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Eschmeyer's Catalog of Fishes
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Species in the genus Icelus: [ 30 ] records


armata, Ochotskia Schmidt [P. Yu.] 1916:612, Fig. 1 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Near St. Jona Island, 56°18'N, 145°04'E, Okhotsk Sea, depth 175 fathoms. Holotype (unique): ZIN 18968. •Valid as Icelus armatus (Schmidt 1916) -- (Nelson 1984:22 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S76 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Parin et al. 2014:253 [ref. 33547], Dyldin & Orlov 2022:45 [ref. 38972]). Current status: Valid as Icelus armatus (Schmidt 1916). Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

beringianus, Icelus bicornis Schmidt [P. Yu.] 1927:7 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Providence Bay, Bering Strait, 63°27' to 64°25'N, 168°48' to 176°27'E, depth 8-40 fathoms. Syntypes: ZIN 21914 (1). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Neelov 1973:597 [ref. 7219], Nelson 1984:36 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Psychrolutidae: Icelinae. Habitat: marine.

bicornis, Cottus Reinhardt [J. C. H.] 1840:9 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) v. 7 (1839); ref. 18617] East Greenland. No types known. Type catalog: Nielsen 1974:81 [ref. 9588]. Published two more times in 1840-41. Cottus bicornis Anonymous (Volta) 1796:clxiv, Pl. 39 (fig. 4) [ref. 30407] from the 'monti del Veronese' (mountains of Province Verona, Italy) is an homonym in fossil fishes; Agassiz 1835:293 [ref. 30408] noted that the original material is lost and the illustration in Volta 1796 is unidentifiable; the name Cottus bicornis Anonymous (Volta) 1796 was not used as valid since 1899, therefore reversal of precedence according to ICZN Art. 23.2 and 23.9 in favor of Cottus bicornis Reinhardt 1840. •Valid as Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:35 [ref. 5391], Fedorov in Whitehead et al. 1986:1250 [ref. 13677], Robins & Ray 1986:282 [ref. 23100], Scott & Scott 1988:498 [ref. 25518], McAllister 1990:198 [ref. 14674], Yabe in Okamura et al. 1995:164 [ref. 22531], Mecklenburg et al. 2002:456 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Dolgov 2013:918 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:253 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:43, 173 [ref. 36199], Mecklenburg et al. 2016:67 [ref. 34440], Mecklenburg et al. 2018:205 [ref. 36370], Love et al. 2021:122 [ref. 39279], Zorina & Chernova. 2022:89 [ref. 40474]). 1°Homonym. Current status: Valid as Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Distribution: Circumpolar in Arctic Ocean, North Atlantic and North Pacific. Habitat: marine.

bispinis, Icelus spatula Andriashev [A. P.] 1937:270, Pl. 6 (fig. 5) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] Penzhina Bay to Chanjanga; northeastern Sachalin Island; and northwestern Kamchatka Peninsula, Okhotsk Sea, depth 55-150 meters and to 240 meters. Syntypes: ZIN 17787 (1), 17790-91 (1, 3), 18060 (1), 21920-23 (5, 6, 6, 2). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Nelson 1984:42 [ref. 5391]). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Psychrolutidae: Icelinae. Habitat: marine.

canaliculatus, Icelus Gilbert [C. H.] 1896:412, Pl. 24 (lower) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North of Unalaska, Albatross station 3329, depth 399 fathoms. Lectotype: USNM or CAS-SU 2182. Paralectotypes (incl. potential lectotype): BMNH 1896.7.23.80 (1); MCZ 28346 (1); CAS-SU 2182 (1, poor condition), 18828 (3); USNM 48590 (1), 126722 (1); ZMUO J1459 (1). Type catalog: Böhlke 1953:130 [ref. 12291], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen from Albatross sta. 3329) in caption to Pl. 286, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable; SU 2182 designated lectotype by Nelson 1984:32 [ref. 5391]. •Valid as Icelus canaliculatus Gilbert 1896 -- (Yabe in Amaoka et al. 1983:281, 337 [ref. 8379], Nelson 1984:32 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:452 [ref. 25968], Nakabo 2002:634 [ref. 26001], Parin et al. 2002:S77 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:253 [ref. 33547], Kells et al. 2016:178 [ref. 35888], Dyldin & Orlov 2022:45 [ref. 38972] with question, Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelus canaliculatus Gilbert 1896. Psychrolutidae: Icelinae. Distribution: North Pacific: northern Hokkaido (Japan) and Sea of Okhotsk north to northern Bering Sea, east to western Aleutian islands (U.S.A.). Habitat: marine.

