Institute for Biodiversity Science and Sustainability
Online Version, Updated 6 November 2023

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Catalog of Fishes

Species that contain: Variola and louti [ 8 ] records

cernipedis, Serranus Miranda Ribeiro [A. de] 1913:Serranidae p. 31 [Arquivos do Museu Nacional de Rio de Janeiro v. 17; ref. 3711] No locality. No types known. On p. 261 when continuously paginated. •Questionably a synonym of Variola louti (Forsskål 1775) -- (Randall & Heemstra 1991:305 [ref. 19601], Heemstra & Randall 1993:304 [ref. 21042], Parenti & Randall 2020:94 [ref. 37354]). •Questionably a synonym of Variola louti (Fabricius 1775). Current status: Uncertain as Variola louti (Fabricius 1775). Epinephelidae. Habitat: marine.

flavimarginatus, Serranus Rüppell [W. P. E. S.] 1830:109 [Atlas zu der Reise im nördlichen Africa. Fische des Rothen Meeres; ref. 3843] Al Muwaylih, Tabuk Province, Saudi Arabia, Red Sea. Holotype (unique): SMF (lost). Non-type: SMF 544 (1). •Synonym of Variola louti (Forsskål 1775) -- (Dor 1984:103 [ref. 29757], Randall & Heemstra 1991:305 [ref. 19601], Heemstra & Randall 1993:304 [ref. 21042], Fricke et al. 2018:158 [ref. 35805], Parenti & Randall 2020:94 [ref. 37354]). •Synonym of Variola louti (Fabricius 1775) -- (Heemstra 2022:78 [ref. 39742]). Current status: Synonym of Variola louti (Fabricius 1775). Epinephelidae. Habitat: marine.

longipinna, Serranus (Variola) Swainson [W.] 1839:202 [The natural history and classification v. 2; ref. 4303] Red Sea. No types known. Based on Serranus louti of Rüppell 1830: Pl. 26 (fig. 2) [ref. 3843], but see Serranus phaenistomus Swainson 1839. •Synonym of Variola louti (Forsskål 1775) -- (Heemstra & Randall 1993:304 [ref. 21042], Hoese et al. 2006:1024 [ref. 28999], Parenti & Randall 2020:94 [ref. 37354]). •Synonym of Variola louti (Fabricius 1775) -- (Heemstra 2022:78 [ref. 39742]). Current status: Synonym of Variola louti (Fabricius 1775). Epinephelidae. Habitat: marine.

louti, Perca Fabricius [J. C.] in Niebuhr (ex Forsskål) 1775:40, xi [Descriptiones animalium (Forsskål); ref. 1351] Jeddah, Saudi Arabia, Red Sea; Al-Luhayya, Yemen, Red Sea. Syntypes: ZMUC P43566 (1, dry skin). Type catalog: Nielsen 1974:61 [ref. 9588]; Fricke 2008:30 [ref. 30182]. Spelled luti by Valenciennes in Cuvier & Valenciennes 1828 [ref. 997]. Surviving syntype figured in Klausewitz & Nielsen 1965: Pl. 9 [ref. 20847]. Authorship according to Fricke 2008:30 [ref. 30182]. •Valid as Variola louti (Forsskål 1775) -- (Kyushin et al. 1977:178 [ref. 19756], Kyushin et al. 1982:156 [ref. 19754], Dor 1984:103 [ref. 29757], Katayama in Masuda et al. 1984:126 [ref. 6441], Heemstra & Randall 1986:536 [ref. 5667], Allen & Swainston 1988:58 [ref. 25477], Paxton et al. 1989:501 [ref. 12442], Winterbottom et al. 1989:27 [ref. 13251], Lee 1990:63 [ref. 19830], Randall et al. 1990:120 [ref. 15987], Francis 1993:160 [ref. 25479], Francis & Randall 1993:119 [ref. 20996], Heemstra & Randall 1993:304 [ref. 21042], Kuiter 1993:134 [ref. 23929], Goren & Dor 1994:27 [ref. 25356], Randall 1995:138 [ref. 22896], Mohsin & Ambak 1996:247 [ref. 27969], Allen 1997:86 [ref. 23977], Chen et al. 1997:41 [ref. 26476], Kuiter 1997:102 [ref. 25488], Randall et al. 1997:120 [ref. 25919], Myers 1999:117 [ref. 23965], Fricke 1999:209 [ref. 24106], Heemstra & Randall 1999:2539 [ref. 24815], Randall 1999:12 [ref. 25829], Allen 2000:87 [ref. 37268], Nakabo 2000:709 [ref. 25086], Randall in Randall & Lim 2000:611 [ref. 25122], Laboute & Grandperrin 2000:193 [ref. 25191], Randall & Earle 2000:12 [ref. 25806], Iwatsuki et al. 2000:98 [ref. 26368], Hutchins 2001:263 [ref. 25841], Hutchins 2001:30 [ref. 25847], Nakabo 2002:709 [ref. 26001], Allen & Adrim 2003:33 [ref. 26830], Manilo & Bogorodsky 2003:S105 [ref. 27377], Myers & Donaldson 2003:622 [ref. 27495], Kimura et al. in Kimura & Matsuura 2003:56 [ref. 27510], Lobel & Lobel 2004:70 [ref. 27576], Randall et al. 2004:12 [ref. 27624], Adrim et al. 2004:120 [ref. 27715], Heemstra et al. 2004:3319 [ref. 27945], Heemstra & Heemstra 2004:173 [ref. 28072], Randall 2005:167 [ref. 28239], Hoese et al. 2006:1024 [ref. 28999], Fricke et al. 2009:49 [ref. 30213], Motomura in Motomura & Satapoomin 2009:95 [ref. 30492], Motomura et al. 2010:101 [ref. 31256], Caratini & Soriano 2011:51 [ref. 39915], Fricke et al. 2011:387 [ref. 31242], Craig et al. 2011:354 [ref. 31786], Allen & Erdmann 2012:285 [ref. 31980], Fricke et al. 2014:66 [ref. 33932], Delrieu-Trottin et al. 2015:5 [ref. 33988], Koeda et al. 2016:29 [ref. 35333], Fricke et al. 2018:158 [ref. 35805], Golani & Fricke 2018:67 [ref. 36273], Ma & Craig 2018:456 [ref. 36100], Fricke et al. 2019:126 [ref. 36673], Zajonz et al. 2019:71 [ref. 36871], Parenti & Randall 2020:94 [ref. 37354], Akhilesh et al. 2021:56 [ref. 38701], Durville et al. 2021:311 [ref. 38802], Kim et al. 2021:132 [ref. 39117], Zajonz et al. 2022:94 [ref. 40522], Moazzam & Osmany 2023:690 [ref. 40722], Murase et al. 2023:39 [ref. 40935]). •Valid as Variola louti (Fabricius 1775) -- (Fricke 2008:29 [ref. 30182], Kovačić et al. 2021:78 [ref. 39014], Heemstra 2022:78 [ref. 39742]). Current status: Valid as Variola louti (Fabricius 1775). Epinephelidae. Distribution: Red Sea; Indo-West Pacific: South Africa, East Africa, Socotra (Yemen), Seychelles, Comoros, Madagascar and Mascarenes (La Réunion, Mauritius, Rodrigues) east to Pitcairn Group, north to southern Japan and Ogasawara Islands (Japan), south to Western Australia, Lord Howe Island (Australia) and Rapa (French Polynesia). Habitat: marine.

