 25182], Laboute & Grandperrin 2000:285 [ref. 25191], Sadovy & Cornish 2000:150 [ref. 25480], Iwatsuki et al. 2000:102 [ref. 26368], Matsuura & Peristiwady in Matsuura & Peristiwady 2000:265 [ref. 27511], Golani 2001:111 [ref. 25530], Sakai et al. 2001:100 [ref. 25693], Shimizu 2001:31 [ref. 25799], Hutchins 2001:264 [ref. 25841], Hutchins 2001:35 [ref. 25847], Randall 2001:3199 [ref. 26109], Kim & Nakaya 2002:130 [ref. 26150], Nakabo 2002:872 [ref. 26193], Allen & Adrim 2003:41 [ref. 26830], Manilo & Bogorodsky 2003:S111 [ref. 27377], Shibukawa et al. in Kimura & Matsuura 2003:113 [ref. 27510], Adrim et al. 2004:122 [ref. 27715], Heemstra et al. 2004:3322 [ref. 27945], Randall 2005:300 [ref. 28239], Randall & Kulbicki 2006:305 [ref. 28802], Hoese & Bray 2006:1273 [ref. 29000], Gomon 2008:591 [ref. 30616], Satapoomin in Imamura et al. 2009:176 [ref. 31157], Uiblein & Heemstra 2010:52 [ref. 30763], Motomura et al. 2010:130 [ref. 31256], Uiblein & Heemstra 2011:587 [ref. 31346], Uiblein & Heemstra 2011:639 [ref. 31501], Yamashita et al. 2011:54 [ref. 31639], Allen & Erdmann 2012:510 [ref. 31980], Uiblein & McGrouther 2012:63 [ref. 32282], Larson et al. 2013:152 [ref. 32988], Kottelat 2013:361 [ref. 32989], Yoshida et al. 2013:159 [ref. 34464], Uiblein & Gouws 2014:670 [ref. 33216], Bos 2014:757 [ref. 33588], Fricke et al. 2014:96 [ref. 33932], Uiblein & Gouws 2015:[18] 1038 [ref. 33992], Satapoomin in Kimura et al. 2015:66 [ref. 34319], Uiblein & White 2015:52 [ref. 33856], Miesen et al. 2016:90 [ref. 34492], Motomura et al. 2017:165 [ref. 35490], Ali et al. 2018:335 [ref. 36183], Gökoglu 2018:57 [ref. 37179], Golani & Fricke 2018:104 [ref. 36273], Satapoomin in Kimura et al. 2018:194 [ref. 36460], Eagderi et al. 2019:89 [ref. 37020], Fricke et al. 2019:176 [ref. 36673], Hoschke et al. 2019:156 [ref. 36626], Uiblein et al. 2019:192 [ref. 37008], Habib & Islam 2020:Supplementary table p. 20 [ref. 38321], Kim et al. 2020:139 [ref. 39118], Psomadakis et al. 2020:506 [ref. 37272], Sonoyama et al. 2020:77 [ref. 37637], Ali in Motomura et al. 2021:37 [ref. 38121], Echreshavi et al. 2022:23 [ref. 39234], Farhana-Azmi et al. 2022:9 [ref. 40448], Vilasri et al. 2023:90 [ref. 40764], Vishnupriya & Nair 2023:119 [ref. 40508]). Current status: Valid as Upeneus tragula Richardson 1846. Mullidae. Distribution: Indo-West Pacific: East Africa and Persian Gulf east to Philippines, New Ireland (Papua New Guinea) and Solomon Islands, north to southern Sea of Japan (Korea, Japan), south to Rottnest Island (Western Australia), Sydney (New South Wales, Australia) and New Caledonia. Former Red Sea records are now assigned to U. niebuhri Guézé 1976. Habitat: brackish, marine.

variegatus, Upeneoides Bleeker [P.] 1849:64 [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 22 [13]; ref. 318] Kagoshima, Japan; Jakarta, Java, Indonesia. Lectotype: RMNH 5720. Paralectotypes: RMNH 25408 or 25498 (some out of 11), SMNS 10617 (2). Type catalog: Fricke 1991:17 [ref. 13474], Fricke 2005:49 [ref. 29864] as 25498. Originally as Upeneoïdes. •Synonym of Upeneus tragula Richardson 1846 -- (Lachner 1954:522 [ref. 10031], Randall & Kulbicki 2006:305 [ref. 28802], Kottelat 2013:362 [ref. 32989]). Current status: Synonym of Upeneus tragula Richardson 1846. Mullidae. Habitat: brackish, marine.


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