Institute for Biodiversity Science and Sustainability
Online Version, Updated 7 Sep 2022

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Catalog of Fishes
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Eschmeyer's Catalog of Fishes - version of 7 Sep 2022 (Continuously updated since the early 1980s, around the turn of the month.)

In this edition, we provide 213 new species in 2022. In 2021 there were 393 new species described.

We record 36,305 valid fish species, including 18,380 valid fish species with the tag ‘Habitat: freshwater’.


Update from Apr. 5, 2022 - Eschmeyer's Catalog of Fishes contributed to Shoal's New Freshwater Fish Species Report 2021, the first in a planned annual series to promote freshwater species awareness and conservation. Check them out at ShoalConservation.org.


Update from Nov. 2, 2020 - We have updated our existing page, Species by Family/Subfamily, to include counts for genera also. The page is now named Genera/Species by Family/Subfamily.

Updated Apr. 9, 2020 - Authors' Count of Species/Subspecies Descriptions. See how many species/subspecies each author has described.
This list was previously posted as a list of "batting averages", comparing total number of species descriptions to number still considered valid. The "batting average" concept was originally conceived by Dr. Eschmeyer for our 2010 paper "Marine fish diversity: history of knowledge and discovery (Pisces)", and was meant to show differences between previous eras and the modern era. It was not meant to be a critique of any individuals, nor a measure of any individual's scientific acumen. The "batting average" concept having served its purpose, it's no longer relevant to present it. In addition, most individuals are only interested in the total number of new descriptions, which is what we present here. - Jon Fong


We are making additions to:

Van der Laan, R., W. N. Eschmeyer & R. Fricke 2014 (11 Nov.), Family-group names of Recent fishes. Zootaxa Monograph 3882 (1), 1-230. DOI 10.11646/zootaxa.3882.1.1

Download the addenda to the family-group list as a PDF.

See Family Group Names for future updates by R. Van der Laan.


By default, search results will not include item 3 names -- unavailable names that detract from the main list of available names. These are mostly names mentioned in synonymy, names in lists, etc. They are still in the database, so if you want to see them, check the box labeled Include unavailable names before searching.


Citing Eschmeyer's Catalog of Fishes

Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds) 2022.  ESCHMEYER'S CATALOG OF FISHES: GENERA, SPECIES, REFERENCES. (http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp). Electronic version accessed dd mmm 2022.

Fricke, R. (ed) 2022.  ESCHMEYER'S CATALOG OF FISHES: REFERENCES. (http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp). Electronic version accessed dd mmm 2022. [Ron Fricke devoted much time to improving references and journals, many of which had not been examined in 15 to 25 years. Please remember that the title of a paper often differs in three places: the title on the paper itself, the contents listed by the journal, and in the PDF provided by the journal. Ron tends to follow the title on the pdf for recent papers and the title on the original for earlier publications. Many of our dates of publication are documented for priority purposes. Some current journals have a target date, but actual publication is often delayed. For nomenclatural date problems, we will help if we can.]

Van der Laan, R. & Fricke, R. 2022.  ESCHMEYER'S CATALOG OF FISHES: FAMILY-GROUP NAMES. (http://www.calacademy.org/scientists/catalog-of-fishes-family-group-names/). Electronic version accessed dd mmm 2022.

Van der Laan, R., Fricke, R. & Eschmeyer, W. N. (eds) 2022.  ESCHMEYER'S CATALOG OF FISHES: CLASSIFICATION. (http://www.calacademy.org/scientists/catalog-of-fishes-classification/). Electronic version accessed dd mmm 2022.

Fricke, R. & Eschmeyer, W. N. 2022.  ESCHMEYER'S CATALOG OF FISHES: JOURNALS. (http://researcharchive.calacademy.org/research/ichthyology/catalog/journals.asp). Electronic version accessed dd mmm 2022. [Includes all journals appearing in the Catalog, including publication information and ISSN numbers.]

