Institute for Biodiversity Science and Sustainability
Online Version, Updated 7 July 2021

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Eschmeyer's Catalog of Fishes
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borysthenicus, Squalius Kessler [K. F.] 1859:545 [Bulletin de la Société Impériale des Naturalistes de Moscou v. 32 (pt 1); ref. 18392] An arm of the Dnieper River at Aleschki, Ukraine. Syntypes: none found at ZIN. Also appeared as new in Kessler 1860:211, Pl. 1 (fig. 3) [ref. 18393]. •Valid as Leuciscus borysthenicus (Kessler 1859) -- (Berg 1949:560 [ref. 20661], Bogutskaya 1996:26 [ref. 22945], Bianco et al. 1996:51 [ref. 26724], Kottelat 1997:64 [ref. 22952], Bogutskaya 1997:172 [ref. 23566], Reshetnikov et al. 1997:738 [ref. 24702], Bogutskaya 1998:63 [ref. 23589], Bogutskaya et al. 2001:43 [ref. 26178], Vassilev & Pehlivanov 2005:169 [ref. 28369]). •Valid as Squalius borysthenicus Kessler 1859 -- (Sanjur et al. 2003:23 [ref. 28742], Doadrio & Carmona 2006:200 [ref. 28975], Parin et al. 2014:95 [ref. 33547], Çiçek et al. 2015:147 [ref. 35151]). •Valid as Petroleuciscus borysthenicus (Kessler 1859) -- (Bogutskaya 2002:236 [ref. 26837], Bogutskaya & Naseka 2004:86 [ref. 28183], Freyhof et al. 2006:418 [ref. 28484], Fricke et al. 2007:43 [ref. 29533], Kottelat & Freyhof 2007:220 [ref. 29996], Ninua & Japoshvili 2008:167 [ref. 31251], Coad & Bogutskaya 2010:37 [ref. 30930], Perea et al. 2010:14 [ref. 31802], Barbieri et al. 2015:59 [ref. 35029], Khaefi et al. 2016:158 [ref. 34748], Mohammadian-Kalat et al. 2017:487 [ref. 35603], Turan et al. 2018:[10] [ref. 35758], Freyhof et al. 2018:131 [ref. 35889], Özuluğ & Saç 2019:30 [ref. 36599], Çiçek et al. 2020:252 [ref. 37644], Kuljanishvili et al. 2020:[5] [ref. 37535]). Current status: Valid as Petroleuciscus borysthenicus (Kessler 1859). Leuciscidae: Leuciscinae. Distribution: Eurasia: Black Sea, Sea of Azov and Sea of Marmara watersheds. Habitat: freshwater.