Eschmeyer's Catalog of Fishes Reference Record:
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Weiss, S. J., D. V. Gonçalves, G. Secci-Petretto, G. K. Englmaier, A. Gomes-Dos-Santos, G. P. J. Denys, H. Persat, A. Antonov, C. Hahn, E. B. Taylor and E. Froufe 2020 (25 Nov.) [ref. 37974] See ref. online
- Global systematic diversity, range distributions, conservation and taxonomic assessments of graylings (Teleostei: Salmonidae; Thymallus spp.). Organisms Diversity & Evolution v. 21: [1-18] 25-42. [First published online, pp. 1-18, on 25 Nov. 2020; volume number and final pagination added in 2021.]