Fricke, R. & Eschmeyer, W. N. 2022  ESCHMEYER'S CATALOG OF FISHES: GUIDE TO FISH COLLECTIONS. (http://researcharchive.calacademy.org/research/ichthyology/catalog/collections.asp). Electronic version accessed dd mmm 2022. [Arranged by museum abbreviation and by country, includes type catalogs and historical publications and www sites where available.]

Fricke, R., Eschmeyer, W. N. & Fong, J. D. 2022  ESCHMEYER'S CATALOG OF FISHES: GENERA/SPECIES BY FAMILY/SUBFAMILY. (http://researcharchive.calacademy.org/research/ichthyology/catalog/SpeciesByFamily.asp). Electronic version accessed dd mmm 2022. [Recalculated with each new version; based on current literature, this provides counts of available genus and species names, valid genera and species, and genera and species described in the last 10 years by family/subfamily.]

Please remember that if you got to Eschmeyer's Catalog of Fishes from Fishbase, you are NOT in Fishbase. Fishbase is a more comprehensive database for a different audience, but they use Eschmeyer's Catalog of Fishes in their authority file.

Some other useful sites for fishes are (1) Fishbase [Fishbase team, Manila]; (2) www.stri.org/sftep -- A comprehensive database on Shorefishes of the Tropical Eastern Pacific (and of the Western Atlantic) [Ross Robertson]; (3) shark-references.com -- A database on elasmobranchs, with over 13,000 references, information on taxa, type specimens, etc. [Jürgen Pollerspöck]; (4) FishWisePro.com -- a comprehensive relational database of more than 99,350 scientific species name combinations and over 34,300 well-identified (mostly marine) fish pictures [Dennis Polack]; (5) Freshwater Fish List, 23rd edition December 2017; (6) ETYFish Project (derivations of Fish Names) by Christopher Scharpf; and (7) BioStor, for Biodiversity Heritage Library references. A very useful site is Worldfish.de [not associated with Fishbase or Worldfish]. Maintained by Michal Mikšik and Erwin Schraml. They post new fish taxa and references as they are published.


Notes

A summary paper was published in 2010 from Eschmeyer's Catalog of Fishes database that might be of interest to users of the Catalog. "Marine fish diversity: history of knowledge and discovery (Pisces)." By William N. Eschmeyer, Ronald Fricke, Jon D. Fong & Dennis A. Polack. Zootaxa, no. 2525:19-50 (2 July 2010, open access, click title to download).

As always, notifying us of missed original descriptions and errors in the database or providing PDFs of recent papers is most welcome.

Bill Eschmeyer edited type localities and standardized many place names. Our style is to use current place names found on an English map (or from Wikipedia and Google sites). Should you need original localities, they can be found in the original descriptions or from particular museum holdings (often online).

Many scientific names of fish species are based on personal names; some were found to be incorrect. For example, Jordan and co-workers spelled Japanese names ending in "a" with a terminal "e" as in matsubarae; that is acceptable, with the name Latinized first. So those names do not end in "i". Remarks are provided by Eschmeyer in this version for most species based on personal names. For a review of this very technical subject, see Dubois 2007: Zootaxa No. 1550:49-68.

New treatment of synonyms: Often an author revises a genus and treats a species in detail but does not mention the species synonyms (if any). If he/she does not mention the synonyms (rather common in recent treatments) and moves the valid “parent” species to a different genus, do we move the synonyms automatically, although the revising author may not have studied or assessed the synonyms? In the past, we did not move the synonyms. Typically, this often resulted in the synonyms being in a different current genus than the valid parent species. We now move the synonyms to the genus of the valid parent species, but no “status reference” showing the new generic placement is found with the synonym. Some synonyms will be moved incorrectly, but it is considered more desirable to have the potential synonyms in the same current genus as the valid parent species.

Some conventions used in Eschmeyer's Catalog

Author plus date and a species code: The code of Zoological Nomenclature does not require a comma between the author and date, only a suggestion. In fact, we consider that confusing as that is traditionally used to show a cited reference in journal publications and not a species authorship and date. Genus+species+author+date (with parentheses as needed) is a (nearly) unique code or formula to define every species [2 known exceptions]. So you will find no comma between the author and date. Some journals are now treating species in this way.