cataphractus, Agonocottus Pavlenko [M. N.] 1910:23, Figs. 2-3 [Trudy Obshchestva Estestvoispytatelej Imperatorskom' Kazanskom' Universiteti, Kazan v. 42 (no. 2); ref. 3393] Near Cape Povorotnyi, Peter the Great Bay, Russia, Sea of Japan, depth 812-875 feet (= 116-125 Russian fathoms). Holotype (unique): not researched. •Synonym of Icelus spiniger Gilbert 1896, but a valid supspecies -- (Lindberg & Krasyukova 1987:190 [ref. 15964]). •Valid as Icelus cataphractus (Pavlenko 1910) -- (Nelson 1984:28 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Kim & Youn 1992:66 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:633 [ref. 25086], Nakabo 2002:633 [ref. 26001], Choi et al. 2003:262, 575 [ref. 26218], Parin et al. 2002:S77 [ref. 28300], Shinohara et al. 2011:43 [ref. 31715], Song et al. 2012:305 [ref. 36295], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:254 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Kim et al. 2020:103 [ref. 39118], Dyldin et al. 2021:64 [ref. 38804], Dyldin & Orlov 2022:45 [ref. 38972]). Current status: Valid as Icelus cataphractus (Pavlenko 1910). Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

crassus, Icelus uncinalis Andriashev [A. P.] 1937:265, Pl. 5 (fig. 2) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] Penzhina Bay, northeastern Sakhalin Island, Okhotsk Sea. Syntypes: ZIN 21900 (1), 21905 (6), 21910 (2), 25281 (1). "Typus" given on caption to Pl. 5 but specimen not distinguished in text. •Synonym in part of Icelus bicornis (Reinhardt 1840) -- (Nelson 1984:42 [ref. 5391]). •Synonym in part of Icelus ochotensis Schmidt 1927 -- (Nelson 1984:50 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300]). •Synonym in part of Icelus spatula Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:28 [ref. 25650]). •Synonym of Icelus ochotensis Schmidt 1927 -- (Parin et al. 2014:255 [ref. 33547]). Current status: Synonym of Icelus ochotensis Schmidt 1927. Psychrolutidae: Icelinae. Habitat: marine.

derjugini, Icelus bicornis Soldatov [V. K.] in Knipowitsch 1926:124 [Trudy Nauchno-issledovatel'skogo instituta po izucheniya Severea. Moskva [= Transactions of the Institute for Scientific Exploration of the North.] No. 27; ref. 22166] Naiden, White Sea. Holotype: NIISM (whereabouts unknown). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Habitat: marine.

ecornis, Icelus Tsutsui [D.] & Yabe [M.] 1996:302, Fig. 2 [Ichthyological Research v. 43 (no. 3); ref. 22432] Kitami-Yamato Bank, southwestern Okhotsk Sea, off Abashiri, Hokkaido, Japan, 44°30.1'N, 144°14.5'E, depth 208 meters. Holotype: HUMZ 125990. Paratypes: CAS 82510 (2); HUMZ 124100-01 (1, 1), 124326 (1), 124832-34 (1, 1, 1), 124836 (1), 124838 (1), 124845 (1), 124848 (1), 124852 (1), 125966-67 (1, 1), 125975 (1), 125980 (1), 125985 (1), 125991 (1), 125999 (1), 126000 (1), 126002 (1), 126006 (1), 126018 (1), 126020-21 (1, 1); NSMT-P 47286-87 (1, 1); UMMZ 230706 (2). •Valid as Icelus ecornis Tsutsui & Yabe 1996 -- (Nakabo 2000:635 [ref. 25086], Nakabo 2002:635 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501], Dyldin & Orlov 2022:46 [ref. 38972]). Current status: Valid as Icelus ecornis Tsutsui & Yabe 1996. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

euryops, Icelus Bean [T. H.] 1890:41 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off Trinity Island, Alaska, U.S.A., Albatross station 2853, 56°N, 154°20'W, depth 159 fathoms. Lectotype: USNM 45367. Paralectotypes: USNM 227363 [ex USNM 45367] (2). Lectotype selected by Nelson 1984:30 [ref. 5391]. •Valid as Icelus euryops Bean 1890 -- (Nelson 1984:30 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:453 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Maslenikov et al. 2013:10 [ref. 37947], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:254 [ref. 33547], Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelus euryops Bean 1890. Psychrolutidae: Icelinae. Distribution: North Pacific: northern Bering Sea to western Gulf of Alaska (U.S.A.). Habitat: marine.

