Institute for Biodiversity Science and Sustainability
Online Version, Updated 8 September 2021

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Eschmeyer's Catalog of Fishes
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granulosus, Benthophilus Kessler [K. F.] 1877:57, Pl. 5 (fig. 14) [The Aralo-Caspian Expedition; ref. 2597] Baku Bay, Caspian Sea, Azerbaijan. Lectotype: ZISP [= ZIN] 2242. Syntypes: (7 + 1) BMNH 1897.7.5.14 [ex ZIN] (1), ZIN 10898 (1, damaged). Lectotype selected by Boldyrev & Bogutskaya 2007:51 [ref. 29150]. •Valid as Benthophilus granulosus Kessler 1877 -- (Berg 1949:1119 [ref. 20662], Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:128 [ref. 23591], Bogutskaya et al. 2001:47 [ref. 26178], Pinchuk & Miller in Miller 2004:192 [ref. 27853], Bogutskaya & Naseka 2004:216 [ref. 28183], Boldyrev & Bogutskaya 2007:49 [ref. 29150], Kottelat & Freyhof 2007:558 [ref. 29996], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:80 [ref. 35254], Esmaeili et al. 2018:61 [ref. 36089], Jouladeh-Roudbar et al. 2020:227 [ref. 37983], Kovačić et al. 2021:62 [ref. 38346]). Current status: Valid as Benthophilus granulosus Kessler 1877. Gobiidae: Gobiinae. Distribution: Eurasia: southern Caspian Sea basin. Habitat: freshwater, brackish.