Figures: Under a new species heading, some journals list (a) only the major figure, some list (b) all figures (including maps, habitats and graphs), and (c) some give figures showing the full view and any illustrated parts of the species. We give all figures that show a full view and any anatomical part of the species, so our listing of figures often differs from the treatment by a journal in the original description (as listed in the species heading).

Type localities: We give a current place name that one will find on a Google search or in an ‘English’ atlas. For some species described many years ago, one may find both the original locality and in parentheses a modern name for that locality. Many collections now have their holdings on-line, and it is there one can usually find the actual original locality if needed. Current localities are often augmented by additional information, such as coordinates.

Authors: We have added initials in brackets to distinguish all individual authors of genera and species.

Other sections

Genera/Species of Fishes by Family/Subfamily -- Total valid taxa reflect new taxa added minus taxa moved into synonymy plus taxa newly moved to valid. The classification is as used in Eschmeyer's Catalog of Fishes. Prepared by Bill Eschmeyer and Jon Fong, with modifications by Richard Van der Laan.

Family-Group Names -- Priority applies to Family-group names. It did not in early years. As classifications are refined with use of more subfamilies and tribes, it is required to use the oldest available family-group name for these new taxa.

Ichthyological Collections -- Presented are those specimen collections mentioned in the main Eschmeyer's Catalog of Fishes database, especially as repositories of type specimens. Besides museum names and abbreviations, this includes WWW sites, cross-referencing of abbreviations, and selection of museums by country, along with type catalogs, historical articles, name changes, transfers, and other items for individual collections. Prepared by Ron Fricke and Bill Eschmeyer.

Ichthyological Journals -- There are nearly 2600 individual journals or monograph series in Eschmeyer's Catalog of Fishes database. We provide full citations, and other information such as changes in titles, published duration, ISSN numbers, WWW sites, etc. Prepared by Ron Fricke and Bill Eschmeyer.