furciger, Icelus Malm [A. W.] 1865:410 [Förhandlingar vid de Skandinaviske Naturforskarnes. v. 9; ref. 17596] Löken, Gullmarfjärden, Sweden. No types known. •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Habitat: marine.

gilberti, Icelus Taranetz [A. Ya.] 1936:149, Fig. [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Sea of Japan (Vladmir and Olga Bays and Yuzhnyy Cape) and Okhotsk Sea (Aniva Cape), depth 56-180 meters. Syntypes: (many) ZIN 25514 (4), 25517 (1), 25328 (3). •Valid as Icelus gilberti Taranetz 1936 -- (Nelson 1984:21 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:192 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:636 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:636 [ref. 26001], Parin et al. 2002:S77 [ref. 28300], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:254 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Dyldin & Orlov 2022:46 [ref. 38972]). Current status: Valid as Icelus gilberti Taranetz 1936. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

hamatus, Icelus Krøyer [H. N.] 1845:253 [Naturhistorisk Tidsskrift (Kjøbenhavn) (n. s.) v. 1; ref. 2689] Belsund, Spitsbergen. Lectotype: ZMUC P811869. Paralectotypes: ?ZMUC 10-11 (1 of 2). Type catalog: Nielsen 1974:81 [ref. 9588]. Lectotype selected by Nelson 1984:35 [ref. 5391]. •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:35 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Habitat: marine.

hypselopterus, Icelus Fukuzawa [H.], Mori [T.], Matsuzaki [K.] & Kai [Y.] 2022:Figs. [Ichthyological Research v. 70 (no. 1); ref. 38955] Shiretoko Peninsula, Hokkaido Island, Japan. Holotype. Paratypes: (3) •Valid as Icelus hypselopterus Fukuzawa, Mori, Matsuzaki & Kai 2022. Current status: Valid as Icelus hypselopterus Fukuzawa, Mori, Matsuzaki & Kai 2022. Psychrolutidae: Icelinae. Distribution: Northwest Pacific: southern Sea of Okhotsk. Habitat: marine.

intermedius, Icelus spiniger Lindberg [G. U.] & Andriashev [A. P.] 1938:517, Figs. 1-2 [Trudy Gidrobiologicheskoi Ekspeditsii Zoologicheskogo instituta AN SSSR No. 1; ref. 19731] Okhotsk Sea, southeastern Kamchatka, Russia, 52°33.3'N, 154°54'E, depth 185-195 meters. Holotype: ZIN 24470 (185 mm). Paratypes: ZIN 24470 (1, with holotype), 27962 (2), 27969 (3). Appeared first as a nomen nudum in Andriashev 1937:301[not researched]. •Synonym of Icelus spiniger Gilbert 1896 -- (Nelson 1984:24 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:257 [ref. 33547]). Current status: Synonym of Icelus spiniger Gilbert 1896. Psychrolutidae: Icelinae. Habitat: marine.

karaensis, Icelus Soldatov [V. K.] 1923:31 [English p. 74], Fig. 34 [Russkij gidrobiologiceskij zurnal: organ Obscestva Issledovatelej Vody i ee Zizni = Russische hydrobiologische Zeitschrift - Saratow v. 3; ref. 15569] Kara Sea, 70°08'N, 56°26'E, Russia. Syntypes: ZIN 22027 (3). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Neelov 1973:597 [ref. 7219], Nelson 1984:42 [ref. 5391], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Psychrolutidae: Icelinae. Habitat: marine.

mandibularis, Icelus Yabe [M.] in Amaoka et al. 1983:283, Pl. 66; Figs. 28-30 [Fishes from the north-eastern Sea of Japan and the Okhotsk Sea off Hokkaido; ref. 8379] Okhotsk Sea, 44°47.0'N, 144°26.5'E, depth 905 meters. Holotype: HMMZ 78676. Paratypes: HUMZ 78675 (1), 78677-78 (2). •Valid as Icelus mandibularis Yabe 1983 -- (Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Nakabo 2002:634 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501]). Current status: Valid as Icelus mandibularis Yabe 1983. Psychrolutidae: Icelinae. Distribution: Western North Pacific. Habitat: marine.

