vardarensis, Squalius cephalus Karaman [S.] 1928:156 [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 6; ref. 15903] Yugoslavia. Syntypes: whereabouts unknown. Also in Karaman 1929:172 [ref. 15904]. •Synonym of Leuciscus cephalus (Linnaeus 1758) -- (Kottelat 1997:66 [ref. 22952]). •Synonym of Squalius cephalus (Linnaeus 1758), but a valid subspecies vardarensis Karaman 1928 -- (Sanjur et al. 2003:23 [ref. 28742], Freyhof et al. 2006:418 [ref. 28484]). •Valid as Squalius vardarensis Karaman 1928 -- (Kottelat & Freyhof 2007:277 [ref. 29996], Bogutskaya & Zupančič 2010:53 [ref. 30905], Perea et al. 2010:14 [ref. 31802], Özuluğ & Freyhof 2011:108 [ref. 31360], Barbieri et al. 2015:67 [ref. 35029], Lorenzoni et al. 2019:252 [ref. 37108], Buj et al. 2019:[16] [ref. 37174]). Current status: Valid as Squalius vardarensis Karaman 1928. Leuciscidae: Leuciscinae. Distribution: Southeastern Europe: Greece and Macedonia. Introduced in Italy. Habitat: freshwater.