Acknowledgements

Acknowledgments for Ron Fricke (2022): Smrutirekha Acharya, Hugo Aguirre, K. V. Akhilesh, Mohammad Alavi-Yeganeh, James Albert, Attaala Ali, Gerald Allen, Larry Allen, Pedro Amorim, Marcia da Silva Anjos, Mariangeles Arce, Carlos Ardila Rodríguez, Mercedes Azpelicueta, Nicolas Bailly, Andrey Balanov, Rafael Bañón, Luís Barcelos, Monica Barone, Dibakar Bhakta, Roger Bills, Jamille Bitencourt, Sergey Bogorodsky, Nina Bogutskaya, Jörg Bohlen, Dmitry Bolshakov, Yana Bolshakova, Ralf Britz, Rodrigo Caires, Darija Ćaleta, Marko Ćaleta, Miguel Carneiro, Fernando Carvalho, Albert Chakona, Bruno Chanet, Chen Xiao-Yong, Yves Cherel, Natalia Chernova, Erdogan Çiçek, Kevin Conway, Peter Coulson, Fabio di Dario, Chengzhi Ding, Guilherme Dutra, Yury Dyldin, Soheil Eagderi, David Ebert, Leandro Eduardo, Karl Einum, Deniz Ergüden, Hamid Esmaeili, Lloyd Findley, Tilmann Fischer, Sergio Floeter, Jon Fong, Malcolm Francis, Jörg Freyhof, Sergei Frolov, Eric Garcia, José García-Marín, Michael Ghedotti, Daniel Golani, Mariano González-Castro, Menachem Goren, Lance Grande, Steven Grant, Michael Hammer, Veryl Hasan, Philip Hastings, Harutaka Hata, Amanda Hay, Albert van der Heiden, Fabian Herder, Eric Hilton, Hans Ho, Douglass Hoese, Nazia Hossain, Mauricio Hoyos Padilla, Robert Hrabik, Manuel Irigoitia, A. K. Jaiswar, Laith Jawad, David Johnson, Jeff Johnson, Mikael Karlsson, Axel Katz, Kouichi Kawamura, David Kelly, Anne Kemp, Jin-Koo Kim, Seishi Kimura, Peter Konstantinidis, Laishram Kosygin, Alexander Kotlyar, Kole Kubicek, Rajan Kumar, Ram Kumar, Ranjith Lakshmanan, Samuel Lalronunga, Oscar Lasso-Alcalá, Peter Last, Robert Lea, Pierre-Yves Le Bail, Kleber Mathubara Leite, Boris Levin, Liao Teyu,Thomas Litz, Tobit Liyandja, Jesús López-Vila, Carlos de Lucena, Manoela Marinho, Rene Martin, Fernanda Martins, Ed Matheson, Mizuki Matsunuma, George Mattox, Marissa McNamara, Lucas Medeiros, Bruno Melo, Marcelo Melo, Rafael Menezes, Michael Mincarone, Swarup Mohanty, Anil Mohapatra, Frederik Mollen, Luis Fernando del Moral Flores, Hamed Mousavi-Sabet, Bruce Mundy, Perica Mustafić, Nozomu Muto, Naohide Nakayama, Muddula Krishna Naranji, Matthew Neilson, Heok Hee Ng, Dalton Nielsen, Jørgen Nielsen, Marianne Nyegaard, Claudio Oliveira, Keltoum Ouassal, Dilruba Öztürk, Lawrence Page, Yvan Papa, Kodeeswaran Paramasiwam, Paolo Parenti, Andy Patel, Gretta Pecl, Teguh Peristiwady, Sabine Petri, Frank Pezold, Wojciech Piasecki, Ted Pietsch, Kyle Piller, John Pogonoski, Dennis Pollack, Lucia Pombo-Ayora, Stuart Poss, Jayasimhan Praveenraj, Francisco Provenzano, Artem Prokofiev, Olga Radchenko, Sally Reader, Lais Reia, Renan Borges dos Reis, Roberto Esser dos Reis, Claude Renaud, Oldřich Říčan, Shannon Robinson, Carla Riva Rossi, Luiz Rocha, Lucas Rodrigues, Vladimir Romanov, Barry Russell, Joao Sales, Srikanta Samanta, Keiichi Sato, Mao Sato, Michelle Schärer, Christopher Scharpf, Frederic Schedel, Nalani Schnell, Erwin Schraml, Burak Seçer, Arya Sen, Jaya Kishor Seth, Ilia Shakhovskoy, Bungdon Shangningam, Sharif Sharifuzzaman, Sergei Shedko, Boris Sheiko, Gento Shinohara, Brian Sidlauskas, Gabriel Silva, Victor da Silva, Mahender Singh, Paul Skelton, David Smith, William Smith-Vaniz, Jos Snoeks, Karla Soares, Rainer Sonnenberg, Maarten van Steenberge, Nir Stern, Yo Su, Sandhya Sukumaran, Sun Zhi-Xian, Sevil Sungur, S. Sureshkumar, David Takhnishvili, James van Tassell, José Tavera Vargas, Luke Tornabene, Eloisa Torres-Hernández, David Townsend, Davut Turan, Franz Uiblein, Eliza Uzunova, Ekaterina Vasil'eva, Luca Vecchioni, Risto Viäinölä, Benjamin Victor, Leandro Villa-Verde, Kirill Vinnikov, Olga Voskoboinikova, Nathan Vranken, Simon Weigmann, David de Weirdt, Livi Wilson, Richard Winterbottom, Peter Wirtz, Sercan Yapici, Mazaher Zamani-Faradonbe, Cláudio Zawadzki, Jouke van der Zee, E Zhang, Mikhail Zhukov.

