intermedius, Gasterosteus Girard [C. F.] 1856:135 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] Cape Flattery, Washington, U.S.A. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.