mororane, Stelgistrum Jordan [D. S.] & Seale [A.] 1906:146, Fig. 4 [Proceedings of the United States National Museum v. 30 (no. 1445); ref. 2496] Mororan harbor, Hokkaido Island, Japan. Holotype (unique): CAS-SU 9260. Type catalog: Böhlke 1953:132 [ref. 12291]. •Synonym of Stelgistrum stejnegeri Jordan & Gilbert 1898 -- (Watanabe 1960:32 [ref. 11959] as morarane, Lindberg & Krasyukova 1987:195 [ref. 15964] as mororanae). •Valid as Icelus mororanis (Jordan & Seale 1906) -- (Tsuruoka et al. 2009:97 [ref. 32107], Parin et al. 2014:255 [ref. 33547], Dyldin & Orlov 2022:46 [ref. 38972]). Current status: Valid as Icelus mororanis (Jordan & Seale 1906). Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Hokkaido (Japan), Kuril Islands and Sakhalin (Russia). Habitat: marine.

ochotensis, Icelus Schmidt [P. Yu.] 1927:4, Figs. 1-2 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Northern Okhotsk Sea, between 58°48.5'N to 53°56'N, 151°18.5'-137°32'E, depth 60-105 meters. Syntypes: (45) ZIN 21898-99 (1, 2), 21901-04 (3, 6, 6, 6). •Valid as Icelus ochotensis Schmidt 1927 -- (Nelson 1984:50 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Kim & Youn 1992:67 [ref. 22287], Suzuki & Uno 1992:4 [ref. 24139], Murai et al. 1994:343 [ref. 21908], Nakabo 2000:635 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:635 [ref. 26001], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S77 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Parin et al. 2014:255 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Kim et al. 2020:103 [ref. 39118], Sonoyama et al. 2020:90 [ref. 37637], Dyldin et al. 2021:65 [ref. 38804], Dyldin & Orlov 2022:46 [ref. 38972]). Current status: Valid as Icelus ochotensis Schmidt 1927. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Sea of Japan to Sea of Okhotsk (Korea, Japan, Russia). Habitat: marine.

omodakae, Icelus Tanaka [S.] 1915:616 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 27; ref. 15023] 50 miles off Hinomisaki, Japan, depth 120 fathoms. Syntypes: (3) ZUMT 6420 (1, lost). •Synonym of Icelus cataphractus (Pavlenko 1910) -- (Nelson 1984:28 [ref. 5391]). Current status: Synonym of Icelus cataphractus (Pavlenko 1910). Psychrolutidae: Icelinae. Habitat: marine.

perminovi, Icelus Taranetz [A. Ya.] 1936:151 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Okhotsk Sea, 56°12'N, 143°49'E. Holotype (unique): No types at ZIN. •Valid as Icelus perminovi Taranetz 1936 -- (Nelson 1984:34 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:634 [ref. 26001], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:72 [ref. 27579], Parin et al. 2014:255 [ref. 33547], Dyldin & Orlov 2022:46 [ref. 38972] with question). Current status: Valid as Icelus perminovi Taranetz 1936. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

rastrinoides, Icelus Taranetz [A. Ya.] 1936:150 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Cave Cape; Olympyad Bay; and 45°05'N, 138°06'E, depth 230-411 meters, coast of Primorye, Russia. Syntypes: (8) ZIN 21445 (3). First mentioned by Schmidt 1935:416 [ref. 3939] in the subgenus Ochotskia Schmidt 1915. See also Nakabo 2002:1525 [ref. 26001]. •Valid as Icelus rastrinoides Taranetz 1936 -- (Nelson 1984:29 [ref. 5391], Lindberg & Krasyukova 1987:193 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Parin et al. 2002:S78 [ref. 28300], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:255 [ref. 33547], Dyldin & Orlov 2022:47 [ref. 38972], Kai et al. 2023:16 [ref. 39903]). Current status: Valid as Icelus rastrinoides Taranetz 1936. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Russia, Japan). Habitat: marine.