Acknowledgments for Richard van der Laan (2021-2022): Felipe Abranches, Emmanuel Abwe, Smrutirekha Acharya, Henry Agudelo-Zamora, Erik Åhlander,Mohammad Alavi-Yeganeh, Bader H. Alhajeri, Marcia Silva Anjos, Mariangeles Arce H., Carlos Ardila Rodríguez, Jairo Arroyave, Muthukumarasamy Arunachalam, Sirachai Arunrugstichai, Calder Atta, Nicolas Bailly, Arcady Balushkin, Yusuf Bektas, Ricardo Betancur, Flávio Bockmann, Pedro Bragança, José Carlos Brito, Marko Ćaleta, Pablo Calviño, Mac Campbell, Tiago de Carvalho Faria, David Černý, Xiao Chen, Brian Coad, Sebastián Cona, Kevin Conway, Neelesh Dahanukar, Fabio di Dario, Arup Das, Vahe Demirjian, Gaël Denys, Eddy Derijst, Omar Domínguez-Domínguez, Laura Donin, Emanuell Duarte-Ribeiro, David Ebert, Guillain Estivals, Dario Faustino-Fuster, Antonia Ford, Acacio Freitas, Jörg Freyhof, Ron Fricke, Dan Fromm, Augusto Frota, Kyoji Fujiwara, Babette van Gool, Harutaka Hata, Elaine Heemstra, Elisabeth Henschel, Hans Ho, Jean Huber, Lily Hughes, Samuel Iglésias, Taiki Ito, Howard Jelks, Wilbert Kadye, Manda Kambikambi, Axel Katz, Daemin Kim, Seishi Kimura, Steen Knudsen, Keita Koeda, Taiga Kunishima, Paul van der Laan, Francisco Langeani, Sébastien Lavoué, Mao-Ying Lee, Eliseo Lescún, Flávio Lima, Thomas Litz, Thiago Silva Loboda, Edgardo Londoño-Cruz, Silvia Lustosa-Costa, Diego Luzzatto, James Maclaine, Katemo Manda, Mizuki Matsunuma, Madhava Meegaskumbura, Bruno Melo, Michal Mikšík, Michael Mincarone, Guido Miranda, Marcos Mirande, Ryo Misawa, Anil Mohapatra, Daniel Mokodongan, Peter Møller, Nozomu Muto, Robert Myers, Dirk Neumann, Makoto Okamoto, Claudio Oliveira, Renildo de Oliveira, Larry Page, Kody Paramasivam, Murilo Pastana, Edson Pereira, Tiago Pessali, Flávia Petean, Dennis Polack, Jayasimhan Praveenraj, Olga Radchenko, Divya Radhakrishnan, Filipe Rangel-Pereira, Lais Reia, Vinícius Reis, Roberto Reis, Ross Robertson, Uwe Römer, Mark Sabaj, David de Santana, Vanessa de Santis, Mudjie Santos, Sérgio Santos, Luisa Sarmento-Soares, Serkan Saygun, Chris Scharpf, Erwin Schraml, Iraj Segherloo, Hiroshi Senou, Emilia Sferco, Bungdon Shangningam, Sergei Shedko, Bakhtiyor Sheraliev, Brian Sidlauskas, Priscilla Caroline Silva, Mahender Singh, Pratima Singh, Dave Smith, Bill Smith-Vaniz, Yan Soares, Gabe Somarriba, Hiranya Sudasinghe, John Sullivan, Milton Tan, Kisekelwa Tchalondawa, Andrea Thomaz, Shinichi Tomiyama, Tom Trnski, Massimiliano Virgilio, Olga Voskoboinikova, Nathan Vranken, Emmanuel Vreven, Simon Weigmann, Jeff Williams, Lei Yang, Jouke van der Zee, W. Zhou, Dan Zimberlin and Elsbeth Zwart.

Acknowledgments for Bill Eschmeyer & Jon Fong (2022): Martin Byttebier, Fábio Vieira.

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Example of
Search Expression
Database
Hits
single word
Salmo
Genera
75
phrase
"valid as Albula vulpes"
Species
1
 
"synonym of Albula vulpes"
Species
21
 
"Comoro Is"
Species
34
x and y
Cuvier and Valenciennes
References
55
x or y
Cuvier or Valenciennes
References
250
x and not y
Salmo and not Salmonidae
Species
109
 
Salmo and not Salmonidae and not Osmeridae
Species
98
x and not (y or z)
Salmo and not (Salmonidae or Osmeridae)
Species
98
wild card (*)
"Boo*" and not book
Genera
14

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