melanotaenia, Serranus Bleeker [P.] 1857:33 [Acta Societatis Regiae Scientiarum Indo-Neêrlandicae v. 2 (art. 7); ref. 356] Ambon Island, Molucca Islands, Indonesia. Holotype (unique): whereabouts unknown. •Synonym of Variola louti (Forsskål 1775) -- (Randall & Heemstra 1991:305 [ref. 19601], Heemstra & Randall 1993:304 [ref. 21042], Parenti & Randall 2020:94 [ref. 37354]). •Synonym of Variola louti (Fabricius 1775). Current status: Synonym of Variola louti (Fabricius 1775). Epinephelidae. Habitat: marine.

phaenistomus, Serranus Swainson [W.] 1839:201 [The natural history and classification v. 2; ref. 4303] No types known. Based on Rüppell 1830:Pl. 26 (fig. 2) [ref. 3843], but perhaps an error because Variola longipinns Swainson on the next page is based on the same figure. •Synonym of Variola louti (Forsskål 1775) -- (Randall & Heemstra 1991:305 [ref. 19601], Heemstra & Randall 1993:304 [ref. 21042], Parenti & Randall 2020:94 [ref. 37354]). •Synonym of Variola louti (Fabricius 1775) -- (Heemstra 2022:78 [ref. 39742]). Current status: Synonym of Variola louti (Fabricius 1775). Epinephelidae. Habitat: marine.

punctulatus, Labrus Lacepède [B. G. E.] (ex Commerson) 1801:431, 477, Pl. 17 (fig. 2) [Histoire naturelle des poissons (Lacepède) v. 3; ref. 2710] Indian Ocean. Lectotype: MNHN A-5724 (dry). Type catalog: Bauchot 1963:100 [ref. 19171], Bauchot et al. 1984:17-18 [ref. 19232]. Based on a Commerson figure. Lectotype selected by Bauchot et al. 1984:17 [ref. 19232]. •Synonym of Variola louti (Forsskål 1775) -- (Randall & Heemstra 1991:305 [ref. 19601], Heemstra & Randall 1993:304 [ref. 21042], Fricke 1999:210 [ref. 24106], Parenti & Randall 2020:94 [ref. 37354]). •Synonym of Variola louti (Fabricius 1775) -- (Heemstra 2022:78 [ref. 39742]). Current status: Synonym of Variola louti (Fabricius 1775). Epinephelidae. Habitat: marine.

roseus, Serranus Valenciennes [A.] in Cuvier & Valenciennes 1828:306 [Histoire naturelle des poissons v. 2; ref. 997] Tahiti, Society Islands. No types known. Based on a Parkinson drawing. •Synonym of Variola louti (Forsskål 1775) -- (Randall & Heemstra 1991:305 [ref. 19601], Parenti & Randall 2020:94 [ref. 37354]). •Synonym of Variola louti (Fabricius 1775) -- (Heemstra 2022:78 [ref. 39742]). Current status: Synonym of Variola louti (Fabricius 1775). Epinephelidae. Habitat: marine.


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