sekii, Icelus Tsuruoka [O.], Munehara [H.] & Yabe [M.] 2006:47, Figs. 1-2 [Ichthyological Research v. 53 (no. 1); ref. 28501] 44°02.1'N, 145°13.2'E, off Rausu, Shiretoko Peninsula, Hokkaido Island, Japan, depth 20 meters. Holotype: HUMZ 187896. Paratypes: HUMZ 179902 (1), 187897 (2), 187898 (1), 188505 (1). •Valid as Icelus sekii Tsuruoka, Munehara & Yabe 2006 -- (Parin et al. 2014:256 [ref. 33547], Dyldin & Orlov 2022:47 [ref. 38972]). Current status: Valid as Icelus sekii Tsuruoka, Munehara & Yabe 2006. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Hokkaido (Japan), Kuril Islands and Sakhalin (Russia). Habitat: marine.

spatula, Icelus Gilbert [C. H.] & Burke [C. V.] 1912:41, Figs. 3, 3a [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off Avatcha Bay, Kamchatka, Russia, 52°47'20"N, 158°44'30"E, Albatross station 4794, depth 58-69 fathoms. Holotype: USNM 74367. Paratypes: CAS-SU 40931 (5), USNM 70904-05 (4, 4). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelus spatula Gilbert & Burke 1912 -- (Neelov 1973:597 [ref. 7219], Nelson 1984:41 [ref. 5391], Fedorov in Whitehead et al. 1986:1251 [ref. 13677], Robins & Ray 1986:282 [ref. 23100], Scott & Scott 1988:499 [ref. 25518], McAllister 1990:199 [ref. 14674], Yabe in Okamura et al. 1995:164 [ref. 22531], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:455 [ref. 25968], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:72 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Mecklenburg et al. 2007:176 [ref. 29448], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:122 [ref. 31212] with comments, Page et al. 2013:123 [ref. 32708], Dolgov 2013:918 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:256 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:45, 174 [ref. 36199], Kells et al. 2016:178 [ref. 35888], Mecklenburg et al. 2016:69 [ref. 34440], Mecklenburg et al. 2018:207 [ref. 36370], Dyldin & Orlov 2022:47 [ref. 38972], Love et al. 2021:122 [ref. 39279], Zorina & Chernova. 2022:89 [ref. 40474]). Current status: Valid as Icelus spatula Gilbert & Burke 1912. Psychrolutidae: Icelinae. Distribution: Circumpolar, Arctic Ocean, northwestern Atlantic and North Pacific. Habitat: marine.

spiniger, Icelus Gilbert [C. H.] 1896:412, Pl. 24 (top) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Alaska, U.S.A., 54°48'30"N, 165°49'00"W, Albatross station 3225, depth 85 fathoms. Lectotype: ?USNM 164258. Paralectotypes: BMNH 1900.7.11.7 (1?); MCZ 28270 (3); MNHN 1896-0484 to 0486 [ex USNM] (3); CAS-SU 2231 (40), 2266 (2), 2269 (16); ZMUO J1463 (3, poor condition). Type catalog: Böhlke 1953:130 [ref. 12291], Blanc & Hureau 1968:54 [ref. 20738], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen from Albatross sta. 3278) in caption to Pl. 286, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable; a lectotype (USNM 164248, sta. 3225) was also designated by Nelson 1984:24 [ref. 5391]. •Valid as Icelus spiniger Gilbert 1896 -- (Watanabe 1960:42 [ref. 11959], Eschmeyer & Herald 1983:174 [ref. 9277], Nelson 1984:24 [ref. 5391], Peden & Hughes 1986:7 [ref. 9714], McAllister 1990:199 [ref. 14674], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:451 [ref. 25968], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:73 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Mecklenburg et al. 2011:122 [ref. 31212] as cf. spiniger with comments, Page et al. 2013:123 [ref. 32708], Parin et al. 2014:256 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Mecklenburg et al. 2016:71 [ref. 34440] see remarks, Kells et al. 2016:180 [ref. 35888], Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelus spiniger Gilbert 1896. Psychrolutidae: Icelinae. Distribution: North Pacific: Sea of Okhotsk north to northern Bering Sea, then south to southern British Columbia (Canada). Habitat: marine.

spitzbergensis, Icelus bicornis Schmidt [P. Yu.] 1927:7 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Spitsbergen. Syntypes: ZIN 11867 (3), 11900-02 (1, 4, 6+), 12750-52 (3, 2, 5), 12754-55 (13, 3), 12807-08 (1, 1), 13993-97 (6+, 6+, 1, 1, 1). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Habitat: marine.

