Institute for Biodiversity Science and Sustainability
Online Version, Updated 8 September 2021

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Eschmeyer's Catalog of Fishes
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quadrimaculata, Clupea Rüppell [W. P. E. S.] 1837:78, Pl. 21 (fig. 3) [Neue Wirbelthiere zu der Fauna von Abyssinien gehörig. Fische des Rothen Meeres; ref. 3844] Bay of Massawa, Eritrea, Red Sea. Lectotype: SMF 4648. Paralectotypes: SMF 78 (1), 4649-52 (4). Lectotype selected by Dor 1984:41 [ref. 29757]. •Valid as Herklotsichthys quadrimaculatus (Rüppell 1837) -- (Dor 1984:41 [ref. 29757], Uyeno & Sato in Masuda et al. 1984:19 [ref. 6441], Whitehead 1985:81 [ref. 5141], Whitehead & Wongratana 1986:201 [ref. 6284], Allen & Swainston 1988:30 [ref. 25477], Paxton et al. 1989:154 [ref. 12442], Randall et al. 1990:47 [ref. 15987], Krishnan & Mishra 1994:274 [ref. 24014], Goren & Dor 1994:10 [ref. 25356], Randall 1995:64 [ref. 22896], Allen 1997:50 [ref. 23977], Carpenter et al. 1997:111 [ref. 23922], Kuiter 1997:32 [ref. 25488], Randall et al. 1997:47 [ref. 25919], Myers 1999:60 [ref. 23965], Munroe et al. 1999:1801 [ref. 24728], Nakabo 2000:246 [ref. 25086], Munroe & Nizinski in Randall & Lim 2000:587 [ref. 25122], Laboute & Grandperrin 2000:342 [ref. 25191], Kimura & Peristiwady in Matsuura & Peristiwady 2000:135 [ref. 27511], Sakai et al. 2001:86 [ref. 25693], Bijukumar & Sushama 2000:184 [ref. 25703], Hutchins 2001:18 [ref. 25847], Nakabo 2002:246 [ref. 26001], Allen & Adrim 2003:24 [ref. 26830], Manilo & Bogorodsky 2003:S96 [ref. 27377], Kimura et al. in Kimura & Matsuura 2003:16 [ref. 27510], Randall et al. 2004:7 [ref. 27624], Randall 2005:55 [ref. 28239], Mundy 2005:148 [ref. 28379], Fricke & Kulbicki 2006:319 [ref. 28867], Paxton et al. 2006:328 [ref. 28995], Randall 2007:100 [ref. 30952], Fricke et al. 2009:22 [ref. 30213], Fricke et al. 2011:355 [ref. 31242], Allen & Erdmann 2012:120 [ref. 31980], Fricke et al. 2013:251 [ref. 32706], Larson et al. 2013:35 [ref. 32988], Kottelat 2013:60 [ref. 32989], Fricke et al. 2014:24 [ref. 33932], Psomadakis et al. 2015:152 [ref. 34104], Hata et al. 2015:40 [ref. 34420], Miesen et al. 2016:83 [ref. 34492], Koeda et al. 2016:10[ref. 35333], Motomura et al. 2017:41 [ref. 35490], Fricke et al. 2018:55 [ref. 35805], Golani & Fricke 2018:30 [ref. 36273], Eagderi et al. 2019:25 [ref. 37020], Fricke et al. 2019:57 [ref. 36673], Zajonz et al. 2019:63 [ref. 36871], Psomadakis et al. 2020:290 [ref. 37272]). Current status: Valid as Herklotsichthys quadrimaculatus (Rüppell 1837). Clupeidae. Distribution: Red Sea, Indo-West Pacific: East Africa, South Africa, Persian Gulf, Socotra, Seychelles, Madagascar and western Mascarenes east to Philippines, Samoa and Tonga, north to southern Japan and Ogasawara Islands, south to Western Australia at 21°38'S, Queensland (Australia) south to 16°35'S, and New Caledonia; introduced in Hawaiian Islands. Habitat: brackish, marine.