stenosomus, Icelus uncinalis Andriashev [A. P.] 1937:266, Pl. 5 (fig. 3) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] 48°08'30"N, 140°08'30'E, Tatar Strait. Holotype: ZIN 17646. Paratypes: ZIN 17641-42 (6, 2), 17645 (4), 17646A (5, with holotype), 17647 (3), 21906-08 (1, 1, 1). "Typus" given on caption to Pl. 5, specimen noted in catalog in 17646 and regarded as holotype. •Synonym of Icelus uncinalis Gilbert & Burke 1912, but a valid subspecies as described -- (Lindberg & Krasyukova 1987:194 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670]). •Valid as Icelus stenosomus Andriashev 1937 -- (Nelson 1984:53 [ref. 5391], Murai et al. 1994 [ref. 21908], Nakabo 2000:635 [ref. 25086], Nakabo 2002:635 [ref. 26001], Parin et al. 2002:S78 [ref. 28300], Ko et al. 2009:61 [ref. 37344], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:257 [ref. 33547], Kim et al. 2020:103 [ref. 39118], Sonoyama et al. 2020:90 [ref. 37637], Dyldin & Orlov 2022:47 [ref. 38972]). Current status: Valid as Icelus stenosomus Andriashev 1937. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia). Habitat: marine.

toyamensis, Ricuzenius (Alloricuzenius) Matsubara [K.] & Iwai [T.] 1951:87, Figs. 1-3 [Miscellaneous Reports of the Research Institute for Natural Resources (Tokyo) Nos 19-21; ref. 2909] Off Uozu, Japan. Holotype: FAKU 14450 (not seen in 1997). Paratypes: FAKU 14451-57 (7, not seen in 1997). •Valid as Ricuzenius toyamensis Matsubara & Iwai 1951 -- (Watanabe 1960:22 [ref. 11959], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:186 [ref. 15964]). •Valid as Icelus toyamensis (Matsubara & Iwai 1951) -- (Nakabo 2000:633 [ref. 25086], Nakabo 2002:633, 1525 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501], Shinohara et al. 2011:44 [ref. 31715], Song et al. 2012:304 [ref. 36295], Shinohara et al. 2014:250 [ref. 33330], Kim et al. 2020:103 [ref. 39118]). •Synonym of Icelus rastrinoides Taranetz 1936 -- (Kai et al. 2023:16 [ref. 39903]). Current status: Synonym of Icelus rastrinoides Taranetz 1936. Psychrolutidae: Icelinae. Habitat: marine.

uncinalis, Icelus Gilbert [C. H.] & Burke [C. V.] 1912:39, Figs. 2, 2a [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Near Attu Island, Aleutian Islands, Bering Sea, 52°55'40"N, 173°26'E, Albatross station 4784, depth 135 fathoms. Holotype: USNM 74366. Paratypes: CAS-SU 25237 (2); USNM 70855 (1), 70885 (2), 70892 (1), 70897 (2), 70912 (3). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelus uncinalis Gilbert & Burke 1912 -- (Nelson 1984:52 [ref. 5391], Kim & Youn 1992:67 [ref. 22287], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:454 [ref. 25968], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S79 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:257 [ref. 33547], Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelus uncinalis Gilbert & Burke 1912. Psychrolutidae: Icelinae. Distribution: North Pacific: Bering Sea north of Commander Islands (Russia) and western Aleutian Islands (U.S.A.). Habitat: marine.

vicinalis, Icelus Gilbert [C. H.] 1896:413 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay, Alaska, U.S.A., 53°33'50"N, 167°46'50"W, Albatross station 3324, depth 200 meters. Lectotype: USNM 48737. Paralectotypes: BMNH 1900.7.11.8 (1); CAS 45714 [ex IU 6887] (2); MCZ 28344 (2); MNHN 1896-0487 and 0488 (2) station 3331; CAS-SU 3018 (12), 3484 (1); USNM 48597 (1), 53038 (2), 103605 (2), 233758-59 (1, 2); ZMUO J1464 (2, dry) station 3331. Type catalog: Böhlke 1953:130 [ref. 12291], Blanc & Hureau 1968:54 [ref. 20738], Pethon 1969:8 [ref. 19268]. Lectotype established by Jordan & Evermann 1898:1917 [ref. 2444], but if USNM 48737 contained more than one specimen at that time then lectotype selected by Nelson 1984:30 [ref. 5391]. •Synonym of Icelus euryops Bean 1890 -- (Nelson 1984:30 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:453 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:254 [ref. 33547]). Current status: Synonym of Icelus euryops Bean 1890. Psychrolutidae: Icelinae. Habitat: marine.


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